225 lines
29 KiB
XML
225 lines
29 KiB
XML
<document id="B34BB5412410C5A9BEBA723B5E77EA51" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501108FFB1FF09AD85187193C8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes orangeanus Ballerio, 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="188" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="187" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="8F22BD7BB5216C8845C59CE98B6F3E3C">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:roleTerm id="0D74AE987BD908B28040348806795AF5">Author</mods:roleTerm>
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<mods:namePart id="8BB49EE5A03C9D0EBF2BB0BAA3F03B86">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="B49A377692E8CA262CC10911EABA58AF">text</mods:typeOfResource>
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<mods:title id="D4919736998B23FB949584C0B1C0F58B">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="054AB2E617DE18483863CAE52FA6788B">
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<mods:date id="82F777D6B12CE4ED448FA0E75097C1B4">2021</mods:date>
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<mods:number id="38A2E776412750E34662BC6CD008CFC4">2021-11-30</mods:number>
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<mods:detail id="C3B73CCC3DDE7C8157C42D2D2283840C" type="volume">
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<mods:number id="C9FDAACE376C7E65AFC69E00B4B4905B">53</mods:number>
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<mods:number id="799D3D254B7F33989560D4F685FB9D77">2</mods:number>
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<mods:start id="02566126B5432B1F0CADFAE0FA13856D">105</mods:start>
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<mods:end id="31AD6A957E6C491425B394BD5AC130DF">282</mods:end>
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<mods:url id="A95855F49CF83B7C83403A07370D5B84">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="8E1A09E9C2BB100FAC2B7AC5F2A3A20E">journal article</mods:classification>
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<mods:identifier id="896681698B1F691184C82BD2F841833D" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="DAF6330AC6887A35C4FE45E93739264F" type="ISSN">2284-4880</mods:identifier>
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<treatment id="0F7475501108FFB1FF09AD85187193C8" LSID="urn:lsid:plazi:treatment:0F7475501108FFB1FF09AD85187193C8" httpUri="http://treatment.plazi.org/id/0F7475501108FFB1FF09AD85187193C8" lastPageId="83" lastPageNumber="188" pageId="82" pageNumber="187">
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<subSubSection id="CFC797CD1108FFB0FF09AD8518D790E8" box="[151,509,692,716]" pageId="82" pageNumber="187" type="nomenclature">
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<paragraph id="8762C4461108FFB0FF09AD8518D790E8" blockId="82.[151,509,692,716]" box="[151,509,692,716]" pageId="82" pageNumber="187">
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<heading id="DC2A732A1108FFB0FF09AD8518D790E8" box="[151,509,692,716]" fontSize="10" level="2" pageId="82" pageNumber="187" reason="5">
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<taxonomicName id="40DDBFC51108FFB0FF09AD85188B90E8" ID-CoL="8SSQV" authorityName="Ballerio" authorityYear="2021" box="[151,417,692,716]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="species" species="orangeanus" status="sp. nov.">
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<emphasis id="B5A918541108FFB0FF09AD85188B90E8" box="[151,417,692,716]" italics="true" pageId="82" pageNumber="187">Goudotostes orangeanus</emphasis>
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</taxonomicName>
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<emphasis id="B5A918541108FFB0FE36AD8518D790E8" bold="true" box="[424,509,692,716]" pageId="82" pageNumber="187">
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<taxonomicNameLabel id="AE9AA52F1108FFB0FE36AD8518D790E8" box="[424,509,692,716]" pageId="82" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="CFC797CD1108FFB0FF09ADE51B3E9108" pageId="82" pageNumber="187" type="description">
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<paragraph id="8762C4461108FFB0FF09ADE51B3E9108" blockId="82.[151,770,724,812]" pageId="82" pageNumber="187">
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(
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<figureCitation id="1FE6D8C31108FFB0FF01ADE5186F90C8" box="[159,325,724,749]" captionStart="Fig" captionStartId="13.[136,167,1778,1797]" captionTargetBox="[243,1357,166,1751]" captionTargetId="figure-4@13.[223,1373,163,1754]" captionTargetPageId="13" captionText="Fig. 9– a, Synarmostes benaraensis sp. nov. female protibia (at: apical teeth); b, Synarmostes benaraensis sp. nov., male protibia; c, Philharmostes scapulatus, protibia; d, Pseudosynarmostes mitsinjo, female protibia; e, Pseudosynarmostes mistinjo, male protibial; f, Goudotostes orangeanus sp. nov., female protibia; g, Goudotostes orangeanus sp. nov., male protibia; h, Philharmostes scapulatus, female mesotibia; i, Synarmostes benaraensis sp. nov., male mesotibia (mias: male inner apical spur); j, Synarmostes benaraensis sp. nov., female mesotibia; k, Synarmostes annamariae sp. nov., male mesotibia l, Goudotostes orangeanus sp. nov. male mesotibia (mias: male inner apical spur); m, Goudotostes orangeanus sp. nov., metatibia; n, Synarmostes benaraensis sp. nov., metatibia; o, Synarmostes annamariae sp. nov., metatibia; p, Pseudosynarmostes mitinjo, metatibia." figureDoi="http://doi.org/10.5281/zenodo.12764182" httpUri="https://zenodo.org/record/12764182/files/figure.png" pageId="82" pageNumber="187">Figs 9 f, g, l, m</figureCitation>
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,
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<figureCitation id="1FE6D8C31108FFB0FECEADE5184190C8" box="[336,363,724,748]" captionStart="Fig" captionStartId="102.[151,182,1885,1904]" captionTargetBox="[154,1448,166,1862]" captionTargetId="figure-13@102.[148,1454,163,1864]" captionTargetPageId="102" captionText="Fig. 37 – Goudotostes orangeanus sp. nov., mouthparts, a–b, mandibles in dorsal view; c–d, mandibles in ventral view; e, labium and labial palpi; f, maxilla and maxillary palpus; g, detail of galeal brush; h, distal epipharynx." figureDoi="http://doi.org/10.5281/zenodo.12764249" httpUri="https://zenodo.org/record/12764249/files/figure.png" pageId="82" pageNumber="187">37</figureCitation>
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a-h, 51 a-i, 77) http://zoobank.org/
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<uri id="F34CC8441108FFB0FEEEADC51B3E9108" pageId="82" pageNumber="187">
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urn:lsid:zoobank.org:act:
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<uuid id="F37BFE931108FFB0FDE3ADC51B3E9108" pageId="82" pageNumber="187">B18888EC- 8B70-4489-A900-11A333FB6EDB</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD1108FFB0FF23AC65186696C8" pageId="82" pageNumber="187" type="materials_examined">
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<paragraph id="8762C4461108FFB0FF23AC65186696C8" blockId="82.[151,772,852,1932]" pageId="82" pageNumber="187">
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<emphasis id="B5A918541108FFB0FF23AC6518699148" bold="true" box="[189,323,852,876]" pageId="82" pageNumber="187">Type series.</emphasis>
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<materialsCitation id="37B5CE1B1108FFB0FED2AC6518609648" collectingDate="2001-02-22" collectingDateMax="2001-02-28" collectingDateMin="2001-02-22" collectorName="San Francisco, CA & de l'Orangea & Fisher & Griswold" country="Madagascar" county="Calif. Academy of Sciences" determinerName="A. Ballerio" elevation="90" latitude="-12.258889" location="Academy of Sciences" longLatPrecision="21" longitude="49.37472" municipality="Province d'Antsiranana" pageId="82" pageNumber="187" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
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<typeStatus id="58667AE41108FFB0FED2AC6518989148" box="[332,434,852,876]" pageId="82" pageNumber="187" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF1108FFB0FE21AC6418FF9148" box="[447,469,853,876]" count="1" pageId="82" pageNumber="187" type="male">♂</specimenCount>
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, deposited in the collection of the
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<collectingRegion id="45190AA41108FFB0FF78AC45187891A8" box="[230,338,884,908]" country="United States of America" name="California" pageId="82" pageNumber="187">California</collectingRegion>
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<location id="8202929D1108FFB0FEC2AC451B7B91A8" LSID="urn:lsid:plazi:treatment:0F7475501108FFB1FF09AD85187193C8:8202929D1108FFB0FEC2AC451B7B91A8" box="[348,593,884,908]" country="Madagascar" county="Calif. Academy of Sciences" latitude="-12.258889" longLatPrecision="21" longitude="49.37472" municipality="Province d'Antsiranana" name="Academy of Sciences" pageId="82" pageNumber="187" stateProvince="California">Academy of Sciences</location>
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,
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<collectorName id="2A28A1901108FFB0FDFFAC4519959188" pageId="82" pageNumber="187">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20274 /
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<collectingCountry id="FFCA84D61108FFB0FD23ACA519F591E9" name="Madagascar" pageId="82" pageNumber="187">Madagascar</collectingCountry>
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||
:
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<collectingMunicipality id="67065E3C1108FFB0FF77AC8518CF91E8" box="[233,485,948,972]" pageId="82" pageNumber="187">Province d’Antsiranana</collectingMunicipality>
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forêt
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<collectorName id="2A28A1901108FFB0FDBDAC851B8591E8" box="[547,687,948,972]" pageId="82" pageNumber="187">de l’Orangea</collectorName>
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,
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||
<locationDeviation id="96D0DD641108FFB0FD24AC8518BF91C8" location="Ramena Elev." pageId="82" pageNumber="187">
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||
<quantity id="402569A31108FFB0FD24AC851A2991E8" box="[698,771,948,972]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="82" pageNumber="187" unit="km" value="3.6">3.6 km</quantity>
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128° SE Ramena Elev.
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</locationDeviation>
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||
<quantity id="402569A31108FFB0FE00ACE518FD91C8" box="[414,471,980,1004]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="82" pageNumber="187" unit="m" value="90.0">
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||
<elevation id="0CF023751108FFB0FE00ACE518FD91C8" box="[414,471,980,1004]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="82" pageNumber="187" unit="m" value="90.0">90 m</elevation>
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||
</quantity>
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||
<date id="F363E2861108FFB0FE7FACE51BBD91C8" box="[481,663,980,1004]" pageId="82" pageNumber="187" value="2001-02-22" valueMax="2001-02-28" valueMin="2001-02-22">
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||
<collectingDate id="E3271B6E1108FFB0FE7FACE51BBD91C8" box="[481,663,980,1004]" pageId="82" pageNumber="187" value="2001-02-22" valueMax="2001-02-28" valueMin="2001-02-22">22-28 Feb. 2001</collectingDate>
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||
</date>
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||
/
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||
<geoCoordinate id="E2E9A2811108FFB0FD2CACE519FC9628" degrees="12" direction="south" minutes="15" orientation="latitude" pageId="82" pageNumber="187" precision="15" seconds="32" value="-12.258889">12° 15’ 32” S</geoCoordinate>
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||
<geoCoordinate id="E2E9A2811108FFB0FF41ACC518AD9628" box="[223,391,1012,1036]" degrees="049" direction="east" minutes="22" orientation="longitude" pageId="82" pageNumber="187" precision="15" seconds="29" value="49.37472">049° 22’ 29” E</geoCoordinate>
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coll.
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<collectorName id="2A28A1901108FFB0FE56ACC51B269628" box="[456,524,1012,1036]" pageId="82" pageNumber="187">Fisher</collectorName>
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,
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<collectorName id="2A28A1901108FFB0FD84ACC51B579628" box="[538,637,1012,1036]" pageId="82" pageNumber="187">Griswold</collectorName>
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et al.
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<collectingCounty id="6E03BCCA1108FFB0FD59ACC518549608" pageId="82" pageNumber="187">Calif. Academy of Sciences</collectingCounty>
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sifted litter littoral rainforest collection code: BLF3200 /
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<taxonomicName id="40DDBFC51108FFB0FE16AB051BB99668" authorityName="Ballerio" authorityYear="2021" box="[392,659,1076,1100]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="species" species="orangeanus" status="sp. nov.">
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<emphasis id="B5A918541108FFB0FE16AB051BB99668" box="[392,659,1076,1100]" italics="true" pageId="82" pageNumber="187">Goudotostes orangeanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AE9AA52F1108FFB0FD04AB041BFE9668" box="[666,724,1077,1100]" pageId="82" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
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det.
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<determinerName id="72BE28821108FFB0FF06AB6518259648" box="[152,271,1108,1132]" pageId="82" pageNumber="187">A. Ballerio</determinerName>
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2020
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||
</materialsCitation>
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<typeStatus id="58667AE41108FFB0FECFAB6518EA9648" box="[337,448,1108,1132]" pageId="82" pageNumber="187" type="holotype">Holotypus</typeStatus>
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||
<specimenCount id="91DB0FCF1108FFB0FE59AB6418F19648" box="[455,475,1109,1132]" count="1" pageId="82" pageNumber="187" type="male">♂</specimenCount>
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/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B1108FFB0FEE0ABA4186296C8" collectingDate="2001-02-22" collectingDateMax="2001-02-28" collectingDateMin="2001-02-22" collectionCode="CASC, ABCB" collectorName="San Francisco, CA & de l'Orangea & Fisher & Griswold" country="Madagascar" county="Calif. Academy of Sciences" determinerName="A. Ballerio" elevation="90" latitude="-12.258889" location="Academy of Sciences" longLatPrecision="21" longitude="49.37472" municipality="Province d'Antsiranana" pageId="82" pageNumber="187" specimenCount="24" specimenCount-female="12" specimenCount-male="12" stateProvince="California" typeStatus="paratype">
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||
<typeStatus id="58667AE41108FFB0FEE0ABA418CC9689" box="[382,486,1173,1197]" pageId="82" pageNumber="187" type="paratype">Paratypes</typeStatus>
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[
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<specimenCount id="91DB0FCF1108FFB0FE6BABA51B1F9688" box="[501,565,1172,1196]" count="2" pageId="82" pageNumber="187" type="male">2 ♂♂</specimenCount>
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dissected]:
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<specimenCount id="91DB0FCF1108FFB0FD26ABA51A299688" box="[696,771,1172,1196]" count="10" pageId="82" pageNumber="187" type="male">10 ♂♂</specimenCount>
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||
and
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<specimenCount id="91DB0FCF1108FFB0FF5AAB85182796E8" box="[196,269,1204,1228]" count="12" pageId="82" pageNumber="187" type="female">12 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A31108FFB0FE82AB85187896E8" box="[284,338,1204,1228]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="82" pageNumber="187" unit="in" value="17.0">17 in</quantity>
|
||
<collectionCode id="E1CC5C831108FFB0FEC6AB8518B496E8" box="[344,414,1204,1228]" pageId="82" pageNumber="187">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A31108FFB0FE4FAB8518D096E8" box="[465,506,1204,1228]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="82" pageNumber="187" unit="in" value="5.0">5 in</quantity>
|
||
<collectionCode id="E1CC5C831108FFB0FE60AB851B6196E8" box="[510,587,1204,1228]" pageId="82" pageNumber="187">ABCB</collectionCode>
|
||
), same collecting data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1108FFB0FF23ABC518F89508" pageId="82" pageNumber="187" type="diagnosis">
|
||
<paragraph id="8762C4461108FFB0FF23ABC518F89508" blockId="82.[151,772,852,1932]" pageId="82" pageNumber="187">
|
||
<emphasis id="B5A918541108FFB0FF23ABC518199728" bold="true" box="[189,307,1268,1292]" pageId="82" pageNumber="187">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC51108FFB0FEA3ABC51B669728" authorityName="Ballerio" authorityYear="2021" box="[317,588,1268,1292]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="species" species="orangeanus" status="sp. nov.">
|
||
<emphasis id="B5A918541108FFB0FEA3ABC51B669728" box="[317,588,1268,1292]" italics="true" pageId="82" pageNumber="187">Goudotostes orangeanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541108FFB0FDC9ABC51B9A9728" bold="true" box="[599,688,1268,1292]" pageId="82" pageNumber="187">
|
||
<taxonomicNameLabel id="AE9AA52F1108FFB0FDC9ABC51B9A9728" box="[599,688,1268,1292]" pageId="82" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="40DDBFC51108FFB0FDC8AA251BF79708" authorityName=", sensu Ballerio" authorityYear="2008" box="[598,733,1300,1324]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541108FFB0FDC8AA251BF79708" box="[598,733,1300,1324]" italics="true" pageId="82" pageNumber="187">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) medium size, b) dorsal ocular area present (vestigial), c) head with a strong tubercle on clypeal disc, d) pronotal anterior carina monolobate, e) pronotal anterior carina obtusely triangular in frontal view, f) pronotal anterior carina raised without giving rise to a longitudinal sharp carina visible in dorsal view, g) pronotal basal carina monolobate, h) pronotum not saddle-shaped in lateral view, i) pronotal disc with gibbosities, in particular the paradiscal ones are strongly developed, and lacking any distinct carinae and/ or tubercles (apart from the monolobate anterior and posterior carinae), j) elytra with dense strongly raised tubercles and short longitudinal carinae. The species falls within the
|
||
<taxonomicName id="40DDBFC51108FFB0FF49A9E5186894C8" baseAuthorityName="Laporte" baseAuthorityYear="1840" box="[215,322,1748,1772]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="species" species="scabrosus">
|
||
<emphasis id="B5A918541108FFB0FF49A9E5186894C8" box="[215,322,1748,1772]" italics="true" pageId="82" pageNumber="187">scabrosus</emphasis>
|
||
</taxonomicName>
|
||
group of species and can be differentiated by the other species of the group by the combination of characters c), e), f), h) and i).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1108FFB0FF23A8051D6A9568" pageId="82" pageNumber="187" type="description">
|
||
<paragraph id="8762C4461108FFB0FF23A8051D6A9568" blockId="82.[151,772,852,1932]" lastBlockId="82.[831,1453,180,1933]" pageId="82" pageNumber="187">
|
||
<emphasis id="B5A918541108FFB0FF23A805186A9568" bold="true" box="[189,320,1844,1868]" pageId="82" pageNumber="187">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A31108FFB0FE5AA8051B329568" box="[452,536,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="82" pageNumber="187" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A31108FFB0FDF1A8051BE99568" box="[623,707,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="187" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A31108FFB0FF09A86519C29548" box="[151,232,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="82" pageNumber="187" unit="mm" value="1.6">1.6 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A31108FFB0FEAEA86518AB9548" box="[304,385,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="82" pageNumber="187" unit="mm" value="2.5">2.5 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A31108FFB0FE5CA8651B399548" box="[450,531,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="82" pageNumber="187" unit="mm" value="2.5">2.5 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A31108FFB0FDC3A8651B859548" box="[605,687,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="82" pageNumber="187" unit="mm" value="2.4">2.4 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Large-sized
|
||
<taxonomicName id="40DDBFC51108FFB0FDE0A8451BD795A8" authorityName=", sensu Ballerio" authorityYear="2008" box="[638,765,1908,1932]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541108FFB0FDE0A8451BD795A8" box="[638,765,1908,1932]" italics="true" pageId="82" pageNumber="187">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Brown with bronze sheen especially on tubercles and carinae, shiny, dorsum with setigerous punctures bearing a very short seta (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A918541108FFB0FAADAE251C419308" box="[1331,1387,276,300]" italics="true" pageId="82" pageNumber="187">Head</emphasis>
|
||
: W/L ratio = 1.66; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides slightly rounded with strongly unaligned genae; dorsal ocular area vestigial, ventral ocular area medium-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface strongly sculptured, with vertex raised with respect to remaining surface and a strongly raised large tubercle on clypeus, surface of tubercle with dense impressed large simple punctures, remaining head surface with dense impressed large simple punctures, surface along clypeal fore margin covered by some transverse irregular lines. Interpunctural distance of punctures inferior than punctural diameter.
|
||
<emphasis id="B5A918541108FFB0FB3FADA41C2C9088" box="[1185,1286,661,684]" italics="true" pageId="82" pageNumber="187">Pronotum</emphasis>
|
||
: subtrapezoidal (dorsal view) (W/L ratio = 1.78), convex, fore angles truncate; pronotal surface with disc covered by dense impressed large simple punctures, interpunctural distance inferior than punctural diameter, interpunctural space raised, giving a granulose appearance to the surface, sides with some horseshoe-shaped punctures with opening directed outwards; pronotal bead not appreciable in dorsal view; pronotal base distinctly and irregularly raised all over its extension; pronotal sculpturing as follows: monolobate basal carina strongly raised to form a tubercle, inverted V-shaped in frontal view and in dorsal view, monolobate anterior carina strongly raised in correspondence with head, generating a tubercle V-shaped in dorsal view and inverted wide V-shaped in frontal view, disc with a depression in the middle, each side of disc with one raised large tubercle and next to it two parallel transverse carinae reaching outer margin; tubercles marked by deep irregular impressed punctures.
|
||
<taxonomicName id="40DDBFC51108FFB0FCE2AB851AF596E8" box="[892,991,1204,1228]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="82" pageNumber="187" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541108FFB0FCE2AB851AF596E8" box="[892,991,1204,1228]" italics="true" pageId="82" pageNumber="187">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: completely covered by deep large transverse short comma-shaped punctures.
|
||
<emphasis id="B5A918541108FFB0FB0FABE51DFA96C8" box="[1169,1232,1236,1260]" italics="true" pageId="82" pageNumber="187">Elytra</emphasis>
|
||
: slightly longer than wide (W/L ratio = 0.97), subcircular (dorsal view), strongly convex; humeral callus indistinct; elytral surface uniformly covered by large horseshoe-shaped punctures with opening directed backwards on proximal third and outwards and inwards on median and distal thirds, interpunctural distance inferior than punctural diameter; striated articular area hardly visible in lateral view, relatively narrow and short; inferi- or sutural stria complete and deep; marginal area large and distinctly protrudent outwards; elytral articular process well developed, smooth and shiny with a line bordering scutellar sides. Elytral sculpturing as follows: base with one tubercle near apex of scutellum (parasutural tubercle), next to it two strongly raised tubercles, one of them marking humeral area, remaining dorsal surface densely covered with several longitudinal raised short carinae, sides and apical area with denser and irregular tubercles; surface of tubercles made of deep irregular large punctures.
|
||
<emphasis id="B5A918541108FFB0FAFDA9C41C8E9528" box="[1379,1444,1781,1804]" italics="true" pageId="82" pageNumber="187">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A918541108FFB0FC32A8251D6D9508" box="[940,1095,1812,1836]" italics="true" pageId="82" pageNumber="187">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A31108FFB0FB73A8251C159508" box="[1261,1343,1812,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="82" pageNumber="187" unit="mm" value="1.2">1.2 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C31108FFB0FC54A8051D309568" box="[970,1050,1844,1868]" captionStart="Fig" captionStartId="116.[151,182,1885,1904]" captionTargetBox="[183,1408,198,1838]" captionTargetId="figure-13@116.[175,1455,180,1865]" captionTargetPageId="116" captionText="Fig. 51 – Goudotostes orangeanus sp. nov., paratype. a, pronotum of rolled up specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f–i, Four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764281" httpUri="https://zenodo.org/record/12764281/files/figure.png" pageId="82" pageNumber="187">Figs 51</figureCitation>
|
||
f-i.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1108FFB0FCFBA8651C2795A8" pageId="82" pageNumber="187" type="etymology">
|
||
<paragraph id="8762C4461108FFB0FCFBA8651C2795A8" blockId="82.[831,1453,180,1933]" pageId="82" pageNumber="187">
|
||
<emphasis id="B5A918541108FFB0FCFBA8651ACC9548" bold="true" box="[869,998,1876,1900]" pageId="82" pageNumber="187">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular meaning “from Orangea”, the
|
||
<typeStatus id="58667AE41108FFB0FB1DA8441D9B95A9" box="[1155,1201,1909,1933]" pageId="82" pageNumber="187">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1109FFB1FF2EAF85187193C8" pageId="83" pageNumber="188" type="distribution">
|
||
<paragraph id="8762C4461109FFB1FF2EAF85187193C8" blockId="83.[136,756,180,492]" pageId="83" pageNumber="188">
|
||
<emphasis id="B5A918541109FFB1FF2EAF8518E492E8" bold="true" box="[176,462,180,204]" pageId="83" pageNumber="188">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE41109FFB1FD58AF841BDE92E9" box="[710,756,181,205]" pageId="83" pageNumber="188">type</typeStatus>
|
||
locality in northern
|
||
<collectingCountry id="FFCA84D61109FFB1FEC7AFE518F092C8" box="[345,474,212,236]" name="Madagascar" pageId="83" pageNumber="188">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="1FE6D8C31109FFB1FE79AFE51B1E92C8" box="[487,564,212,236]" captionStart="Fig" captionStartId="168.[151,182,1832,1851]" captionTargetBox="[151,1451,166,1794]" captionTargetPageId="168" captionText="Fig. 93 – Habitat, a–d, Ambondrobe forest (photos by Brian Fisher), habitat of Goudotostes litoralis sp. nov.; e–f, Makirovana forest (photos by Brian Fisher), habitat of Synarmostes makirovanae sp. nov., Goudotostes laevis sp. nov., Philharmostes rotundatus sp. nov., and Philharmostes scapulatus sp. nov., and g–h, Orangea forest (g, photo by Brian Fisher in the year 2001 and h, photo by Alberto Ballerio in the year 2005), habitat of Goudotostes andreonei sp. nov. and Goudotostes orangeanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12764384" httpUri="https://zenodo.org/record/12764384/files/figure.png" pageId="83" pageNumber="188">Fig. 93</figureCitation>
|
||
g-h). The
|
||
<typeStatus id="58667AE41109FFB1FD3FAFE41BE592C9" box="[673,719,213,237]" pageId="83" pageNumber="188">type</typeStatus>
|
||
series was collected by sifting litter in littoral forest, although the vegetation cover of the area is also classified as “Dry deciduous forest” (also known as “Western dry forest”,
|
||
<bibRefCitation id="E34CB9B71109FFB1FF16AE6418449348" author="Moat J. & Smith P." box="[136,366,340,364]" pageId="83" pageNumber="188" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
). The forest cover of Oronjia (also known as Orangea) is low and degraded and sometimes takes the form of a dry thicket (
|
||
<bibRefCitation id="E34CB9B71109FFB1FE4BAEA51B909388" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[469,698,404,428]" pageId="83" pageNumber="188" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
). The area falls within the protected area “Paysage Harmonieux Protegé d’Oronjia”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |