224 lines
28 KiB
XML
224 lines
28 KiB
XML
<document id="77C849863535FF1F6C257C44B399A244" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501102FFBBFF09ABE51BA79168" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes rakotonirinai Ballerio, 2021, sp. nov." docType="treatment" docUuid="8C3A8BF0-80F1-467E-B1EE-8C600BC42F1D" docUuidSource="ZooBank" docVersion="2" lastPageNumber="194" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="193" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="88D8683AAD0B340B177B1E7E69FD0713">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:roleTerm id="E6D99B37DA8FB96C0E1FED8C9C3A655B">Author</mods:roleTerm>
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<mods:namePart id="908B9077B2F1B134DDB3106BEDAEF68A">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="B84F00D1FB94B46CC28BA7143F1A38ED">text</mods:typeOfResource>
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<mods:title id="8CCD80CD6C1BE67C4AACA52B30B245A0">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="967EC3EC4FF7D367CA956B5F567DD147">
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<mods:date id="FB60108D4E056DB31F7820A581E1359A">2021</mods:date>
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<mods:number id="A836764BD36040745D5B9DDD222CA9B6">2021-11-30</mods:number>
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<mods:number id="8217A5CEDAF53A7C497C6B5571017C03">53</mods:number>
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<mods:number id="684CD676DD6FE37D90B3C4AAC8725811">2</mods:number>
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<mods:start id="95FAFC856F7F2558356CC2D190497A60">105</mods:start>
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<mods:url id="ACCE2A7E3E75E002E1B295AFFE5728E8">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="5F7679342648175BBB74E125A522F3AF">journal article</mods:classification>
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<mods:identifier id="7F7307260927E865DDFB8CF46ED57A47" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="7E730E96E81E1A0A6509637911976491" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="65B7A887F82D7B017BDC3327395554FE" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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<treatment id="0F7475501102FFBBFF09ABE51BA79168" LSID="urn:lsid:zoobank.org:act:8C3A8BF0-80F1-467E-B1EE-8C600BC42F1D" httpUri="http://treatment.plazi.org/id/0F7475501102FFBBFF09ABE51BA79168" lastPageId="89" lastPageNumber="194" pageId="88" pageNumber="193">
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<subSubSection id="CFC797CD1102FFBAFF09ABE51B2496C8" box="[151,526,1236,1260]" pageId="88" pageNumber="193" type="nomenclature">
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<paragraph id="8762C4461102FFBAFF09ABE51B2496C8" blockId="88.[151,526,1236,1260]" box="[151,526,1236,1260]" pageId="88" pageNumber="193">
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<heading id="DC2A732A1102FFBAFF09ABE51B2496C8" box="[151,526,1236,1260]" fontSize="10" level="2" pageId="88" pageNumber="193" reason="5">
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<taxonomicName id="40DDBFC51102FFBAFF09ABE5189B96C8" ID-CoL="8SSN3" authorityName="Ballerio" authorityYear="2021" box="[151,433,1236,1260]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="rakotonirinai" status="sp. nov.">
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<emphasis id="B5A918541102FFBAFF09ABE5189B96C8" box="[151,433,1236,1260]" italics="true" pageId="88" pageNumber="193">Goudotostes rakotonirinai</emphasis>
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</taxonomicName>
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<emphasis id="B5A918541102FFBAFE26ABE51B2496C8" bold="true" box="[440,526,1236,1260]" pageId="88" pageNumber="193">
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<taxonomicNameLabel id="AE9AA52F1102FFBAFE26ABE51B2496C8" box="[440,526,1236,1260]" pageId="88" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD1102FFBAFF09ABC51B3C9768" pageId="88" pageNumber="193" type="description">
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<paragraph id="8762C4461102FFBAFF09ABC51B3C9768" blockId="88.[151,771,1268,1356]" pageId="88" pageNumber="193">
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(
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<figureCitation id="1FE6D8C31102FFBAFF01ABC519DA9728" box="[159,240,1268,1292]" captionStart="Fig" captionStartId="130.[151,183,1858,1877]" captionTargetBox="[201,1390,159,1827]" captionTargetId="figure-13@130.[147,1455,149,1846]" captionTargetPageId="130" captionText="Fig. 65 – Goudotostes rakotonirinai sp. nov., holotype. a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, pronotum of rolled up specimen. g–k, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764313" httpUri="https://zenodo.org/record/12764313/files/figure.png" pageId="88" pageNumber="193">Figs 65</figureCitation>
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a-k, 77) http://zoobank.org/
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<uri id="F34CC8441102FFBAFEECAA251B3C9768" pageId="88" pageNumber="193">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="F37BFE931102FFBAFD1FAA251B3C9768" pageId="88" pageNumber="193">8C3A8BF0- 80F1-467E-B1EE-8C600BC42F1D</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="CFC797CD1102FFBAFF23AA45185F9528" pageId="88" pageNumber="193" type="materials_examined">
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<paragraph id="8762C4461102FFBAFF23AA45185F9528" blockId="88.[151,772,1396,1932]" pageId="88" pageNumber="193">
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<emphasis id="B5A918541102FFBAFF23AA45186997A8" bold="true" box="[189,323,1396,1420]" pageId="88" pageNumber="193">Type series.</emphasis>
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<materialsCitation id="37B5CE1B1102FFBAFED2AA4519E69488" collectingDate="2005-11-18" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="125" latitude="-16.672222" location="Manompana" longLatPrecision="21" longitude="49.70111" municipality="Reserve Ambodiriana" pageId="88" pageNumber="193" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
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<typeStatus id="58667AE41102FFBAFED2AA45189897A8" box="[332,434,1396,1420]" pageId="88" pageNumber="193" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF1102FFBAFE21AA4418FF97A8" box="[447,469,1397,1420]" count="1" pageId="88" pageNumber="193" type="male">♂</specimenCount>
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, deposited in the collection of the California
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<collectingCounty id="6E03BCCA1102FFBAFEF8AAA51B4B9788" box="[358,609,1428,1452]" pageId="88" pageNumber="193">Academy of Sciences</collectingCounty>
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,
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<collectorName id="2A28A1901102FFBAFDEAAAA519C397E8" pageId="88" pageNumber="193">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20279 / HT: CASENT 8135325 /
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<collectingCountry id="FFCA84D61102FFBAFE24AAE51B1597C8" box="[442,575,1492,1516]" name="Madagascar" pageId="88" pageNumber="193">Madagascar</collectingCountry>
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:
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<collectingRegion id="45190AA41102FFBAFDD3AAE51A2E97C8" box="[589,772,1492,1516]" country="Madagascar" name="Toamasina" pageId="88" pageNumber="193">Toamasina Prov.</collectingRegion>
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<collectingMunicipality id="67065E3C1102FFBAFF09AAC418AC9428" box="[151,390,1524,1548]" pageId="88" pageNumber="193">Reserve Ambodiriana</collectingMunicipality>
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<quantity id="402569A31102FFBAFE0CAAC518C89428" box="[402,482,1524,1548]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="88" pageNumber="193" unit="km" value="4.8">4.8 km</quantity>
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306°
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<location id="8202929D1102FFBAFDB1AAC41B9D9428" LSID="urn:lsid:plazi:treatment:0F7475501102FFBBFF09ABE51BA79168:8202929D1102FFBAFDB1AAC41B9D9428" box="[559,695,1525,1548]" country="Madagascar" county="Academy of Sciences" latitude="-16.672222" longLatPrecision="21" longitude="49.70111" municipality="Reserve Ambodiriana" name="Manompana" pageId="88" pageNumber="193" stateProvince="Toamasina">Manompana</location>
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, along
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<location id="8202929D1102FFBAFF09A924187E9408" LSID="urn:lsid:plazi:treatment:0F7475501102FFBBFF09ABE51BA79168:8202929D1102FFBAFF09A924187E9408" box="[151,340,1556,1580]" country="Madagascar" county="Academy of Sciences" latitude="-16.672222" longLatPrecision="21" longitude="49.70111" municipality="Reserve Ambodiriana" name="Manompana river" pageId="88" pageNumber="193" stateProvince="Toamasina">Manompana river</location>
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elev
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<quantity id="402569A31102FFBAFE12A92518E59408" box="[396,463,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="88" pageNumber="193" unit="m" value="125.0">
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<elevation id="0CF023751102FFBAFE12A92518E59408" box="[396,463,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="88" pageNumber="193" unit="m" value="125.0">125 m</elevation>
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</quantity>
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<date id="F363E2861102FFBAFE4BA9251B489408" box="[469,610,1556,1580]" pageId="88" pageNumber="193" value="2005-11-18">
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<collectingDate id="E3271B6E1102FFBAFE4BA9251B489408" box="[469,610,1556,1580]" pageId="88" pageNumber="193" value="2005-11-18">18 Nov. 2005</collectingDate>
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</date>
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||
/
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||
<geoCoordinate id="E2E9A2811102FFBAFDE8A9251A299408" box="[630,771,1556,1580]" degrees="16" direction="south" minutes="40" orientation="latitude" pageId="88" pageNumber="193" precision="15" seconds="20" value="-16.672222">16° 40’ 20” S</geoCoordinate>
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<geoCoordinate id="E2E9A2811102FFBAFF09A90518199468" box="[151,307,1588,1612]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="88" pageNumber="193" precision="15" seconds="04" value="49.70111">049° 42’ 04” E</geoCoordinate>
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<location id="8202929D1102FFBAFEA7A9051BA49468" LSID="urn:lsid:plazi:treatment:0F7475501102FFBBFF09ABE51BA79168:8202929D1102FFBAFEA7A9051BA49468" box="[313,654,1588,1612]" country="Madagascar" county="Academy of Sciences" latitude="-16.672222" longLatPrecision="21" longitude="49.70111" municipality="Reserve Ambodiriana" name="California Academy of Sciences" pageId="88" pageNumber="193" stateProvince="Toamasina">California Academy of Sciences</location>
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coll.
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<collectorName id="2A28A1901102FFBAFD56A90519DD9448" pageId="88" pageNumber="193">Brian L. Fisher</collectorName>
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et al. sifted litter, rainforest collection code: BLF 12706/
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<taxonomicName id="40DDBFC51102FFBAFF75A9451B2394A8" authorityName="Ballerio" authorityYear="2021" box="[235,521,1652,1676]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="rakotonirinai" status="sp. nov.">
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<emphasis id="B5A918541102FFBAFF75A9451B2394A8" box="[235,521,1652,1676]" italics="true" pageId="88" pageNumber="193">Goudotostes rakotonirinai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AE9AA52F1102FFBAFD8DA9441B7A94A8" box="[531,592,1653,1676]" pageId="88" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
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det.
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<determinerName id="72BE28821102FFBAFD17A9451A2994A8" box="[649,771,1652,1676]" pageId="88" pageNumber="193">A. Ballerio</determinerName>
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2020
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</materialsCitation>
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<typeStatus id="58667AE41102FFBAFF4BA9A5186E9488" box="[213,324,1684,1708]" pageId="88" pageNumber="193" type="holotype">Holotypus</typeStatus>
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<specimenCount id="91DB0FCF1102FFBAFED2A9A4184A9488" box="[332,352,1685,1708]" count="1" pageId="88" pageNumber="193" type="male">♂</specimenCount>
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/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B1102FFBAFE90A9E4185B9528" collectingDate="2005-11-18" collectionCode="CASC" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="125" latitude="-16.672222" location="Manompana" longLatPrecision="21" longitude="49.70111" municipality="Reserve Ambodiriana" pageId="88" pageNumber="193" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
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<typeStatus id="58667AE41102FFBAFE90A9E4185094C9" box="[270,378,1749,1773]" pageId="88" pageNumber="193" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="91DB0FCF1102FFBAFE1AA9E5188694C8" box="[388,428,1748,1772]" count="1" pageId="88" pageNumber="193" type="male">1 ♂</specimenCount>
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and
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<specimenCount id="91DB0FCF1102FFBAFE7FA9E51B2394C8" box="[481,521,1748,1772]" count="1" pageId="88" pageNumber="193" type="female">1 ♀</specimenCount>
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||
(
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<collectionCode id="E1CC5C831102FFBAFD86A9E51B4994C8" box="[536,611,1748,1772]" pageId="88" pageNumber="193">CASC</collectionCode>
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), same collecting data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD1102FFBAFF23A8251DDF90E8" pageId="88" pageNumber="193" type="diagnosis">
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<paragraph id="8762C4461102FFBAFF23A8251DDF90E8" blockId="88.[151,772,1396,1932]" lastBlockId="88.[831,1452,180,1932]" pageId="88" pageNumber="193">
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<emphasis id="B5A918541102FFBAFF23A82518049508" bold="true" box="[189,302,1812,1836]" pageId="88" pageNumber="193">Diagnosis.</emphasis>
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<taxonomicName id="40DDBFC51102FFBAFEA7A8251B7B9508" authorityName="Ballerio" authorityYear="2021" box="[313,593,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="rakotonirinai" status="sp. nov.">
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<emphasis id="B5A918541102FFBAFEA7A8251B7B9508" box="[313,593,1812,1836]" italics="true" pageId="88" pageNumber="193">Goudotostes rakotonirinai</emphasis>
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</taxonomicName>
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<emphasis id="B5A918541102FFBAFDC5A8251B989508" bold="true" box="[603,690,1812,1836]" pageId="88" pageNumber="193">
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<taxonomicNameLabel id="AE9AA52F1102FFBAFDC5A8251B989508" box="[603,690,1812,1836]" pageId="88" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can be differentiated from all other species of
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<taxonomicName id="40DDBFC51102FFBAFDA8A8051B939568" authorityName=", sensu Ballerio" authorityYear="2008" box="[566,697,1844,1868]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="genus">
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<emphasis id="B5A918541102FFBAFDA8A8051B939568" box="[566,697,1844,1868]" italics="true" pageId="88" pageNumber="193">Goudotostes</emphasis>
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</taxonomicName>
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by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection pointed, d) clypeal punctation made of dense impressed transverse comma-shaped punctures, e) fronto-clypeal tubercle present, f) pronotal anterior carina bilobate, g) pronotal basal carina bilobate, h) pronotal disc covered by sparse (interpunctural distance larger than punctural diameter) small simple impressed punctures, i) elytral dorsal area covered by longitudinal rows of irregular blunt carinae of variable length but, on average, short, j) elytral first interstria (area between sutural stria and first longitudinal carina) with sparse punctation (interpunctural distance inferior than punctural diameter), k) left paramere with ventral margin invaginated (a character which places this species in the
|
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<taxonomicName id="40DDBFC51102FFBAFA96AD251C6F9008" authorityName="Ballerio" authorityYear="2021" box="[1288,1349,532,556]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="laevis">
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||
<emphasis id="B5A918541102FFBAFA96AD251C6F9008" box="[1288,1349,532,556]" italics="true" pageId="88" pageNumber="193">laevis</emphasis>
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||
</taxonomicName>
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group of species), l) the distinctive shape of parameres (as in
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<figureCitation id="1FE6D8C31102FFBAFAC3AD051C869068" box="[1373,1452,564,588]" captionStart="Fig" captionStartId="130.[151,183,1858,1877]" captionTargetBox="[201,1390,159,1827]" captionTargetId="figure-13@130.[147,1455,149,1846]" captionTargetPageId="130" captionText="Fig. 65 – Goudotostes rakotonirinai sp. nov., holotype. a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, pronotum of rolled up specimen. g–k, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764313" httpUri="https://zenodo.org/record/12764313/files/figure.png" pageId="88" pageNumber="193">Figs 65</figureCitation>
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g-k). Within the
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<taxonomicName id="40DDBFC51102FFBAFC75AD651D029048" authorityName="Ballerio" authorityYear="2021" box="[1003,1064,596,620]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A918541102FFBAFC75AD651D029048" box="[1003,1064,596,620]" italics="true" pageId="88" pageNumber="193">laevis</emphasis>
|
||
</taxonomicName>
|
||
species group,
|
||
<taxonomicName id="40DDBFC51102FFBAFB56AD651A5390A8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="species" species="rakotonirinai" status="sp. nov.">
|
||
<emphasis id="B5A918541102FFBAFB56AD651A5390A8" italics="true" pageId="88" pageNumber="193">Goudotostes rakotonirinai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541102FFBAFC1FAD451AFC90A8" bold="true" box="[897,982,628,652]" pageId="88" pageNumber="193">
|
||
<taxonomicNameLabel id="AE9AA52F1102FFBAFC1FAD451AFC90A8" box="[897,982,628,652]" pageId="88" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unmistakable because of the combination of transverse comma-shaped punctures on head and sparse simple impressed punctures on pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1102FFBBFCFBADE518CD9008" lastPageId="89" lastPageNumber="194" pageId="88" pageNumber="193" type="description">
|
||
<paragraph id="8762C4461102FFBBFCFBADE518CD9008" blockId="88.[831,1452,180,1932]" lastBlockId="89.[135,757,180,845]" lastPageId="89" lastPageNumber="194" pageId="88" pageNumber="193">
|
||
<emphasis id="B5A918541102FFBAFCFBADE51AC590C8" bold="true" box="[869,1007,724,748]" pageId="88" pageNumber="193">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A31102FFBAFBE7ADE51DFA90C8" box="[1145,1232,724,748]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="88" pageNumber="193" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A31102FFBAFAB4ADE51CAB90C8" box="[1322,1409,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="88" pageNumber="193" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A31102FFBAFCC9ADC51A849128" box="[855,942,756,780]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="88" pageNumber="193" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A31102FFBAFB9CADC51D739128" box="[1026,1113,756,780]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="88" pageNumber="193" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; EL = 2.0 mm; EW =
|
||
<quantity id="402569A31102FFBAFACCADC51C829128" box="[1362,1448,756,780]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="88" pageNumber="193" unit="mm" value="1.9">1.9 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC51102FFBAFCA1AC051AE99168" authorityName=", sensu Ballerio" authorityYear="2008" box="[831,963,820,844]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541102FFBAFCA1AC051AE99168" box="[831,963,820,844]" italics="true" pageId="88" pageNumber="193">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A918541102FFBAFC24ACA51AD99188" box="[954,1011,916,940]" italics="true" pageId="88" pageNumber="193">Head</emphasis>
|
||
: W/L ratio = 1.60; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides almost rectilinear with slightly unaligned genae in correspondence with the genal suture; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae rounded, distinctly protruding outwards, genal suture indistinct; head dorsal surface with vertex covered by medium-sized sparse impressed simple punctures becoming larger and transversely comma-shaped on frons and clypeus, fore margin marked by some irregular transverse lines. Interpunctural distance equal to or inferior than punctural diameter. Fronto-clypeal tubercle distinctly raised, small, pointed and smooth.
|
||
<emphasis id="B5A918541102FFBAFB33AA241C3C9708" box="[1197,1302,1301,1324]" italics="true" pageId="88" pageNumber="193">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 1.90), convex, fore angles distinctly subtruncate; pronotal surface covered by sparse large deep simple punctures becoming comma-shaped at sides, area near fore angles with few irregular lines, interpunctural distance mostly twice to three times punctural diameter; pronotal bead present only along sides and fore angles; bilobate basal carina raised; bilobate anterior carina raised; pronotal sculpturing made of large and gently raised tubercles and carinae as follows: disc with two gently raised tubercles, each side with one gently raised tubercle and one transverse carina reaching outer margin.
|
||
<taxonomicName id="40DDBFC51102FFBAFA9CA9451C4094A8" box="[1282,1386,1652,1676]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="88" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541102FFBAFA9CA9451C4094A8" box="[1282,1386,1652,1676]" italics="true" pageId="88" pageNumber="193">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: covered by sparse impressed transverse large comma-shaped punctures.
|
||
<emphasis id="B5A918541102FFBAFC26A9851AD094E8" box="[952,1018,1716,1740]" italics="true" pageId="88" pageNumber="193">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 1.08), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface uniformly covered by large sparse deep comma-shaped punctures with opening directed backwards mixed to large impressed simple punctures, interpunctural distance about equal to punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria visible complete; marginal area medium-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large tubercle below articular process (parasutural tubercle), next to it a humeral oblique large raised tubercle (humeral tubercle), dorsal area of elytra with three to four irregular and asymmetrical rows of weakly raised smooth carinae, sides and apical third with some more raised tubercles and longitudinal short carinae.
|
||
<emphasis id="B5A918541103FFBBFF17AEC419E39028" box="[137,201,501,524]" italics="true" pageId="89" pageNumber="194">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A918541103FFBBFED6AEC518CF9028" box="[328,485,500,524]" italics="true" pageId="89" pageNumber="194">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A31103FFBBFD06AEC51BC79028" box="[664,749,500,524]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="89" pageNumber="194" unit="mm" value="1.2">1.2 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C31103FFBBFEF6AD25189D9008" box="[360,439,532,556]" captionStart="Fig" captionStartId="130.[151,183,1858,1877]" captionTargetBox="[201,1390,159,1827]" captionTargetId="figure-13@130.[147,1455,149,1846]" captionTargetPageId="130" captionText="Fig. 65 – Goudotostes rakotonirinai sp. nov., holotype. a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, pronotum of rolled up specimen. g–k, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764313" httpUri="https://zenodo.org/record/12764313/files/figure.png" pageId="89" pageNumber="194">Figs 65</figureCitation>
|
||
g-k.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1103FFBBFF30AD051BCE90A8" pageId="89" pageNumber="194" type="etymology">
|
||
<paragraph id="8762C4461103FFBBFF30AD051BCE90A8" blockId="89.[135,757,180,845]" pageId="89" pageNumber="194">
|
||
<emphasis id="B5A918541103FFBBFF30AD0518009068" bold="true" box="[174,298,564,588]" pageId="89" pageNumber="194">Etymology.</emphasis>
|
||
Noun in the genitive case. Named after Jean-Claude Rakotonirina (
|
||
<collectingCountry id="FFCA84D61103FFBBFE3EAD651B359048" box="[416,543,596,620]" name="Madagascar" pageId="89" pageNumber="194">Madagascar</collectingCountry>
|
||
Biodiversity Center,
|
||
<collectingRegion id="45190AA41103FFBBFF16AD45183390A8" box="[136,281,628,652]" country="Madagascar" name="Antananarivo" pageId="89" pageNumber="194">Antananarivo</collectingRegion>
|
||
), member of the Malagasy lab team of CAS.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1103FFBBFF30ADA51BA79168" pageId="89" pageNumber="194" type="distribution">
|
||
<paragraph id="8762C4461103FFBBFF30ADA51BA79168" blockId="89.[135,757,180,845]" pageId="89" pageNumber="194">
|
||
<emphasis id="B5A918541103FFBBFF30ADA518E69088" bold="true" box="[174,460,660,684]" pageId="89" pageNumber="194">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE41103FFBBFD58ADA41BDE9089" box="[710,756,661,685]" pageId="89" pageNumber="194">type</typeStatus>
|
||
locality (Reserve Ambodiriana) along the coast of Eastern
|
||
<collectingCountry id="FFCA84D61103FFBBFF16ADE5182190C8" box="[136,267,724,748]" name="Madagascar" pageId="89" pageNumber="194">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE41103FFBBFED4ADE4185290C9" box="[330,376,725,749]" pageId="89" pageNumber="194">type</typeStatus>
|
||
series was collected by sifting litter in rainforest. The vegetation cover of the area consists of “humid forest” (
|
||
<bibRefCitation id="E34CB9B71103FFBBFEC0AC251B699108" author="Moat J. & Smith P." box="[350,579,788,812]" pageId="89" pageNumber="194" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
) also known as “moist evergreen forest” (
|
||
<bibRefCitation id="E34CB9B71103FFBBFE05AC051BA89168" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[411,642,820,844]" pageId="89" pageNumber="194" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |