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<document id="31AA7091D66A1E0B19E62D4FBFEA68B3" ID-DOI="10.5281/zenodo.178251" ID-GBIF-Dataset="905dea16-dd69-45f9-9fe2-5b1aa6af93b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="178251" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459886785444" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Vilkamaa, Pekka" docDate="2007" docId="03DB45439C7BFFC043DDF9D2FAF9FC2B" docLanguage="en" docName="zt01556p050.pdf" docOrigin="Zootaxa 1556" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Keilbachia flabellata Hippa &amp; Vilkamaa, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFE23D3B9C70FFCC434AFFE0FFC7FFA9" masterDocTitle="The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar" masterLastPageNumber="50" masterPageNumber="31" pageNumber="42" updateTime="1698227854892" updateUser="plazi">
<mods:mods id="28D38CE7164F946F4F9E4DC5F8398F79" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="823BDE94D2C82E76B32919A2AFADE009">
<mods:title id="3468313CBC3E9457077522E7FA0CD738">The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar</mods:title>
</mods:titleInfo>
<mods:name id="F772D9B9E38F74DADE44BA7E680CA935" type="personal">
<mods:role id="A0BD5E77EC16E4B983FBC02EDF5692B1">
<mods:roleTerm id="D84A90D60F8C2204415E1889BBCF557C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61BC3094CC67B7FF75182110B3650129">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:name id="9C7282ACC29B6A10C3E98F0214F5F663" type="personal">
<mods:role id="2F455D1A15F6BEA3C3CB817EC20E6A05">
<mods:roleTerm id="0DCD0978A86E7D28137E33F0B29D8B34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2640086EA21EF483C531A9FAE8CDD5F7">Vilkamaa, Pekka</mods:namePart>
</mods:name>
<mods:typeOfResource id="104A304D07C969A36052E24E7AA50A1A">text</mods:typeOfResource>
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<mods:titleInfo id="4D0ECB27A332088D5E406F40B40335EC">
<mods:title id="EA59644FC281C460CAF63ACC842CBD86">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3DF20AD16410558DD762D34C3EC6B194">
<mods:date id="956D45B64520873BE4A100961B49A3B7">2007</mods:date>
<mods:detail id="565B0586554AD3325D4350921EDA021F" type="volume">
<mods:number id="B6C387244D904BE53137A80F6A306283">1556</mods:number>
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<mods:classification id="12E9D396B51676D2A45C992375DBC827">journal article</mods:classification>
<mods:identifier id="0FF8E666E161EA8E8F699885A10ACCA5" type="DOI">10.5281/zenodo.178251</mods:identifier>
<mods:identifier id="F4354F1C408A818DBE97B9B214006B41" type="GBIF-Dataset">905dea16-dd69-45f9-9fe2-5b1aa6af93b6</mods:identifier>
<mods:identifier id="CA1B8546E6AF0F0ABA505735BDABDDAF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1840E55630958F008159A4B0010C68F5" type="Zenodo-Dep">178251</mods:identifier>
</mods:mods>
<treatment id="03DB45439C7BFFC043DDF9D2FAF9FC2B" ID-DOI="http://doi.org/10.5281/zenodo.6241298" ID-GBIF-Taxon="119371646" ID-Zenodo-Dep="6241298" LSID="urn:lsid:plazi:treatment:03DB45439C7BFFC043DDF9D2FAF9FC2B" httpUri="http://treatment.plazi.org/id/03DB45439C7BFFC043DDF9D2FAF9FC2B" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42">
<subSubSection id="C368A7DE9C7BFFC743DDF9D2FE79F9C6" pageId="11" pageNumber="42" type="nomenclature">
<paragraph id="8BCDF4559C7BFFC743DDF9D2FE1CF9E5" blockId="11.[151,475,1586,1647]" box="[151,475,1586,1612]" pageId="11" pageNumber="42">
<heading id="D08543399C7BFFC743DDF9D2FE1CF9E5" bold="true" box="[151,475,1586,1612]" fontSize="11" level="1" pageId="11" pageNumber="42" reason="1">
<emphasis id="B90628479C7BFFC743DDF9D2FE1CF9E5" bold="true" box="[151,475,1586,1612]" pageId="11" pageNumber="42">
<taxonomicName id="4C728FD69C7BFFC743DDF9D2FE55F9E5" ID-CoL="3R4LK" box="[151,402,1586,1612]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="flabellata" status="sp. nov.">
<emphasis id="B90628479C7BFFC743DDF9D2FE55F9E5" bold="true" box="[151,402,1586,1612]" italics="true" pageId="11" pageNumber="42">Keilbachia flabellata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235953C9C7BFFC742D3F9D2FE1CF9E5" box="[409,475,1586,1612]" pageId="11" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCDF4559C7BFFC743DDF9B5FE79F9C6" blockId="11.[151,475,1586,1647]" box="[151,446,1621,1647]" pageId="11" pageNumber="42">
(
<figureCitation id="1349E8D09C7BFFC743D5F9B5FF21F9C6" box="[159,230,1621,1647]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="11" pageNumber="42">Figs 1</figureCitation>
B, 3E, 4A, 7A–D)
</paragraph>
</subSubSection>
<subSubSection id="C368A7DE9C7BFFC043DDF945FE5EFDC3" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42" type="materials_examined">
<paragraph id="8BCDF4559C7BFFC743DDF945FB6BF94E" blockId="11.[151,1436,1701,2007]" pageId="11" pageNumber="42">
<emphasis id="B90628479C7BFFC743DDF945FEAAF916" bold="true" box="[151,365,1701,1727]" pageId="11" pageNumber="42">Material studied.</emphasis>
<emphasis id="B90628479C7BFFC7423FF945FDE6F917" box="[373,545,1701,1726]" italics="true" pageId="11" pageNumber="42">
<typeStatus id="54C94AF79C7BFFC7423FF945FE27F917" box="[373,480,1701,1726]" pageId="11" pageNumber="42" type="holotype">Holotype</typeStatus>
male
</emphasis>
(in
<collectionCode id="ED636C909C7BFFC7411BF945FD6EF916" box="[593,681,1701,1727]" pageId="11" pageNumber="42">NRMS</collectionCode>
). N.E.
<collectingCountry id="F365B4C59C7BFFC741B0F945FC8DF916" box="[762,842,1701,1727]" name="Myanmar" pageId="11" pageNumber="42">Burma</collectingCountry>
(
<collectingCountry id="F365B4C59C7BFFC74013F945FC16F916" box="[857,977,1701,1727]" name="Myanmar" pageId="11" pageNumber="42">Myanmar</collectingCountry>
), Kambaiti,
<quantity id="4C8A59B09C7BFFC7472BF945FB7DF916" box="[1121,1210,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="11" pageNumber="42" unit="ft" value="7000.0">7000 ft</quantity>
.,
<date id="FFCCD2959C7BFFC74781F945FAF3F916" box="[1227,1332,1701,1727]" pageId="11" pageNumber="42" value="1934-06-08">8.6.1934</date>
, R. Malaise.
<emphasis id="B90628479C7BFFC74399F92DFE8DF94F" box="[211,330,1741,1766]" italics="true" pageId="11" pageNumber="42">
<typeStatus id="54C94AF79C7BFFC74399F92DFE8DF94F" box="[211,330,1741,1766]" pageId="11" pageNumber="42" type="paratype">Paratypes</typeStatus>
</emphasis>
(in
<collectionCode id="ED636C909C7BFFC74233F92DFE16F94E" box="[377,465,1741,1767]" pageId="11" pageNumber="42">NRMS</collectionCode>
).
<specimenCount id="9D743FDC9C7BFFC742AEF92DFDF9F94E" box="[484,574,1741,1767]" pageId="11" pageNumber="42" type="male">2 males</specimenCount>
with same data as
<typeStatus id="54C94AF79C7BFFC74056F92DFC44F94E" box="[796,899,1741,1767]" pageId="11" pageNumber="42" type="holotype">holotype</typeStatus>
but dated 25.5. and 26.5.
</paragraph>
<paragraph id="8BCDF4559C7BFFC0438CF915FEFFFDEB" blockId="11.[151,1436,1701,2007]" lastBlockId="12.[151,1436,152,898]" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42">
Male.
<emphasis id="B90628479C7BFFC7425FF915FEACF8A6" bold="true" box="[277,363,1781,1807]" pageId="11" pageNumber="42">Colour</emphasis>
. The specimen is strongly faded, almost unicolorous pale yellowish-brown.
<emphasis id="B90628479C7BFFC74652F915FA9CF8A6" bold="true" box="[1304,1371,1781,1807]" pageId="11" pageNumber="42">Head</emphasis>
. Eye bridge 4–5 facets wide. Face (prefrons) with 1–3 setae. Clypeus non-setose. Maxillary palpus (
<figureCitation id="1349E8D09C7BFFC7464EF8FDFA87F89E" box="[1284,1344,1821,1847]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="11" pageNumber="42">Fig.1</figureCitation>
B) with one palpomere, with one lateral seta, hyaline sensilla widely scattered on dorsal surface, no sensory pit. Antennal flagellomere 4 ca. one-and-a-half times as long as broad, rather deformed in both specimens and not drawn.
<emphasis id="B90628479C7BFFC743A5F875FE8BF806" bold="true" box="[239,332,1941,1967]" pageId="11" pageNumber="42">Thorax</emphasis>
. Anterior pronotum and episternum 1 with 1–2 setae each in the
<typeStatus id="54C94AF79C7BFFC74707F875FB70F806" box="[1101,1207,1941,1967]" pageId="11" pageNumber="42" type="holotype">holotype</typeStatus>
, not possible to see in the
<typeStatus id="54C94AF79C7BFFC743AAF85DFE93F87E" box="[224,340,1981,2007]" pageId="11" pageNumber="42" type="paratype">paratypes</typeStatus>
.
<emphasis id="B90628479C7BFFC74214F85DFE66F87E" bold="true" box="[350,417,1981,2007]" pageId="11" pageNumber="42">Wing</emphasis>
,
<figureCitation id="1349E8D09C7BFFC742FBF85DFE31F87E" box="[433,502,1981,2007]" captionStart="FIGURE 4" captionStartId="6.[151,255,1974,1998]" captionTargetBox="[302,1325,197,1973]" captionTargetId="figure@6.[271,1328,160,2005]" captionTargetPageId="6" captionText="FIGURE 4. Wing, dorsal view. A, Keilbachia flabellata sp. n. (holotype). B, K. foveolata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. rima sp. n. (paratype). Scale 1.0 mm." httpUri="https://zenodo.org/record/178255/files/figure.png" pageId="11" pageNumber="42">Fig. 4</figureCitation>
A. Both wings lost in one of the
<typeStatus id="54C94AF79C7BFFC7403BF85DFC22F87E" box="[881,997,1981,2007]" pageId="11" pageNumber="42" type="paratype">paratypes</typeStatus>
. Length
<quantity id="4C8A59B09C7BFFC74700F85DFB30F87E" box="[1098,1271,1981,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" metricValueMax="1.95" metricValueMin="1.67" pageId="11" pageNumber="42" unit="mm" value="1.81" valueMax="1.95" valueMin="1.67">1.67–1.95 mm</quantity>
. Width/length 0.42–0.48. R1/R 1.00. The meeting point of R1 and C in basal half of wing not as in the other species. c/
<geoCoordinate id="EE4692929C7CFFC046CDFF78FF0BFF73" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="555" value="-0.8">w 0.80</geoCoordinate>
–0.87. r-m shorter than bM, r-m/bM ca. 0.81–0.87, r-m with 1–2 setae, bM non-setose.
<emphasis id="B90628479C7CFFC047F5FF5FFB3DFF70" bold="true" box="[1215,1274,191,217]" pageId="12" pageNumber="43">Legs</emphasis>
. Apex of fore tibia,
<figureCitation id="1349E8D09C7CFFC04391FF08FED8FEAB" box="[219,287,232,258]" captionStart="FIGURE 3" captionStartId="5.[151,255,1905,1929]" captionTargetBox="[188,1393,195,1864]" captionTargetId="figure@5.[163,1428,158,1890]" captionTargetPageId="5" captionText="FIGURE 3. Apical part of fore tibia, prolateral view. A, Keilbachia oligonema sp. n. (Burma). B, K. fasciata sp. n. (paratype). C, K. rima sp. n. (holotype). D, K. acamptochaeta sp. n. (paratype). E, K. flabellata sp. n. (holotype). F, K. foveolata sp. n. (holotype). G, K. filigera sp. n. (holotype). H, K. bicuspis sp. n. (holotype). I, K. gigas sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178254/files/figure.png" pageId="12" pageNumber="43">Fig. 3</figureCitation>
E. Length of fore basitarsomere/length of fore tibia 0.72–0.78. Length of fore tibia/length of fore femur 1.56–1.58. Length of hind tibia/length of scutum+scutellum 1.95–2.16.
<emphasis id="B90628479C7CFFC04773FEEFFB74FE80" bold="true" box="[1081,1203,271,297]" pageId="12" pageNumber="43">Abdomen</emphasis>
. Sternite
<quantity id="4C8A59B09C7CFFC0466DFEF0FA92FE83" box="[1319,1365,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="12" pageNumber="43" unit="in" value="8.0">8 in</quantity>
<typeStatus id="54C94AF79C7CFFC04614FEF0FF0DFEFB" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
with 10 setae, in a transverse posterior row, with one of the setae near the lateral margin placed more anteriorly, one of the
<typeStatus id="54C94AF79C7CFFC042D0FE80FDCDFED3" box="[410,522,352,378]" pageId="12" pageNumber="43" type="paratype">paratypes</typeStatus>
apparently similar but the exact number difficult see and the other
<typeStatus id="54C94AF79C7CFFC0467DFE80FA5BFED3" box="[1335,1436,352,378]" pageId="12" pageNumber="43" type="paratype">paratype</typeStatus>
with 13 setae in similar pattern but with three setae placed medially in a more anterior position.
<emphasis id="B90628479C7CFFC047B8FE67FA5BFE08" bold="true" box="[1266,1436,391,417]" pageId="12" pageNumber="43">Hypopygium.</emphasis>
<figureCitation id="1349E8D09C7CFFC043DDFE50FF26FE63" box="[151,225,432,458]" captionStart="FIGURE 7" captionStartId="10.[151,255,1818,1842]" captionTargetBox="[166,1425,193,1794]" captionTargetId="figure@10.[151,1436,176,1794]" captionTargetPageId="10" captionText="FIGURE 7. Keilbachia flabellata sp. n. (A, B, and C paratypes, D holotype). A, C, part of hypopygium, ventral view. B, D, gonostylus, ventral view. Scale 0.10 mm." httpUri="https://zenodo.org/record/178258/files/figure.png" pageId="12" pageNumber="43">Fig. 7</figureCitation>
: Gonostylus with 5 long mesial megasetae arising from a common basal body, the megasetae curved except on one side in the
<typeStatus id="54C94AF79C7CFFC04280FE38FDF7FE5B" box="[458,560,472,498]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
where they are unusually straight, with no additional megasetae but with two stronger setae diverging from other nearby setae near the base of the apical tooth. Tegmen rather simple, subtriangular.
</paragraph>
<paragraph id="8BCDF4559C7CFFC0438CFDB0FE5EFDC3" blockId="12.[151,1436,152,898]" box="[198,409,592,618]" pageId="12" pageNumber="43">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C368A7DE9C7CFFC0438CFD97FAF9FC2B" pageId="12" pageNumber="43" type="discussion">
<paragraph id="8BCDF4559C7CFFC0438CFD97FAF9FC2B" blockId="12.[151,1436,152,898]" pageId="12" pageNumber="43">
<emphasis id="B90628479C7CFFC0438CFD97FE95FD38" bold="true" box="[198,338,631,657]" pageId="12" pageNumber="43">Discussion.</emphasis>
<taxonomicName id="4C728FD69C7CFFC04210FD98FD93FD38" box="[346,596,632,657]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis id="B90628479C7CFFC04210FD98FD93FD38" box="[346,596,632,657]" italics="true" pageId="12" pageNumber="43">Keilbachia flabellata</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C728FD69C7CFFC041B9FD98FC0EFD38" box="[755,969,632,657]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="acamptochaeta">
<emphasis id="B90628479C7CFFC041B9FD98FC0EFD38" box="[755,969,632,657]" italics="true" pageId="12" pageNumber="43">K. acamptochaeta</emphasis>
</taxonomicName>
in having a completely one-segmented maxillary palpus, but otherwise the species are not especially similar.
<taxonomicName id="4C728FD69C7CFFC040ACFD40FBBCFD10" box="[998,1147,672,697]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis id="B90628479C7CFFC040ACFD40FBBCFD10" box="[998,1147,672,697]" italics="true" pageId="12" pageNumber="43">K. flabellata</emphasis>
</taxonomicName>
is distinguished e.g. by having in the gonostylus five long curved megasetae instead of two short ones and by lacking strong megasetae between the curved ones and the apical tooth. The gonostylus of
<taxonomicName id="4C728FD69C7CFFC040FAFD10FB84FCA0" box="[944,1091,752,777]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis id="B90628479C7CFFC040FAFD10FB84FCA0" box="[944,1091,752,777]" italics="true" pageId="12" pageNumber="43">K. flabellata</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C728FD69C7CFFC04661FD10FA5BFCA0" box="[1323,1436,752,777]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flagria">
<emphasis id="B90628479C7CFFC04661FD10FA5BFCA0" box="[1323,1436,752,777]" italics="true" pageId="12" pageNumber="43">K. flagria</emphasis>
</taxonomicName>
from
<collectingCountry id="F365B4C59C7CFFC0439DFCF8FEDAFC9B" box="[215,285,792,818]" name="Nepal" pageId="12" pageNumber="43">Nepal</collectingCountry>
and there are no clear distinguishing characters. The tegmen of both species is also similar except that it is relatively longer in
<taxonomicName id="4C728FD69C7CFFC042BFFCA0FDADFCF0" box="[501,618,832,857]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flagria">
<emphasis id="B90628479C7CFFC042BFFCA0FDADFCF0" box="[501,618,832,857]" italics="true" pageId="12" pageNumber="43">K. flagria</emphasis>
</taxonomicName>
.
<taxonomicName id="4C728FD69C7CFFC04136FCA0FD36FCF0" box="[636,753,832,857]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="flagria">
<emphasis id="B90628479C7CFFC04136FCA0FD36FCF0" box="[636,753,832,857]" italics="true" pageId="12" pageNumber="43">K. flagria</emphasis>
</taxonomicName>
differs by its three-segmented palpus and by having the whole length of r-m–bM setose instead of being non-setose but with a couple of setae on r-m near Rs.
</paragraph>
</subSubSection>
</treatment>
</document>
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