267 lines
34 KiB
XML
267 lines
34 KiB
XML
<document id="0F554652A5E804E84EC00313F805EFC9" ID-CLB-Dataset="54167" ID-DOI="10.6620/ZS.2021.60-35" ID-GBIF-Dataset="dab15414-b59f-48a1-96ce-2515b2143001" ID-PMC="PMC8652416" ID-PubMed="34963788" ID-Zenodo-Dep="8070503" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" checkinTime="1687183603144" checkinUser="valdenar" docAuthor="Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés & Pañinao-Monsálvez, Laura" docDate="2021" docId="03A087D3FFCDFF8BB5CFFBD9FD65EE25" docLanguage="en" docName="ZoolStud.60.35.1-18.pdf" docOrigin="Zoological Studies 60 (35)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Holops anarayae Barahona-Segovia & Guzmán & Pañinao-Monsálvez 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="4" masterDocId="FF99FFABFFCFFF88B50DFF8BFFB4EC30" masterDocTitle="Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1699526456147" updateUser="plazi">
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<mods:titleInfo id="C176B25995D050771FC838DAC6EFEF6A">
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<mods:title id="C8859E1431880E6F0511B541195614F5">Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina</mods:title>
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<mods:namePart id="02820BCC67D5C8FF8E81D6D5B08E70E3">Barahona-Segovia, Rodrigo M.</mods:namePart>
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<mods:namePart id="07F766273924F4838D922C3F36B92F2F">Guzmán, Vicente Valdés</mods:namePart>
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<mods:namePart id="5C863D979371F00EBC7D417E1515C1A2">Pañinao-Monsálvez, Laura</mods:namePart>
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<mods:typeOfResource id="5AEBE198C423FAD89860EB508DD043FE">text</mods:typeOfResource>
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<mods:title id="7F84834DFEC4A09A01C74A7B44AF1604">Zoological Studies</mods:title>
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<mods:part id="E3311D778D8C8BDE772FDDD7C03A2A77">
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<mods:date id="DE2F56599B9056528A57CD6D52B674F4">2021</mods:date>
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<mods:number id="95B31CCFD1C2FF3AB4201644FA189CE3">2021-07-07</mods:number>
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<mods:number id="2B772FD604C3364D955E4A38D12794F2">60</mods:number>
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<mods:number id="1C0E4C94174404417746274212137D5B">35</mods:number>
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<treatment id="03A087D3FFCDFF8BB5CFFBD9FD65EE25" ID-DOI="http://doi.org/10.5281/zenodo.8055912" ID-GBIF-Taxon="210383222" ID-Zenodo-Dep="8055912" LSID="urn:lsid:plazi:treatment:03A087D3FFCDFF8BB5CFFBD9FD65EE25" httpUri="http://treatment.plazi.org/id/03A087D3FFCDFF8BB5CFFBD9FD65EE25" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="C313654EFFCDFF8AB5CFFBD9FD47E85B" box="[194,755,1105,1131]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BB636C5FFCDFF8AB5CFFBD9FD47E85B" blockId="2.[194,755,1105,1215]" box="[194,755,1105,1131]" pageId="2" pageNumber="3">
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<heading id="D0FE81A9FFCDFF8AB5CFFBD9FD47E85B" bold="true" box="[194,755,1105,1131]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
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<emphasis id="B97DEAD7FFCDFF8AB5CFFBD9FD47E85B" bold="true" box="[194,755,1105,1131]" pageId="2" pageNumber="3">
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<taxonomicName id="4C094D46FFCDFF8AB5CFFBD9FE2CE85B" ID-CoL="9HZB7" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[194,408,1106,1131]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anarayae" status="sp. nov.">
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<emphasis id="B97DEAD7FFCDFF8AB5CFFBD9FE2CE85B" bold="true" box="[194,408,1106,1131]" italics="true" pageId="2" pageNumber="3">Holops anarayae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57ACFFCDFF8AB492FBD9FDB5E85B" box="[415,513,1106,1131]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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Barahona-Segovia
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313654EFFCDFF8BB469FBF9FD65EE25" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BB636C5FFCDFF8AB469FBF9FD93E88F" blockId="2.[194,755,1105,1215]" pageId="2" pageNumber="3">
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(
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<figureCitation id="13322A40FFCDFF8AB461FBF9FE66E8BA" box="[364,466,1138,1162]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Figs. 1–2</figureCitation>
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,
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<figureCitation id="13322A40FFCDFF8AB4D0FBF9FE4EE8BA" box="[477,506,1138,1162]" captionStart="Figs" captionStartId="10.[174,213,1924,1943]" captionTargetBox="[175,1451,1033,1895]" captionTargetId="figure-6@10.[159,1468,991,1918]" captionTargetPageId="10" captionText="Figs. 15–20. Holops wing details: (15) H. anarayae sp. nov.; (16) H. cyanea (Philippi); (17) H. frauenfeldi Schiner; (18) H. grezi sp. nov.; (19) H. pullomen sp. nov., and (20) H. virens." figureDoi="http://doi.org/10.5281/zenodo.8070513" httpUri="https://zenodo.org/record/8070513/files/figure.png" pageId="2" pageNumber="3">15</figureCitation>
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,
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<figureCitation id="13322A40FFCDFF8AB708FBF9FDFDE8BA" box="[517,585,1138,1162]" captionStart="Figs" captionStartId="11.[136,175,1897,1916]" captionTargetBox="[238,1303,1064,1871]" captionTargetId="figure-5@11.[133,1450,993,1938]" captionTargetPageId="11" captionText="Figs. 21–26. Genitalia of Holops and Villalus: (21) dorsal view of H. cyanea (Philippi), female; (22) dorsal view of H. pullomen sp. nov.; (23) dorsal and lateral view of H. frauenfeldi Schiner, male; (24) dorsal view of V. inanis (Philippi), female; (25) lateral view of H. pullomen, male; (26) dorsal view of gonocoxa of H. anarayae sp. nov. epa = epandrium; gncx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="2" pageNumber="3">25–26</figureCitation>
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)
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<uri id="FF983AC7FFCDFF8AB5DCFB05FD93E88F" pageId="2" pageNumber="3">
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||
urn:lsid:zoobank.org:act:
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<uuid id="FFAF0C10FFCDFF8AB4B7FB05FD93E88F" pageId="2" pageNumber="3">E641677F-66FA-46A0-B797- 5FA1073404D2</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB5E7FB6EFE85E98D" blockId="2.[174,775,1253,1950]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB5E7FB6EFE0FE8CD" box="[234,443,1253,1277]" italics="true" pageId="2" pageNumber="3">Material examined</emphasis>
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:
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<materialsCitation id="3B613C98FFCDFF8AB4C1FB6EFE85E98D" collectingDate="1963-09-08" collectionCode="UACH, MNHNCL" collectorName="Leg. E. Krahmer" pageId="2" pageNumber="3" specimenCount="2" typeStatus="holotype">
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<emphasis id="B97DEAD7FFCDFF8AB4C1FB6EFD84E8CD" box="[460,560,1253,1277]" italics="true" pageId="2" pageNumber="3">
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<typeStatus id="54B28867FFCDFF8AB4C1FB6EFD84E8CD" box="[460,560,1253,1277]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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</emphasis>
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: male, in excellent condition, found in the Ernesto Krahmer’s collection from
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<collectionCode id="ED18AE00FFCDFF8AB5E0FAADFE8EE90E" box="[237,314,1318,1342]" pageId="2" pageNumber="3">UACH</collectionCode>
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and finally deposited in the
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<collectionCode id="ED18AE00FFCDFF8AB784FAADFCB7E90E" box="[649,771,1318,1342]" pageId="2" pageNumber="3">MNHNCL</collectionCode>
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; with the following labels: “Prov. Valdivia / Santo Domingo /
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<date id="FFB71005FFCDFF8AB421FAEEFE2AE94D" box="[300,414,1381,1405]" pageId="2" pageNumber="3" value="1963-09-08">
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<collectingDate id="EFF3E9EDFFCDFF8AB421FAEEFE2AE94D" box="[300,414,1381,1405]" pageId="2" pageNumber="3" value="1963-09-08">08.ix.1963</collectingDate>
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</date>
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/
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<collectorName id="26FC5313FFCDFF8AB4B5FAEDFDC2E94D" box="[440,630,1381,1406]" pageId="2" pageNumber="3">Leg. E. Krahmer</collectorName>
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”; “
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<typeStatus id="54B28867FFCDFF8AB798FAEEFCB3E94D" box="[661,775,1381,1405]" pageId="2" pageNumber="3" type="holotype">Holotypus</typeStatus>
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/
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<taxonomicName id="4C094D46FFCDFF8AB5B1FA0EFEC1E9AD" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[188,373,1413,1437]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anarayae">
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<emphasis id="B97DEAD7FFCDFF8AB5B1FA0EFEC1E9AD" box="[188,373,1413,1437]" italics="true" pageId="2" pageNumber="3">Holops anarayae</emphasis>
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</taxonomicName>
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/ sp. nov. ³ / det.
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<bibRefCitation id="EF984B34FFCDFF8AB74FFA0EFF52E98E" author="Barahona-Segovia RM & Valdes-Guzman V & Paninao-Monsalvez L." pageId="2" pageNumber="3" pagination="913234" refId="ref11464" refString="Barahona-Segovia RM, Valdes-Guzman V, Paninao-Monsalvez L. 2020. The species of the genus Physoconops Szilady (Diptera: Conopidae) from Chile, with the description of a new species: Physoconops tentenvilu nov. sp. Rev Mex Biodiv 91: e 913234. doi: 10.22201 / ib. 20078706 e. 2020.91.3234." type="journal article" year="2020">Barahona-Segovia 2020</bibRefCitation>
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” [red].
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB5E7FA4DFD00EA4D" blockId="2.[174,775,1253,1950]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB5E7FA4DFEE7E9ED" box="[234,339,1478,1501]" italics="true" pageId="2" pageNumber="3">
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<typeStatus id="54B28867FFCDFF8AB5E7FA4DFEE7E9ED" box="[234,339,1478,1501]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
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</emphasis>
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: male, in excellent condition, found in the Ernesto Krahmer’s collection from UACH and finally deposited in the MNHNCL; with the following labels: “Prov. Valdivia / Santo Domingo /
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<date id="FFB71005FFCDFF8AB78EF9AEFD42EA0D" box="[643,758,1573,1597]" pageId="2" pageNumber="3" value="1987-09-27">27.ix.1987</date>
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/ Leg. E. Krahmer”; “
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<typeStatus id="54B28867FFCDFF8AB49CF9CDFE47EA6E" box="[401,499,1606,1630]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
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/
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<taxonomicName id="4C094D46FFCDFF8AB703F9CEFD79EA6D" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[526,717,1605,1629]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anarayae">
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<emphasis id="B97DEAD7FFCDFF8AB703F9CEFD79EA6D" box="[526,717,1605,1629]" italics="true" pageId="2" pageNumber="3">Holops anarayae</emphasis>
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</taxonomicName>
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/ sp. nov. ³ / det.
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<bibRefCitation id="EF984B34FFCDFF8AB432F9EEFDF6EA4E" author="Barahona-Segovia RM & Valdes-Guzman V & Paninao-Monsalvez L." box="[319,578,1637,1662]" pageId="2" pageNumber="3" pagination="913234" refId="ref11464" refString="Barahona-Segovia RM, Valdes-Guzman V, Paninao-Monsalvez L. 2020. The species of the genus Physoconops Szilady (Diptera: Conopidae) from Chile, with the description of a new species: Physoconops tentenvilu nov. sp. Rev Mex Biodiv 91: e 913234. doi: 10.22201 / ib. 20078706 e. 2020.91.3234." type="journal article" year="2020">Barahona-Segovia 2020</bibRefCitation>
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” [yellow].
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB5E7F90DFE13EA8D" blockId="2.[174,775,1253,1950]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB5E7F90DFEC1EAAD" box="[234,373,1669,1693]" italics="true" pageId="2" pageNumber="3">Type locality</emphasis>
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: Valdivia Province, Santo Domingo,
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<collectingRegion id="49CDF827FFCDFF8AB5A3F92DFEEFEA8D" box="[174,347,1701,1726]" country="Chile" name="Los Rios" pageId="2" pageNumber="3">Los Ríos región</collectingRegion>
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,
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<collectingCountry id="F31E7655FFCDFF8AB46BF92EFE17EA8D" box="[358,419,1701,1725]" name="Chile" pageId="2" pageNumber="3">Chile</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB5E7F94EFEC5EB0D" blockId="2.[174,775,1253,1950]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB5E7F94EFEE9EAED" box="[234,349,1733,1757]" italics="true" pageId="2" pageNumber="3">Etymology</emphasis>
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: the specific epithet “
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<taxonomicName id="4C094D46FFCDFF8AB75DF94DFD01EAED" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[592,693,1734,1757]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anarayae">
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<emphasis id="B97DEAD7FFCDFF8AB75DF94DFD01EAED" box="[592,693,1734,1757]" italics="true" pageId="2" pageNumber="3">anarayae</emphasis>
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</taxonomicName>
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” refers to Ana Clara Luz Araya (1941–2020), the grandmother of the first author, which raised and helped pay for their university studies.
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB5E7F8CDFAB0EDA5" blockId="2.[174,775,1253,1950]" lastBlockId="2.[850,1452,253,1974]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB5E7F8CDFEE2EB6D" box="[234,342,1862,1885]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
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: dark blue species. Scutum with brown-yellowish pilosity. Postocellar lobe dark blue as well as all thorax segment excepting anepimeron, which is violet-brownish. Coxa and trochanter brownish. Femora bicolor (brown-yellow). Wings hyaline. M
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<subScript id="178D3480FFCDFF8AB011FEA1FA90ED08" attach="left" box="[1308,1316,298,312]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
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||
shorter than M
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<subScript id="178D3480FFCDFF8AB666FEC1FCC7ED68" attach="right" box="[875,883,330,344]" fontSize="6" pageId="2" pageNumber="3">2</subScript>
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. Cell r
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<subScript id="178D3480FFCDFF8AB6C8FEC1FC69ED68" attach="left" box="[965,989,330,344]" fontSize="6" pageId="2" pageNumber="3">4+5</subScript>
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elongated and thin. Abdomen elongated longitudinally and at same level of the maximum margins of thorax. All tergites dark blue.
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</paragraph>
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<paragraph id="8BB636C5FFCDFF8AB682FE15FBBFEBA5" blockId="2.[850,1452,253,1974]" pageId="2" pageNumber="3">
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<emphasis id="B97DEAD7FFCDFF8AB682FE15FBABED85" box="[911,1055,414,437]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
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:
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<emphasis id="B97DEAD7FFCDFF8AB139FE16FBD9ED85" box="[1076,1133,413,437]" italics="true" pageId="2" pageNumber="3">male</emphasis>
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: Length:
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<quantity id="4CF19B20FFCDFF8AB1E1FE16FAFAED85" box="[1260,1358,413,437]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="3" unit="mm" value="5.1">5.1 mm</quantity>
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; width:
|
||
<quantity id="4CF19B20FFCDFF8AB65FFE35FC17EDE5" box="[850,931,446,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="mm" value="1.8">1.8 mm</quantity>
|
||
(head),
|
||
<quantity id="4CF19B20FFCDFF8AB6F7FE36FBFFEDE5" box="[1018,1099,445,469]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="3" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(thorax) and
|
||
<quantity id="4CF19B20FFCDFF8AB1D7FE36FA9FEDE5" box="[1242,1323,445,469]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="3" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(abdomen);
|
||
<emphasis id="B97DEAD7FFCDFF8AB65FFE55FC3CEDC5" box="[850,904,478,501]" italics="true" pageId="2" pageNumber="3">wing</emphasis>
|
||
: 5.0 mm;
|
||
<emphasis id="B97DEAD7FFCDFF8AB109FE56FB8EEDC5" box="[1028,1082,477,501]" italics="true" pageId="2" pageNumber="3">head</emphasis>
|
||
: eyes brownish (
|
||
<figureCitation id="13322A40FFCDFF8AB00FFE56FAC5EDC6" box="[1282,1393,477,502]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Figs. 1–2</figureCitation>
|
||
); the size is more than 3/4 of the maximum height of the thorax (
|
||
<figureCitation id="13322A40FFCDFF8AB6A6FD96FC45EE06" box="[939,1009,541,566]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
); antenna inserted in the middle of the head; scape and pedicelum blackish with similar size; flagellum styliform approximately four times longer than pedicelum and scape together; antennae with similar length to head; eyes with long and brown pilosity; three ocelli dark bluish with ocellar triangle shiny blue somewhat protuberant; occiput dark and shiny blue with long and brownish pilosity; mouth parts yellowish with abundant brownish hairs;
|
||
<emphasis id="B97DEAD7FFCDFF8AB02EFC96FADEEF05" box="[1315,1386,797,821]" italics="true" pageId="2" pageNumber="3">thorax</emphasis>
|
||
: dark blue scutum with some shiny parts; long, dense and brownish pilosity, sparse but uniformly distributed (
|
||
<figureCitation id="13322A40FFCDFF8AB07BFCD6FCCBEFA6" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Figs. 1–2</figureCitation>
|
||
); postocellar lobe dark blue with 8–12 long and yellowish hairs (
|
||
<figureCitation id="13322A40FFCDFF8AB10AFC16FBFFEF86" box="[1031,1099,925,950]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
); anterior spiracle barely visible and darkish; anepisternum, katepisternum, katatergite, meron, katepimeron and anatergite shiny blue with long and yellowish pilosity (
|
||
<figureCitation id="13322A40FFCDFF8AB146FC76FB39E826" box="[1099,1165,1021,1046]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
); anepimeron violet-bluish with some yellowish hairs; subalar sclerite brownish and; scutellum shiny blue with long and brownish hairs;
|
||
<emphasis id="B97DEAD7FFCDFF8AB65FFBD6FCCBE845" box="[850,895,1117,1141]" italics="true" pageId="2" pageNumber="3">legs</emphasis>
|
||
: coxa and trochanters dark blue; femora almost completely dark bluish excepting anterior which yellow (
|
||
<figureCitation id="13322A40FFCDFF8AB657FB16FC28E886" box="[858,924,1181,1206]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
); tibiae yellow and tarsi brownish with dense and short pale pilosity (
|
||
<figureCitation id="13322A40FFCDFF8AB12BFB36FBDEE8E6" box="[1062,1130,1213,1238]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
); black claws;
|
||
<emphasis id="B97DEAD7FFCDFF8AB002FB35FAF9E8E5" box="[1295,1357,1214,1237]" italics="true" pageId="2" pageNumber="3">wings</emphasis>
|
||
: hyaline and membranous appearance; all veins dark brown; R
|
||
<subScript id="178D3480FFCDFF8AB0A9FB61FA18E8C8" attach="left" box="[1444,1452,1258,1272]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
|
||
and R
|
||
<subScript id="178D3480FFCDFF8AB69EFA81FC1FE928" attach="left" box="[915,939,1290,1304]" fontSize="6" pageId="2" pageNumber="3">2+3</subScript>
|
||
closer between them; R
|
||
<subScript id="178D3480FFCDFF8AB1BFFA81FB0EE928" attach="left" box="[1202,1210,1290,1304]" fontSize="6" pageId="2" pageNumber="3">4</subScript>
|
||
and R
|
||
<subScript id="178D3480FFCDFF8AB00FFA81FABEE928" attach="left" box="[1282,1290,1290,1304]" fontSize="6" pageId="2" pageNumber="3">5</subScript>
|
||
not petiolated, inward position (
|
||
<figureCitation id="13322A40FFCDFF8AB105FA96FBE1E905" box="[1032,1109,1309,1333]" captionStart="Figs" captionStartId="10.[174,213,1924,1943]" captionTargetBox="[175,1451,1033,1895]" captionTargetId="figure-6@10.[159,1468,991,1918]" captionTargetPageId="10" captionText="Figs. 15–20. Holops wing details: (15) H. anarayae sp. nov.; (16) H. cyanea (Philippi); (17) H. frauenfeldi Schiner; (18) H. grezi sp. nov.; (19) H. pullomen sp. nov., and (20) H. virens." figureDoi="http://doi.org/10.5281/zenodo.8070513" httpUri="https://zenodo.org/record/8070513/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
|
||
) and separated in the distal part for the half of length of each vein; M
|
||
<subScript id="178D3480FFCDFF8AB1EBFAC1FB5AE968" attach="left" box="[1254,1262,1354,1368]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
|
||
, M
|
||
<subScript id="178D3480FFCDFF8AB019FAC1FAA8E968" attach="left" box="[1300,1308,1354,1368]" fontSize="6" pageId="2" pageNumber="3">2</subScript>
|
||
and M
|
||
<subScript id="178D3480FFCDFF8AB064FAC1FA35E968" attach="left" box="[1385,1409,1354,1368]" fontSize="6" pageId="2" pageNumber="3">3+4</subScript>
|
||
not reaching the wing margin; M
|
||
<subScript id="178D3480FFCDFF8AB19CFAE1FB2DE948" attach="left" box="[1169,1177,1386,1400]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
|
||
shorter than M
|
||
<subScript id="178D3480FFCDFF8AB04EFAE1FAFFE948" attach="left" box="[1347,1355,1386,1400]" fontSize="6" pageId="2" pageNumber="3">2</subScript>
|
||
; cell r
|
||
<subScript id="178D3480FFCDFF8AB09EFAE1FA1FE948" attach="left" box="[1427,1451,1386,1400]" fontSize="6" pageId="2" pageNumber="3">4+5</subScript>
|
||
elongated and thin, with 3/4 length of R
|
||
<subScript id="178D3480FFCDFF8AB00DFA01FAADE9A8" attach="left" box="[1280,1305,1418,1432]" fontSize="6" pageId="2" pageNumber="3">2+3</subScript>
|
||
vein and five sides; cell m
|
||
<subScript id="178D3480FFCDFF8AB6D2FA21FC53E988" attach="left" box="[991,999,1450,1464]" fontSize="6" pageId="2" pageNumber="3">3</subScript>
|
||
with 1/3 length of r
|
||
<subScript id="178D3480FFCDFF8AB1C1FA21FB51E988" attach="left" box="[1228,1253,1450,1464]" fontSize="6" pageId="2" pageNumber="3">4+5</subScript>
|
||
connected to cell bm with a very short projection of m-cu vein; halters brownish with apex some yellowish; calypter rim blackish and inner part is whitish with dense and long yellowish pilosity;
|
||
<emphasis id="B97DEAD7FFCDFF8AB123F996FB27EA05" box="[1070,1171,1565,1589]" italics="true" pageId="2" pageNumber="3">abdomen</emphasis>
|
||
: not curved (
|
||
<figureCitation id="13322A40FFCDFF8AB022F996FA29EA06" box="[1327,1437,1565,1590]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Figs. 1–2</figureCitation>
|
||
); oval shape similar in size to head and thorax together (
|
||
<figureCitation id="13322A40FFCDFF8AB656F9D6FC14EA46" box="[859,928,1629,1654]" captionStart="Figs" captionStartId="3.[136,176,1927,1946]" captionTargetBox="[174,1412,1084,1900]" captionTargetId="figure-5@3.[136,1413,1051,1900]" captionTargetPageId="3" captionText="Figs. 1–4. Holops anarayae sp. nov.: (1) dorsal view and (2) lateral view; Holops cyaneus (Philippi, 1865): (3) dorsal view and (4) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070505" httpUri="https://zenodo.org/record/8070505/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
); in dorsal view, the abdomen not protruded laterally from the thorax margins; all tergites completely rough and covered with short and brownish pilosity: tergites 1–4 with similar width; all sternites darkish;
|
||
<emphasis id="B97DEAD7FFCDFF8AB65FF956FC0CEAC5" box="[850,952,1757,1781]" italics="true" pageId="2" pageNumber="3">genitalia</emphasis>
|
||
: not detached and partially everted; cerci, part of the epandrium and gonocoxa visible; cerci light yellow with many short and yellow pilosity; aedeagus yellowish; gonocoxa brownish, enlarged (
|
||
<figureCitation id="13322A40FFCDFF8AB02FF8B6FAC7EB65" box="[1314,1395,1853,1877]" captionStart="Figs" captionStartId="11.[136,175,1897,1916]" captionTargetBox="[238,1303,1064,1871]" captionTargetId="figure-5@11.[133,1450,993,1938]" captionTargetPageId="11" captionText="Figs. 21–26. Genitalia of Holops and Villalus: (21) dorsal view of H. cyanea (Philippi), female; (22) dorsal view of H. pullomen sp. nov.; (23) dorsal and lateral view of H. frauenfeldi Schiner, male; (24) dorsal view of V. inanis (Philippi), female; (25) lateral view of H. pullomen, male; (26) dorsal view of gonocoxa of H. anarayae sp. nov. epa = epandrium; gncx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="2" pageNumber="3">Fig. 25</figureCitation>
|
||
) and bifurcated (
|
||
<figureCitation id="13322A40FFCDFF8AB6DDF8D6FB95EB45" box="[976,1057,1885,1909]" captionStart="Figs" captionStartId="11.[136,175,1897,1916]" captionTargetBox="[238,1303,1064,1871]" captionTargetId="figure-5@11.[133,1450,993,1938]" captionTargetPageId="11" captionText="Figs. 21–26. Genitalia of Holops and Villalus: (21) dorsal view of H. cyanea (Philippi), female; (22) dorsal view of H. pullomen sp. nov.; (23) dorsal and lateral view of H. frauenfeldi Schiner, male; (24) dorsal view of V. inanis (Philippi), female; (25) lateral view of H. pullomen, male; (26) dorsal view of gonocoxa of H. anarayae sp. nov. epa = epandrium; gncx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="2" pageNumber="3">Fig. 26</figureCitation>
|
||
) with several setae in ventral view;
|
||
<emphasis id="B97DEAD7FFCDFF8AB65FF8F6FC2DEBA5" box="[850,921,1917,1941]" italics="true" pageId="2" pageNumber="3">female</emphasis>
|
||
: unknown
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFCDFF8BB682F816FF58ED26" blockId="2.[850,1452,253,1974]" lastBlockId="3.[136,737,253,534]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="B97DEAD7FFCDFF8AB682F816FBA2EB85" box="[911,1046,1949,1973]" italics="true" pageId="2" pageNumber="3">Distribution</emphasis>
|
||
: only known from the
|
||
<typeStatus id="54B28867FFCDFF8AB02DF815FAFBEB86" box="[1312,1359,1950,1974]" pageId="2" pageNumber="3">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13322A40FFCCFF8BB59DFF76FF54ED26" box="[144,224,253,278]" captionStart="Fig" captionStartId="12.[175,206,1952,1971]" captionTargetId="figure-5@12.[322,1303,539,1925]" captionTargetPageId="12" captionText="Fig. 27. Distributional map of Holops species." figureDoi="http://doi.org/10.5281/zenodo.8070517" httpUri="https://zenodo.org/record/8070517/files/figure.png" pageId="3" pageNumber="4">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<footnote id="E8122ACBFFCDFF8AB191F86EFA1FEBC7" box="[1180,1451,2020,2039]" pageId="2" pageNumber="3">
|
||
<paragraph id="8BB636C5FFCDFF8AB191F86EFA1FEBC7" blockId="2.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="2" pageNumber="3">© 2021 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<paragraph id="8BB636C5FFCCFF8BB5C8FE96FD65EE25" blockId="3.[136,737,253,534]" pageId="3" pageNumber="4">
|
||
<emphasis id="B97DEAD7FFCCFF8BB5C8FE96FE9BED05" box="[197,303,285,309]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
|
||
: this endemic species belongs to Valdivian district inside Valdivian forest province, subantarctic subregion (
|
||
<bibRefCitation id="EF984B34FFCCFF8BB482FED5FD86ED45" author="Morrone JJ" box="[399,562,349,374]" pageId="3" pageNumber="4" pagination="207 - 236" refId="ref11905" refString="Morrone JJ. 2015. Biogeographical regionalisation of the Andean region. Zootaxa 3936: 207 - 236. doi: 10.11646 / zootaxa. 3936.2.3." type="journal article" year="2015">Morrone 2015</bibRefCitation>
|
||
). The Valdivian evergreen forest is mainly dominated by
|
||
<taxonomicName id="4C094D46FFCCFF8BB76DFEF5FE20ED85" authority="Ruiz et Pav." authorityName="Ruiz et Pav." class="Magnoliopsida" family="Aextoxicaceae" genus="Aextoxicum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Berberidopsidales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="punctatum">
|
||
<emphasis id="B97DEAD7FFCCFF8BB76DFEF5FF49ED85" italics="true" pageId="3" pageNumber="4">Aextoxicum punctatum</emphasis>
|
||
Ruiz et Pav.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D46FFCCFF8BB4AFFE15FDA3ED86" box="[418,535,414,438]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||
trees species and
|
||
<taxonomicName id="4C094D46FFCCFF8BB585FE36FEEBEDE5" box="[136,351,445,469]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B97DEAD7FFCCFF8BB585FE36FEB7EDE5" box="[136,259,445,469]" italics="true" pageId="3" pageNumber="4">Nothofagus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The species present high variability in color legs and length of R
|
||
<subScript id="178D3480FFCCFF8BB4B3FE61FE72EDC8" attach="left" box="[446,454,490,504]" fontSize="6" pageId="3" pageNumber="4">4</subScript>
|
||
, R
|
||
<subScript id="178D3480FFCCFF8BB4EBFE61FE5AEDC8" attach="left" box="[486,494,490,504]" fontSize="6" pageId="3" pageNumber="4">5</subScript>
|
||
and M veins of wings (Shaun Winterton, pers. comm.). The host is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |