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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.520.9581" ID-PMC="PMC4591719" ID-Pensoft-Pub="1313-2970-520-1" ID-Pensoft-UUID="7B52C11DFFEEA14A16685707FF94530E" ID-PubMed="26448711" ID-Zenodo-Dep="579014" ID-ZooBank="B96D0F7DCCC1488DBF144111C90FF9CC" ModsDocID="1313-2970-520-1" checkinTime="1451243998483" checkinUser="pensoft" docAuthor="Pamungkas, Joko &amp; Glasby, Christopher J." docDate="2015" docId="2AB78BC010895740B531BE7DA685FE1A" docLanguage="en" docName="ZooKeys 520: 1-25" docOrigin="ZooKeys 520" docPubDate="2015-09-08" docSource="http://dx.doi.org/10.3897/zookeys.520.9581" docTitle="Neanthes gisserana" docType="treatment" docVersion="3" id="7B52C11DFFEEA14A16685707FF94530E" lastPageNumber="8" masterDocId="7B52C11DFFEEA14A16685707FF94530E" masterDocTitle="Taxonomy of reproductive Nereididae (Annelida) in multispecies swarms at Ambon Island, Indonesia" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1668160663365" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomy of reproductive Nereididae (Annelida) in multispecies swarms at Ambon Island, Indonesia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pamungkas, Joko</mods:namePart>
<mods:affiliation>Research Center for Deep Sea, Indonesian Institute of Sciences, Jl. Y. Syaranamual, Guru-Guru, Poka, Ambon 97233, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glasby, Christopher J.</mods:namePart>
<mods:affiliation>Museum and Art Gallery of the Northern Territory, GPO Box 4646, Darwin NT 0801, Australia</mods:affiliation>
<mods:nameIdentifier type="email">chris.glasby@nt.gov.au</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-09-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>520</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.520.9581</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.520.9581</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-520-1</mods:identifier>
<mods:identifier type="ZooBank">B96D0F7DCCC1488DBF144111C90FF9CC</mods:identifier>
<mods:identifier type="Pensoft-UUID">7B52C11DFFEEA14A16685707FF94530E</mods:identifier>
<mods:identifier type="Zenodo-Dep">579014</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182234420" LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A" httpUri="http://treatment.plazi.org/id/2AB78BC010895740B531BE7DA685FE1A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection pageId="6" pageNumber="7" type="nomenclature">
<paragraph pageId="6" pageNumber="7">
<taxonomicName LSID="2AB78BC0-1089-5740-B531-BE7DA685FE1A" authority="(Horst, 1924)" baseAuthorityName="Horst" baseAuthorityYear="1924" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes gisserana" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="gisserana">Neanthes sp. cf. Neanthes gisserana (Horst, 1924)</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nereidid epitokes, preserved specimens, dorsal view. A Neanthes unifasciata, male B Neanthes unifasciata, female C Neanthes sp. cf. Neanthes gisserana male, inset showing close up of pygidial rosette D Neanthes sp. cf. Neanthes gisserana female. mdc = modified dorsal cirri. Scales bars: 2 mm (A, B), 3 mm (C), 4 mm (D)." pageId="6" pageNumber="7">Fig. 4C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="reference_group">
<paragraph pageId="6" pageNumber="7">
<taxonomicName baseAuthorityName="Horst" baseAuthorityYear="1924" class="Polychaeta" family="Nereididae" genus="Nereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nereis (Lycoris) gisserana" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="gisserana" subGenus="Lycoris">Nereis (Lycoris) gisserana</taxonomicName>
Horst, 1924: 151-152, pl. 30, figs 6, 7.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph pageId="6" pageNumber="7">Material examined.</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<specimenCount type="male">5 males</specimenCount>
(RCLA.Ann.031),
<materialsCitation collectingDate="2014-03-18" collectorName="R. Alik" country="Ambon Island" latitude="-3.6272779" location="Ann." longLatPrecision="1" longitude="128.30695" municipality="Suli" specimenCount="3" specimenCount-female="3">
<specimenCount type="female">3 females</specimenCount>
(RCLA.
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:19590B851D50C7AA5EBF660D8965BF06" country="Ambon Island" latitude="-3.6272779" longLatPrecision="1" longitude="128.30695" municipality="Suli" name="Ann.">Ann.</location>
032),
<collectingMunicipality>Suli</collectingMunicipality>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<geoCoordinate degrees="3" direction="south" minutes="37" orientation="latitude" precision="1" seconds="38.2" value="-3.6272779">3°37'38.2&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="18" orientation="longitude" precision="1" seconds="25.0" value="128.30695">128°18'25.0&quot;E</geoCoordinate>
, coll.
<collectorName>R. Alik</collectorName>
,
<collectingDate value="2014-03-18">18 March 2014</collectingDate>
</materialsCitation>
;
<specimenCount type="male">
<pageBreakToken pageId="7" pageNumber="8" start="start">1</pageBreakToken>
male
</specimenCount>
(RCLA.Ann.033),
<materialsCitation collectingDate="2014-03-19" collectorName="R. Alik" country="Ambon Island" latitude="-3.6272779" location="Ann." longLatPrecision="1" longitude="128.30695" municipality="Suli" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
(RCLA.
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:0DAD8B9DABFF0C86FCAA4945F4E0EEB2" country="Ambon Island" latitude="-3.6272779" longLatPrecision="1" longitude="128.30695" municipality="Suli" name="Ann.">Ann.</location>
034),
<collectingMunicipality>Suli</collectingMunicipality>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<geoCoordinate degrees="3" direction="south" minutes="37" orientation="latitude" precision="1" seconds="38.2" value="-3.6272779">3°37'38.2&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="18" orientation="longitude" precision="1" seconds="25.0" value="128.30695">128°18'25.0&quot;E</geoCoordinate>
, coll.
<collectorName>R. Alik</collectorName>
,
<collectingDate value="2014-03-19">19 March 2014</collectingDate>
</materialsCitation>
;
<specimenCount type="male">3 males</specimenCount>
(RCLA.Ann.035),
<materialsCitation collectingDate="2014-03-18" collectorName="A. S. Leatemia" country="Ambon Island" latitude="-3.7451668" location="Ann." longLatPrecision="1" longitude="128.19017" municipality="Mahia" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2 females</specimenCount>
(RCLA.
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:8423CEFE4842876BDFAB55FFBE7AF3B6" country="Ambon Island" latitude="-3.7451668" longLatPrecision="1" longitude="128.19017" municipality="Mahia" name="Ann.">Ann.</location>
036),
<collectingMunicipality>Mahia</collectingMunicipality>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<geoCoordinate degrees="3" direction="south" minutes="44" orientation="latitude" precision="1" seconds="42.6" value="-3.7451668">3°44'42.6&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.6" value="128.19017">128°11'24.6&quot;E</geoCoordinate>
, coll.
<collectorName>A.S. Leatemia</collectorName>
,
<collectingDate value="2014-03-18">18 March 2014</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="2014-03-19" collectorName="A. S. Leatemia" country="Ambon Island" latitude="-3.7451668" location="Ann." longLatPrecision="1" longitude="128.19017" municipality="Mahia" specimenCount="2" specimenCount-male="2">
<specimenCount type="male">2 males</specimenCount>
(RCLA.
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:499CCE2B757C3F49DDB9C3EFF412C44F" country="Ambon Island" latitude="-3.7451668" longLatPrecision="1" longitude="128.19017" municipality="Mahia" name="Ann.">Ann.</location>
037),
<collectingMunicipality>Mahia</collectingMunicipality>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<geoCoordinate degrees="3" direction="south" minutes="44" orientation="latitude" precision="1" seconds="42.6" value="-3.7451668">3°44'42.6&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.6" value="128.19017">128°11'24.6&quot;E</geoCoordinate>
, coll.
<collectorName>A.S. Leatemia</collectorName>
,
<collectingDate value="2014-03-19">19 March 2014</collectingDate>
</materialsCitation>
;
<specimenCount type="male">2 males</specimenCount>
(MZB.Pol.00167),
<materialsCitation collectingDate="2014-03-18" collectorName="R. Alik" country="Ambon Island" latitude="-3.6272779" location="Pol." longLatPrecision="1" longitude="128.30695" municipality="Suli" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2 females</specimenCount>
(MZB.
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:6E975DBD5C9F82E736122C051A74907F" country="Ambon Island" latitude="-3.6272779" longLatPrecision="1" longitude="128.30695" municipality="Suli" name="Pol.">Pol.</location>
00168),
<collectingMunicipality>Suli</collectingMunicipality>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<geoCoordinate degrees="3" direction="south" minutes="37" orientation="latitude" precision="1" seconds="38.2" value="-3.6272779">3°37'38.2&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="18" orientation="longitude" precision="1" seconds="25.0" value="128.30695">128°18'25.0&quot;E</geoCoordinate>
, coll.
<collectorName>R. Alik</collectorName>
,
<collectingDate value="2014-03-18">18 March 2014</collectingDate>
</materialsCitation>
;
<materialsCitation accessionNumber="W23797" collectingDate="2009-03-14" collectorName="J. Pamungkas" country="Ambon Island" latitude="-3.7717223" location="Alang" longLatPrecision="1" longitude="128.00684" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1 male</specimenCount>
(NTM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/W23797">W23797</accessionNumber>
),
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:F27D5A4F46B55F3CF915178708AF03F1" country="Ambon Island" latitude="-3.7717223" longLatPrecision="1" longitude="128.00684" name="Alang">Alang</location>
,
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
<geoCoordinate degrees="3" direction="south" minutes="46" orientation="latitude" precision="1" seconds="18.2" value="-3.7717223">3°46'18.2&quot;S</geoCoordinate>
,
<geoCoordinate degrees="128" direction="east" minutes="00" orientation="longitude" precision="1" seconds="24.6" value="128.00684">128°00'24.6&quot;E</geoCoordinate>
, coll.
<collectorName>J. Pamungkas</collectorName>
,
<collectingDate value="2009-03-14">14 March 2009</collectingDate>
</materialsCitation>
;
<materialsCitation accessionNumber="W23804" collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectorName="D. S. Hoedt" country="Ambon Island" county="Fig." location="Naturalis" municipality="Leiden" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
(specimen extracted from a jar of many hundred worms in the collection of
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:73CA202A0812087D141168C8D47EEB79" country="Ambon Island" county="Fig." municipality="Leiden" name="Naturalis">Naturalis</location>
,
<collectingMunicipality>Leiden</collectingMunicipality>
);
<collectingCounty>Fig.</collectingCounty>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sample of mixed-species nereidid epitokes collected Dutch biologist D. S. Hoedt in 1866, as found in Naturalis, Leiden." pageId="7" pageNumber="8">2</figureCitation>
(NTM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/W23804">W23804</accessionNumber>
),
<collectingCountry name="Ambon Island">Ambon Island</collectingCountry>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
, coll.
<collectorName>D.S. Hoedt</collectorName>
</materialsCitation>
, 1866.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="materials_examined">
<paragraph pageId="7" pageNumber="8">Comparative material.</paragraph>
<paragraph pageId="7" pageNumber="8">
<taxonomicName baseAuthorityName="Horst" baseAuthorityYear="1924" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes gisserana" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="gisserana">
<emphasis italics="true" pageId="7" pageNumber="8">Neanthes gisserana</emphasis>
</taxonomicName>
(Horst, 1924).
<materialsCitation collectingDate="1899-08-26" country="Indonesia" elevation="18" location="Gisser" municipality="Seram-Laut" specimenCount="2" stateProvince="Maluku" typeStatus="Syntypes">
<typeStatus>Syntypes</typeStatus>
<specimenCount type="generic">2 ex.</specimenCount>
(ZMA VPol 0854), Siboga Stn. 172,
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:95160C9C34C4D47C07CD4E94D9172998" country="Indonesia" municipality="Seram-Laut" name="Gisser" stateProvince="Maluku">Gisser</location>
anchorage, between this island and
<collectingMunicipality>Seram-Laut</collectingMunicipality>
,
<collectingRegion country="Indonesia" name="Maluku Utara">Maluku</collectingRegion>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="1.8" unit="m" value="18.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="1.8" unit="m" value="18.0">18 m</elevation>
</quantity>
, coll.
<collectingDate value="1899-08-26">26 August 1899</collectingDate>
.
<taxonomicName baseAuthorityName="Horst" baseAuthorityYear="1924" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes gisserana" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="gisserana">Neanthes cf gisserana</taxonomicName>
</materialsCitation>
.
<materialsCitation accessionNumber="W22501" collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" country="Australia" location="Lizard Island" municipality="northern Great Barrier Reef" specimenCount="1" typeStatus="syntype">
<specimenCount type="generic">1 ex.</specimenCount>
(NTM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/W22501">W22501</accessionNumber>
),
<location LSID="urn:lsid:plazi:treatment:2AB78BC010895740B531BE7DA685FE1A:72604FE84D6CEE39C3C07E06FFBCF741" country="Australia" municipality="northern Great Barrier Reef" name="Lizard Island">Lizard Island</location>
,
<collectingMunicipality>northern Great Barrier Reef</collectingMunicipality>
,
<collectingCountry name="Australia">Australia</collectingCountry>
, coll. CREEFS surveys 2008-2010
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="size range">
<paragraph pageId="7" pageNumber="8">Size range.</paragraph>
<paragraph pageId="7" pageNumber="8">Male: length (16-25 mm), maximum width (2.0-4.0 mm). Female: length (16-32 mm), maximum width (3.0-4.0 mm).</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="diagnosis">
<paragraph pageId="7" pageNumber="8">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="8">Neanthes</emphasis>
</taxonomicName>
species having brown band on dorsal surface of chaetiger 4 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nereidid epitokes, preserved specimens, dorsal view. A Neanthes unifasciata, male B Neanthes unifasciata, female C Neanthes sp. cf. Neanthes gisserana male, inset showing close up of pygidial rosette D Neanthes sp. cf. Neanthes gisserana female. mdc = modified dorsal cirri. Scales bars: 2 mm (A, B), 3 mm (C), 4 mm (D)." pageId="7" pageNumber="8">4C, D</figureCitation>
). Paragnaths comprise cones and bars arranged as follows: Areas I: 4 (in a longitudinal line); II: 14; III: 19-22 (including lateral groups); IV: 26 (includes cones and bars); V: 0; VI: 3-6; VII-VIII: 5-7 in transverse line. Male and female epitokes with similar body pigmentation and natatory region extending to pygidium, but differ in number of pre-natatory chaetigers and number modified dorsal and ventral cirri. Male having pre-natatory region comprising 21-22 chaetigers; basally swollen dorsal cirri on chaetigers 1 to 7; basally swollen ventral cirri on chaetigers 1 to 5 or 6. Female having pre-natatory region comprising 26 chaetigers; basally swollen dorsal cirri on chaetigers 1 to 6; basally swollen ventral cirri on chaetigers 1 to 5. Also, male has pygidial rosette (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nereidid epitokes, preserved specimens, dorsal view. A Neanthes unifasciata, male B Neanthes unifasciata, female C Neanthes sp. cf. Neanthes gisserana male, inset showing close up of pygidial rosette D Neanthes sp. cf. Neanthes gisserana female. mdc = modified dorsal cirri. Scales bars: 2 mm (A, B), 3 mm (C), 4 mm (D)." pageId="7" pageNumber="8">4C</figureCitation>
) and sub-distally swollen (not scalloped) dorsal cirri, lacking in female.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="remarks">
<paragraph pageId="7" pageNumber="8">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="8">
The specimens examined in this study were compared with Horst's, syntypes, which they resemble closely, especially in the paragnath count and pattern; however, the present specimens are much larger than those of
<normalizedToken originalValue="Horsts">Horst's</normalizedToken>
, the modified chaetigers start later in the present specimens, and Horst does not mention the presence of a brown band which we observed on chaetiger 4. Therefore our material possibly represents a new species; it is very similar to specimens also referred to as
<taxonomicName class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="8">Neanthes</emphasis>
</taxonomicName>
sp. cf
<taxonomicName baseAuthorityName="Horst" baseAuthorityYear="1924" class="Polychaeta" family="Nereididae" genus="Neanthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanthes gisserana" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="gisserana">
<emphasis italics="true" pageId="7" pageNumber="8">Neanthes gisserana</emphasis>
</taxonomicName>
by
<bibRefCitation author="Glasby, CJ" journalOrPublisher="Bulletin of the British Museum (Natural History, Zoology)" pageId="18" pageNumber="19" refId="B9" refString="Glasby, CJ, 2015. Nereididae (Annelida: Phyllodocida) from Lizard Island, northern Great Barrier Reef, Australia. In: Zootaxa." title="Nereididae (Annelida: Phyllodocida) from Lizard Island, northern Great Barrier Reef, Australia." volumeTitle="Zootaxa." year="2015">Glasby (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>