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<document id="5F690043A1B99043F4A94119C96D7F4B" ID-DOI="10.11865/zs.201718" ID-ISSN="2095-6827" ID-Zenodo-Dep="5366783" ID-ZooBank="25D9FD67-3B6B-4F95-88D1-190FDD5C2E54" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630257354491" checkinUser="diego" docAuthor="Niu, Zeqing, Yuan, Feng &amp; Zhu, Chaodong" docDate="2017" docId="03C38791FFCCCC248CD9FE8273EAFECB" docLanguage="en" docName="Taxonomic study of the genus Tetralonioidella Strand from China (Hymenoptera_ Apidae_ Melectini).pdf" docOrigin="Zoological Systematics 42 (4)" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Tetralonioidella maniwengensis Niu &amp; Zhu 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="439" masterDocId="FFFAFFE9FFDFCC318C51FFD97754FFA7" masterDocTitle="Taxonomic study of the genus Tetralonioidella Strand from China (Hymenoptera: Apidae: Melectini)" masterLastPageNumber="445" masterPageNumber="418" pageNumber="437" updateTime="1699199799450" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C991917A1B7E3732E0F4868B4C730F06">Taxonomic study of the genus Tetralonioidella Strand from China (Hymenoptera: Apidae: Melectini)</mods:title>
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<mods:namePart id="D17F8EA8BAEA50CA4442876F87E15900">Niu, Zeqing</mods:namePart>
<mods:affiliation id="133C3BFD8FE306D8FC3E6059F17DEFFA">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China; E-mails: niuzq @ ioz. ac. cn; yuanfeng @ ioz. ac. cn</mods:affiliation>
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<mods:namePart id="C50B7AF08FD7C1C59FD5633C0DD76B7A">Yuan, Feng</mods:namePart>
<mods:affiliation id="6C6DDCEA4C64A9A312FD02D83C4D50CA">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China; E-mails: niuzq @ ioz. ac. cn; yuanfeng @ ioz. ac. cn</mods:affiliation>
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<mods:namePart id="0F831F422C96F229D72BEB15F72215A9">Zhu, Chaodong</mods:namePart>
<mods:affiliation id="F49C0FE5099FDD626147591905877399">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China; E-mails: niuzq @ ioz. ac. cn; yuanfeng @ ioz. ac. cn &amp; University of Chinese Academy of Sciences, Beijing 100049, P. R. China Corresponding author, E-mail: zhucd @ ioz. ac. cn</mods:affiliation>
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<subSubSection id="C370650CFFCCCC228CD9FE82743FFED4" box="[136,875,347,371]" pageId="19" pageNumber="437" type="nomenclature">
<paragraph id="8BD53687FFCCCC228CD9FE82743FFED4" blockId="19.[136,875,347,371]" box="[136,875,347,371]" pageId="19" pageNumber="437">
<heading id="D09D81EBFFCCCC228CD9FE82743FFED4" box="[136,875,347,371]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="437" reason="2">
<emphasis id="B91EEA95FFCCCC228CD9FE827598FED4" bold="true" box="[136,716,347,371]" pageId="19" pageNumber="437">
<taxonomicName id="4C6A4D04FFCCCC228CD9FE82753EFED4" ID-CoL="7BSRV" authority="Niu &amp; Zhu" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[136,618,347,371]" class="Insecta" family="Apidae" genus="Tetralonioidella" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="437" phylum="Arthropoda" rank="species" species="maniwengensis" status="sp. nov.">
<emphasis id="B91EEA95FFCCCC228CD9FE8276B2FED4" bold="true" box="[136,486,347,371]" italics="true" pageId="19" pageNumber="437">Tetralonioidella maniwengensis</emphasis>
Niu &amp; Zhu
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, 
<taxonomicNameLabel id="A22D57EEFFCCCC228E27FE827598FED4" box="[630,716,347,371]" pageId="19" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13512A02FFCCCC228EBBFE827430FED4" box="[746,868,347,371]" captionStart-0="Figure 16" captionStart-1="Figure 17" captionStartId-0="19.[154,217,2025,2047]" captionStartId-1="20.[153,216,1995,2017]" captionTargetBox-0="[188,1398,766,1976]" captionTargetBox-1="[187,1397,736,1946]" captionTargetId-0="figure-270@19.[188,1398,766,1976]" captionTargetId-1="figure-285@20.[187,1397,736,1946]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." captionText-1="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi-0="http://doi.org/10.5281/zenodo.5366815" figureDoi-1="http://doi.org/10.5281/zenodo.5366817" httpUri-0="https://zenodo.org/record/5366815/files/figure.png" httpUri-1="https://zenodo.org/record/5366817/files/figure.png" pageId="19" pageNumber="437">Figs 16–17</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C370650CFFCCCC248CECFE4A73EAFECB" lastPageId="21" lastPageNumber="439" pageId="19" pageNumber="437" type="diagnosis">
<paragraph id="8BD53687FFCCCC228CECFE4A75F1FE6A" blockId="19.[136,1452,403,704]" pageId="19" pageNumber="437">
Diagnosis. Apical margin of labrum slightly concave, similar to male of 
<taxonomicName id="4C6A4D04FFCCCC228FECFE4D731BFE0C" authorityName="Lieftinck" authorityYear="1983" box="[957,1103,403,427]" class="Insecta" family="Apidae" genus="Tetralonioidella" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="437" phylum="Arthropoda" rank="species" species="fukienensis">
<emphasis id="B91EEA95FFCCCC228FECFE4D7498FE0C" box="[957,972,404,427]" italics="true" pageId="19" pageNumber="437">T</emphasis>
. 
<emphasis id="B91EEA95FFCCCC228F89FE4A731BFE0C" box="[984,1103,403,427]" italics="true" pageId="19" pageNumber="437">fukienensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6A4D04FFCCCC2288D3FE4D73A9FE0C" baseAuthorityName="Lieftinck" baseAuthorityYear="1944" box="[1154,1277,403,427]" class="Insecta" family="Apidae" genus="Tetralonioidella" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="437" phylum="Arthropoda" rank="species" species="insidiosa">
<emphasis id="B91EEA95FFCCCC2288D3FE4D73C5FE0C" box="[1154,1169,404,427]" italics="true" pageId="19" pageNumber="437">T</emphasis>
. 
<emphasis id="B91EEA95FFCCCC2288CCFE4A73A9FE0C" box="[1181,1277,403,427]" italics="true" pageId="19" pageNumber="437">insidiosa</emphasis>
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, but they can be distinguished by the different S7, S8 and genitalia.
</paragraph>
<paragraph id="8BD53687FFCCCC258CECFE0172C4FD25" blockId="19.[136,1452,403,704]" lastBlockId="20.[136,1452,236,676]" lastPageId="20" lastPageNumber="438" pageId="19" pageNumber="437">
Description. Male. BL = 
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(
<figureCitation id="13512A02FFCCCC228E6DFE0175CEFE57" box="[572,666,472,496]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437">Fig. 16a</figureCitation>
); head broader than long, HW: HL = 64: 47 (
<figureCitation id="13512A02FFCCCC2288C5FE0173A0FE57" box="[1172,1268,472,496]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437">Fig. 16b</figureCitation>
); gena distinctly narrower than eye, GW: EW= 9:19; width of metasoma slightly broader than the width between the tegulae, MtW: MsW=76: 74. Clypeus broader than long (
<figureCitation id="13512A02FFCCCC228D89FDC47561FD92" box="[472,565,541,565]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437">Fig. 16b</figureCitation>
); apical margin of labrum slightly concave (
<figureCitation id="13512A02FFCCCC22885BFDC47331FD92" box="[1034,1125,541,565]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437">Fig. 16c</figureCitation>
); antenna short, reaching front margin of tegula, scape conically broadened, longer than F1 to F3 together, flagellomere equal in breadth, F1 longer than broad, nearly 1.2 times as long as broad, F2 long than broad, nearly 1.5 times as long as broad, F3 longer than broad, nearly 1.2 times as long as broad, F4, F5, F6 and F10 equal in length about 1.1 times as long as broad, F7–F9 equal in length, nearly as long as broad, F11 rounded apically, equal in length with F2, about 1.5 times as long as broad (
<figureCitation id="13512A02FFCCCC2288CCFD7173AEFD67" box="[1181,1274,680,704]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437">Fig. 16d</figureCitation>
); fore wing with distinct numerous papillae apically (
<figureCitation id="13512A02FFCBCC258E54FF35750BFEA2" box="[517,607,236,261]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17a</figureCitation>
); scutellar spines short and broad, blunt rounded apically, barely visible between pubescence; inner ramus of hind tarsal claw axe-shaped, arolium present (
<figureCitation id="13512A02FFCBCC258FC9FED674A1FE80" box="[920,1013,271,295]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16b</figureCitation>
); ventral surface of apical part of S7 with dense setae laterally, median part slightly concave, with sparse setae (
<figureCitation id="13512A02FFCBCC258FDAFEEB74BDFEED" box="[907,1001,306,330]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17c</figureCitation>
); apical margin of median process of S8 distinctly concave (
<figureCitation id="13512A02FFCBCC258D08FE8D76E1FECB" box="[345,437,340,364]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17d</figureCitation>
); genitalia as illustrated in 
<figureCitation id="13512A02FFCBCC258E86FE8D747AFECB" box="[727,814,340,364]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17e</figureCitation>
(in dorsal view) and 
<figureCitation id="13512A02FFCBCC258845FE8D733CFECB" box="[1044,1128,340,364]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17f</figureCitation>
(in lateral view), basal part of gonostylus with crescent-shaped process dorsally, and the process with dense long hairs along dorsal margin (
<figureCitation id="13512A02FFCBCC258915FEAE72F4FE28" box="[1348,1440,375,399]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17f</figureCitation>
). Clypeus black (
<figureCitation id="13512A02FFCBCC258D62FE4376C7FE15" box="[307,403,410,434]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16b</figureCitation>
); mandible reddish-brown; labrum dark yellowish-brown except apical margin black (
<figureCitation id="13512A02FFCBCC258911FE4372CAFE15" box="[1344,1438,410,434]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16c</figureCitation>
); antenna black beneath (
<figureCitation id="13512A02FFCBCC258DDBFE6576BDFE73" box="[394,489,444,468]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16d</figureCitation>
); tegula yellowish-brown (
<figureCitation id="13512A02FFCBCC258F5FFE65743FFE73" box="[782,875,444,468]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16e</figureCitation>
); all legs dark reddish-brown; hind tarsal claw black apically (
<figureCitation id="13512A02FFCBCC258CBFFE067619FE50" box="[238,333,479,503]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">Fig. 17b</figureCitation>
). Scutum pubescence yellowish (
<figureCitation id="13512A02FFCBCC258EE8FE067461FE50" box="[697,821,479,503]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Figs 16a, e</figureCitation>
); clypeus covered with white hairs, periphery of antennal socket, supraclypeal area, paraocular area and front surface of scape covered with yellowish-white hairs (
<figureCitation id="13512A02FFCBCC2588A6FDDB7206FDBD" box="[1271,1362,514,538]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16b</figureCitation>
);vertex, genal area, scutum, scutellum, metanotum and episternum covered with dense and long yellowish hairs (
<figureCitation id="13512A02FFCBCC2588ADFDFD722CFD9B" box="[1276,1400,548,572]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Figs 16a, e</figureCitation>
); T1 covered with thin and short yellowish-brown hairs (
<figureCitation id="13512A02FFCBCC258EE3FD9E745DFDF8" box="[690,777,583,607]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Fig. 16f</figureCitation>
), T2-T4with short, yellowish-white hair bands before marginal zones, and the bands gradually narrowed medially; disc of T2–T4 covered dark yellowish-brown hairs (
<figureCitation id="13512A02FFCBCC25888BFDB3721BFD25" box="[1242,1359,617,642]" captionStart="Figure 16" captionStartId="19.[154,217,2025,2047]" captionTargetBox="[188,1398,766,1976]" captionTargetId="figure-270@19.[188,1398,766,1976]" captionTargetPageId="19" captionText="Figure 16. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars=1mm." figureDoi="http://doi.org/10.5281/zenodo.5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="20" pageNumber="438">Figs 16a, f</figureCitation>
, 
<figureCitation id="13512A02FFCBCC25890BFDB372D0FD25" box="[1370,1412,618,642]" captionStart="Figure 17" captionStartId="20.[153,216,1995,2017]" captionTargetBox="[187,1397,736,1946]" captionTargetId="figure-285@20.[187,1397,736,1946]" captionTargetPageId="20" captionText="Figure 17. Tetralonioidella maniwengensis Niu &amp; Zhu, sp. nov., male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f= 1mm; b, c, d=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438">17a</figureCitation>
).
</paragraph>
<caption id="DF15660FFFCCCC228CCBF8307322F7B9" ID-DOI="http://doi.org/10.5281/zenodo.5366815" ID-Zenodo-Dep="5366815" httpUri="https://zenodo.org/record/5366815/files/figure.png" pageId="19" pageNumber="437" startId="19.[154,217,2025,2047]" targetBox="[188,1398,766,1976]" targetPageId="19">
<paragraph id="8BD53687FFCCCC228CCBF8307322F7B9" blockId="19.[154,1431,2025,2079]" pageId="19" pageNumber="437">
Figure 16. 
<taxonomicName id="4C6A4D04FFCCCC228D54F83075F8F858" authority="Niu &amp; Zhu" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[261,684,2025,2047]" class="Insecta" family="Apidae" genus="Tetralonioidella" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="437" phylum="Arthropoda" rank="species" species="maniwengensis" status="sp. nov.">
<emphasis id="B91EEA95FFCCCC228D54F830756CF858" box="[261,568,2025,2047]" italics="true" pageId="19" pageNumber="437">Tetralonioidella maniwengensis</emphasis>
Niu &amp; Zhu
</taxonomicName>
, 
<emphasis id="B91EEA95FFCCCC228EE6F8307450F859" bold="true" box="[695,772,2025,2046]" pageId="19" pageNumber="437">
<taxonomicNameLabel id="A22D57EEFFCCCC228EE6F8307450F859" box="[695,772,2025,2046]" pageId="19" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. a. Body, lateral view. b. Head, frontal view. c. Labrum, frontal view. d. Antenna. e. Mesosoma, dorsal view. f. Metasoma, dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BD53687FFCBCC258CECFD5576DEFD03" blockId="20.[136,1452,236,676]" box="[189,394,652,676]" pageId="20" pageNumber="438">Female. Unknown.</paragraph>
<caption id="DF15660FFFCBCC258CC8F8127622F786" ID-DOI="http://doi.org/10.5281/zenodo.5366817" ID-Zenodo-Dep="5366817" httpUri="https://zenodo.org/record/5366817/files/figure.png" pageId="20" pageNumber="438" startId="20.[153,216,1995,2017]" targetBox="[187,1397,736,1946]" targetPageId="20">
<paragraph id="8BD53687FFCBCC258CC8F8127622F786" blockId="20.[153,1430,1995,2081]" pageId="20" pageNumber="438">
Figure 17. 
<taxonomicName id="4C6A4D04FFCBCC258D55F81275FFF846" authority="Niu &amp; Zhu" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[260,683,1995,2017]" class="Insecta" family="Apidae" genus="Tetralonioidella" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="438" phylum="Arthropoda" rank="species" species="maniwengensis" status="sp. nov.">
<emphasis id="B91EEA95FFCBCC258D55F8127563F846" box="[260,567,1995,2017]" italics="true" pageId="20" pageNumber="438">Tetralonioidella maniwengensis</emphasis>
Niu &amp; Zhu
</taxonomicName>
, 
<emphasis id="B91EEA95FFCBCC258EE4F8127457F847" bold="true" box="[693,771,1995,2016]" pageId="20" pageNumber="438">
<taxonomicNameLabel id="A22D57EEFFCBCC258EE4F8127457F847" box="[693,771,1995,2016]" pageId="20" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. a. Fore wing, showing the papillae. b. Claw, showing the inner ramus. c. S7, ventral view. d. S8, ventral view. e. Genitalia, dorsal view. f. Genitalia, lateral view. Scale bars: a, e, f = 1 mm; b, c, d =0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD53687FFCACC248CECFF3576C4FE81" blockId="21.[136,1452,236,365]" pageId="21" pageNumber="439">
<materialsCitation id="3B023CDAFFCACC248CECFF3576C4FE81" collectingDate="1979-06-31" collectorName="Gentao Jin &amp; Jianyi Wu." country="China" county="Material" elevation="1050" latitude="28.733334" location="Maniweng" longLatPrecision="1230" longitude="95.01667" municipality="Medog" pageId="21" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<collectingCounty id="62B44E0BFFCACC248CECFF35764CFEA3" box="[189,280,236,260]" pageId="21" pageNumber="439">Material</collectingCounty>
examined. 
<typeStatus id="54D18825FFCACC248DC0FF3576ACFEA3" box="[401,504,236,260]" pageId="21" pageNumber="439" type="holotype">Holotype</typeStatus>
. 
<specimenCount id="9D6CFD0EFFCACC248E51FF297540FEA3" box="[512,532,240,260]" pageId="21" pageNumber="439" type="male">♂</specimenCount>
, 
<collectingCountry id="F37D7617FFCACC248E71FF357537FEA3" box="[544,611,236,260]" name="China" pageId="21" pageNumber="439">China</collectingCountry>
, 
<collectingRegion id="49AEF865FFCACC248E3DFF3575E8FEA3" box="[620,700,236,260]" country="China" name="Xizang" pageId="21" pageNumber="439">Xizang</collectingRegion>
, 
<collectingMunicipality id="6BB1ACFDFFCACC248E94FF357447FEA3" box="[709,787,236,260]" pageId="21" pageNumber="439">Mêdog</collectingMunicipality>
, 
<location id="8EB5605CFFCACC248F4DFF3574DAFEA3" LSID="urn:lsid:plazi:treatment:03C38791FFCCCC248CD9FE8273EAFECB:8EB5605CFFCACC248F4DFF3574DAFEA3" box="[796,910,236,260]" country="China" county="Material" latitude="28.733334" longLatPrecision="1230" longitude="95.01667" municipality="Medog" name="Maniweng" pageId="21" pageNumber="439" stateProvince="Xizang">Maniweng</location>
(
<geoCoordinate id="EE5E5040FFCACC248FCAFF3474A2FEA3" box="[923,1014,237,261]" degrees="28" direction="north" minutes="44" orientation="latitude" pageId="21" pageNumber="439" precision="925" value="28.733334">28º44′N</geoCoordinate>
, 
<geoCoordinate id="EE5E5040FFCACC248FAEFF357301FEA3" box="[1023,1109,236,260]" degrees="95" direction="east" minutes="01" orientation="longitude" pageId="21" pageNumber="439" precision="925" value="95.01667">95º01′E</geoCoordinate>
; elev. 
<quantity id="4C929B62FFCACC2488C7FF3573B2FEA2" box="[1174,1254,236,261]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="21" pageNumber="439" unit="m" value="1050.0">
<elevation id="0047D1B4FFCACC2488C7FF3573B2FEA2" box="[1174,1254,236,261]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="21" pageNumber="439" unit="m" value="1050.0">1050 m</elevation>
</quantity>
), 
<date id="FFD41047FFCACC2488A6FF347227FEA2" box="[1271,1395,237,261]" pageId="21" pageNumber="439" value="1979-06-31">
<collectingDate id="EF90E9AFFFCACC2488A6FF347227FEA2" box="[1271,1395,237,261]" pageId="21" pageNumber="439" value="1979-06-31">31.VI.1979</collectingDate>
</date>
, coll. 
<collectorName id="269F5351FFCACC248CD9FED677AFFE80" box="[136,251,271,295]" pageId="21" pageNumber="439">Gentao Jin</collectorName>
&amp; 
<collectorName id="269F5351FFCACC248D4CFED676C4FE81" box="[285,400,271,295]" pageId="21" pageNumber="439">Jianyi Wu.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD53687FFCACC248CECFEEB76ACFEED" blockId="21.[136,1452,236,365]" box="[189,504,306,330]" pageId="21" pageNumber="439">
Distribution. 
<collectingCountry id="F37D7617FFCACC248D1DFEEB76D8FEED" box="[332,396,306,330]" name="China" pageId="21" pageNumber="439">China</collectingCountry>
(
<collectingRegion id="49AEF865FFCACC248DCBFEEB76B9FEED" box="[410,493,306,330]" country="China" name="Xizang" pageId="21" pageNumber="439">Xizang</collectingRegion>
).
</paragraph>
<paragraph id="8BD53687FFCACC248CECFE8D73EAFECB" blockId="21.[136,1452,236,365]" box="[189,1214,340,365]" pageId="21" pageNumber="439">
Etymology. The 
<typeStatus id="54D18825FFCACC248D3EFE8C76C9FECA" box="[367,413,341,365]" pageId="21" pageNumber="439">type</typeStatus>
location Maniweng (Mêdog, 
<collectingRegion id="49AEF865FFCACC248E8AFE8D747FFECB" box="[731,811,340,364]" country="China" name="Xizang" pageId="21" pageNumber="439">Xizang</collectingRegion>
, 
<collectingCountry id="F37D7617FFCACC248F64FE8D7422FECB" box="[821,886,340,364]" name="China" pageId="21" pageNumber="439">China</collectingCountry>
) is given as the specific name.
</paragraph>
</subSubSection>
</treatment>
</document>