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<document id="7B45F0759102FC3058EF581A03C717E4" ID-CLB-Dataset="298366" ID-DOI="10.11646/zootaxa.5458.3.5" ID-GBIF-Dataset="ebea9f7d-8042-4071-9150-0188619a167d" ID-ISSN="1175-5326" ID-Zenodo-Dep="11547603" ID-ZooBank="F263E968-63D0-4721-B09F-0CA4A6B50484" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1718021326182" checkinUser="plazi" docAuthor="Silva, Iara Siqueira Santos, Santos, Adalberto J. &amp; Carvalho, Leonardo Sousa" docDate="2024" docId="FC1B87881264FFE800A2E2F662ABA75F" docLanguage="en" docName="zootaxa.5458.3.5.pdf" docOrigin="Zootaxa 5458 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5458.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rowlandius ufpi Silva &amp; Santos &amp; Carvalho 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="417" masterDocId="0022FFF01266FFE70035E2616162A642" masterDocTitle="A potentially endangered new species of the genus Rowlandius (Arachnida: Schizomida: Hubbardiidae) from Northeastern Brazil" masterLastPageNumber="419" masterPageNumber="403" pageNumber="405" updateTime="1718122122975" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="90EF1F85C77C4FDB8AA9C1B4AB1062B6">A potentially endangered new species of the genus Rowlandius (Arachnida: Schizomida: Hubbardiidae) from Northeastern Brazil</mods:title>
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<mods:namePart id="620D09A487A095F902834B24A16C98D4">Silva, Iara Siqueira Santos</mods:namePart>
<mods:affiliation id="3588F6905F8597410D88656CC0AFF7E6">Programa de Pós-Graduação em Biodiversidade e Conservação, Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, BR 343, KM 3.5, Bairro Meladão, CEP 64808 - 605, Floriano, PI, Brazil</mods:affiliation>
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<mods:namePart id="A6F86E3D53A68116C6807C09E8D8D6CF">Santos, Adalberto J.</mods:namePart>
<mods:affiliation id="B2E773CD45073F60D591E17B2365C25D">Departamento de Zoologia, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, Av. Antônio Carlos, 6627, CEP 31270 - 901, Belo Horizonte, Minas Gerais, Brazil</mods:affiliation>
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<mods:namePart id="D12BEB3F64DB63FDBA2C90C1AB297091">Carvalho, Leonardo Sousa</mods:namePart>
<mods:affiliation id="F2F2CAE52F9D3FCB665B788E03BCC76C">Programa de Pós-Graduação em Biodiversidade e Conservação, Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, BR 343, KM 3.5, Bairro Meladão, CEP 64808 - 605, Floriano, PI, Brazil &amp; Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, BR 343, KM 3.5, Bairro Meladão, CEP 64808 - 605, Floriano, Piauí, Brazil &amp; Programa de Pós-Graduação em Biodiversidade e Conservação, Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, BR 343, KM 3.5, Bairro Meladão, CEP 64808 - 605, Floriano, PI, Brazil</mods:affiliation>
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<mods:date id="E9EC315124EB2198FACEE441F08ADEA2">2024</mods:date>
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<taxonomicName id="B3B24D1D1264FFE500A2E2F6603DA6F3" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[151,351,151,178]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1264FFE500A2E2F6603DA6F3" bold="true" box="[151,351,151,178]" italics="true" pageId="2" pageNumber="405">Rowlandius ufpi</emphasis>
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<figureCitation id="EC892A1B1264FFE500A2E2DA6048A694" box="[151,298,187,214]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–14" captionStart-10="FIGURE 58" captionStart-2="FIGURES 15–19" captionStart-3="FIGURES 20–26" captionStart-4="FIGURES 27–35" captionStart-5="FIGURES 36–42" captionStart-6="FIGURES 43–46" captionStart-7="FIGURES 47–50" captionStart-8="FIGURES 51–54" captionStart-9="FIGURES 55–57" captionStartId-0="3.[151,264,1496,1520]" captionStartId-1="4.[151,264,1887,1911]" captionStartId-10="14.[151,250,1079,1103]" captionStartId-2="5.[151,264,1577,1601]" captionStartId-3="6.[151,264,1442,1466]" captionStartId-4="7.[151,264,1939,1963]" captionStartId-5="8.[151,264,1911,1935]" captionStartId-6="10.[151,264,1884,1908]" captionStartId-7="11.[151,264,1226,1250]" captionStartId-8="12.[151,264,1112,1136]" captionStartId-9="13.[151,264,1467,1491]" captionTargetBox-0="[151,1436,179,1470]" captionTargetBox-1="[151,1436,179,1861]" captionTargetBox-10="[176,1413,187,1050]" captionTargetBox-2="[151,1436,423,1551]" captionTargetBox-3="[237,1349,239,1415]" captionTargetBox-4="[289,1299,522,1913]" captionTargetBox-5="[151,1436,630,1885]" captionTargetBox-6="[169,1420,313,1854]" captionTargetBox-7="[151,1436,342,1200]" captionTargetBox-8="[152,1436,234,1086]" captionTargetBox-9="[151,1436,187,1443]" captionTargetId-0="figure-19@3.[151,1436,179,1470]" captionTargetId-1="figure-23@4.[151,1436,179,1861]" captionTargetId-10="figure-23@14.[171,1417,181,1055]" captionTargetId-2="figure-148@5.[151,1436,414,1551]" captionTargetId-3="figure-56@6.[228,1359,234,1415]" captionTargetId-4="figure-157@7.[289,1299,522,1913]" captionTargetId-5="figure-246@8.[151,1436,630,1885]" captionTargetId-6="figure-85@10.[151,1436,306,1858]" captionTargetId-7="figure-93@11.[151,1436,342,1200]" captionTargetId-8="figure-52@12.[152,1436,234,1086]" captionTargetId-9="figure-19@13.[151,1436,181,1443]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="14" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." captionText-1="FIGURES 7–14. Rowlandius ufpi sp. nov. heteromorphic male, holotype (CHNUFPI 4056), dorsal (7–8), ventral (9–10) and lateral (11–12) habitus; prolateral (13) and retrolateral (14) views of the right pedipalp. Scale bars: 0.5 mm." captionText-10="FIGURE 58. Known geographic distribution of described schizomid species from continental South America, including the Rowlandius ufpi sp. nov. (red circle), the remaining Rowlandius taxa (yellow squares) and other taxa (blue triangles)." captionText-2="FIGURES 15–19. Rowlandius ufpi sp. nov., heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19)." captionText-3="FIGURES 20–26. Rowlandius ufpi sp. nov., heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm." captionText-4="FIGURES 27–35. Rowlandius ufpi sp. nov. homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm." captionText-5="FIGURES 36–42. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4064), habitus (36–39) and flagellum (40–42), in dorsal (36–37, 40) and ventral (38–39, 41) and retrolateral (42) views. Scale bars: 0.5 mm." captionText-6="FIGURES 43–46. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4057, flagellum (43–45): dorsal (43), ventral (44) and lateral (45) views; and internal genitalia (46). Scale bars: 0.5 mm (43–45), 0.1 mm (46). Abbreviations: APA, anterior portion of the chitinized arch; B, bulb; G, gonopod; LL, lateral lobes; ML, median lobes; PPA, posterior portion of the chitinized arch; S, stalk." captionText-7="FIGURES 47–50. Rowlandius ufpi sp. nov. female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm." captionText-8="FIGURES 51–54. Environments at the type-locality of Rowlandius ufpi sp. nov.. 51–52: Riparian zone, where the examined specimens were found. 53–54: Phytophysiognomies where no schizomid was observed, savanna (Cerrado stricto sensu; 53) and seasonally dry tropical forest (54)." captionText-9="FIGURES 55–57. Sites sampled in the search for Rowlandius ufpi sp. nov. specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where R. ufpi sp. nov. specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where R. ufpi sp. nov., no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth." figureDoi-0="http://doi.org/10.5281/zenodo.11547605" figureDoi-1="http://doi.org/10.5281/zenodo.11547607" figureDoi-10="http://doi.org/10.5281/zenodo.11547630" figureDoi-2="http://doi.org/10.5281/zenodo.11547612" figureDoi-3="http://doi.org/10.5281/zenodo.11547614" figureDoi-4="http://doi.org/10.5281/zenodo.11547616" figureDoi-5="http://doi.org/10.5281/zenodo.11547618" figureDoi-6="http://doi.org/10.5281/zenodo.11547620" figureDoi-7="http://doi.org/10.5281/zenodo.11547622" figureDoi-8="http://doi.org/10.5281/zenodo.11547626" figureDoi-9="http://doi.org/10.5281/zenodo.11547628" httpUri-0="https://zenodo.org/record/11547605/files/figure.png" httpUri-1="https://zenodo.org/record/11547607/files/figure.png" httpUri-10="https://zenodo.org/record/11547630/files/figure.png" httpUri-2="https://zenodo.org/record/11547612/files/figure.png" httpUri-3="https://zenodo.org/record/11547614/files/figure.png" httpUri-4="https://zenodo.org/record/11547616/files/figure.png" httpUri-5="https://zenodo.org/record/11547618/files/figure.png" httpUri-6="https://zenodo.org/record/11547620/files/figure.png" httpUri-7="https://zenodo.org/record/11547622/files/figure.png" httpUri-8="https://zenodo.org/record/11547626/files/figure.png" httpUri-9="https://zenodo.org/record/11547628/files/figure.png" pageId="2" pageNumber="405">Figures 1–58</figureCitation>
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<emphasis id="46C6EA8C1264FFE500A2E361606DA75A" box="[151,271,256,280]" italics="true" pageId="2" pageNumber="405">Rowlandius</emphasis>
sp.
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,
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<bibRefCitation id="10234B6F1264FFE5010AE3616372A75A" author="Ontano, A. Z. &amp; Gainett, G. &amp; Aharon, S. &amp; Ballesteros, J. A. &amp; Benavides, L. R. &amp; Corbett, K. F. &amp; Sharma, P. P." box="[319,528,256,280]" pageId="2" pageNumber="405" pagination="2446 - 2467" refId="ref8179" refString="Ontano, A. Z., Gainett, G., Aharon, S., Ballesteros, J. A., Benavides, L. R., Corbett, K. F. &amp; Sharma, P. P. (2021) Taxonomic Sampling and Rare Genomic Changes Overcome Long-Branch Attraction in the Phylogenetic Placement of Pseudoscorpions. Molecular Biology and Evolution, 38 (6), 2446 - 2467. https: // doi. org / 10.1093 / molbev / msab 038" type="journal article" year="2021">
Ontano
<emphasis id="46C6EA8C1264FFE501A5E36160A1A75A" box="[400,451,256,280]" italics="true" pageId="2" pageNumber="405">et al.</emphasis>
(2021)
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: 2459
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, fig.
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<emphasis id="46C6EA8C1264FFE500A2E3276025A71D" bold="true" box="[151,327,325,352]" pageId="2" pageNumber="405">Type material.</emphasis>
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:
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, riverine forest along the
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margins,
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<quantity id="B34A9B7B1264FFE50207E30B63E7A7C6" box="[562,645,362,388]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="405" unit="m" value="109.0">
<elevation id="FF9FD1AD1264FFE50207E30B63E7A7C6" box="[562,645,362,388]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="405" unit="m" value="109.0">109 m</elevation>
</quantity>
.
</specimenCount>
a.s.l. (
<geoCoordinate id="118650591264FFE502F8E30B6239A7C7" box="[717,859,362,389]" degrees="6" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="405" precision="1" seconds="33.2" value="-6.759222">6°45’33.2”S</geoCoordinate>
,
<geoCoordinate id="118650591264FFE5035EE30B6567A7C6" box="[875,1029,362,388]" degrees="43" direction="west" minutes="3" orientation="longitude" pageId="2" pageNumber="405" precision="1" seconds="17.9" value="-43.05497">43°3’17.9”W</geoCoordinate>
),
<collectorName id="D94753481264FFE50428E30B65DDA7C6" box="[1053,1215,362,388]" pageId="2" pageNumber="405">L.S. Carvalho</collectorName>
leg.,
<date id="000C105E1264FFE50533E30B641CA7C6" box="[1286,1406,362,388]" pageId="2" pageNumber="405" value="2020-02-20">
<collectingDate id="1048E9B61264FFE50533E30B641CA7C6" box="[1286,1406,362,388]" pageId="2" pageNumber="405" value="2020-02-20">20/II/2020</collectingDate>
</date>
, 1 heteromorphic
<specimenCount id="62B4FD171264FFE50174E3EF6034A7EA" box="[321,342,398,424]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
(CHNUFPI 4056)
</materialsCitation>
.
<materialsCitation id="C4DA3CC31264FFE50201E3EC6006A78F" ID-GBIF-Occurrence="4876020306" pageId="2" pageNumber="405">
<emphasis id="46C6EA8C1264FFE50201E3EC63D9A7E5" bold="true" box="[564,699,397,423]" pageId="2" pageNumber="405">
<typeStatus id="AB09883C1264FFE50201E3EC63D5A7E5" box="[564,695,397,423]" pageId="2" pageNumber="405" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
same locality and collector as
<typeStatus id="AB09883C1264FFE50426E3EF651AA7EA" box="[1043,1144,398,424]" pageId="2" pageNumber="405" type="holotype">holotype</typeStatus>
,
<date id="000C105E1264FFE504B6E3EF659EA7EA" box="[1155,1276,398,424]" pageId="2" pageNumber="405" value="2020-02-13">13/II/2020</date>
,
<specimenCount id="62B4FD171264FFE50532E3EF6453A7EA" box="[1287,1329,398,424]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
,
<specimenCount id="62B4FD171264FFE5050AE3EF64FFA7EA" box="[1343,1437,398,424]" count="17" pageId="2" pageNumber="405" type="generic">17 imm.</specimenCount>
(CHNUFPI4055),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5015CE3D36577A78F" ID-GBIF-Occurrence="4876020302" box="[361,1045,434,461]" pageId="2" pageNumber="405">
<date id="000C105E1264FFE5015CE3D36080A78E" box="[361,482,434,460]" pageId="2" pageNumber="405" value="2020-02-20">20/II/2020</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE5029FE3D363DDA78E" box="[682,703,434,460]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE502F6E3D36387A78E" box="[707,741,434,460]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f
<specimenCount id="62B4FD171264FFE502CDE3D36227A78E" box="[760,837,434,460]" count="2" pageId="2" pageNumber="405" type="generic">2 imm.</specimenCount>
(CHNUFPI4057),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5042FE3D36378A7B2" ID-GBIF-Occurrence="4876020317" collectingDate="2020-02-12" pageId="2" pageNumber="405">
<date id="000C105E1264FFE5042FE3D365F1A78E" box="[1050,1171,434,460]" pageId="2" pageNumber="405" value="2020-02-12">
<collectingDate id="1048E9B61264FFE5042FE3D365F1A78E" box="[1050,1171,434,460]" pageId="2" pageNumber="405" value="2020-02-12">12/II/2020</collectingDate>
</date>
,
<specimenCount id="62B4FD171264FFE504AEE3D365DFA78E" box="[1179,1213,434,460]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4211), L.S.
<location id="716D60451264FFE500F9E3B76051A7B2" LSID="urn:lsid:plazi:treatment:FC1B87881264FFE800A2E2F662ABA75F:716D60451264FFE500F9E3B76051A7B2" box="[204,307,470,496]" country="Brazil" name="Carvalho" pageId="2" pageNumber="405" stateProvince="Piaui">Carvalho</location>
&amp;
<collectionCode id="12A3AE5B1264FFE50163E3B76009A7B2" box="[342,363,470,496]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="405">R</collectionCode>
.
<collectorName id="D94753481264FFE5015AE3B76086A7B2" box="[367,484,470,496]" pageId="2" pageNumber="405">F. Ferreira</collectorName>
leg.,
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50214E3B765EAA7B2" ID-GBIF-Occurrence="4876020307" box="[545,1160,470,496]" collectingDate="2020-02-20" pageId="2" pageNumber="405">
<date id="000C105E1264FFE50214E3B763F8A7B2" box="[545,666,470,496]" pageId="2" pageNumber="405" value="2020-02-20">
<collectingDate id="1048E9B61264FFE50214E3B763F8A7B2" box="[545,666,470,496]" pageId="2" pageNumber="405" value="2020-02-20">20/II/2020</collectingDate>
</date>
, 1 heteromorphic
<specimenCount id="62B4FD171264FFE50356E3B7621AA7B2" box="[867,888,470,496]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE5034AE3B762C1A7B2" box="[895,931,470,496]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4064),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE504BAE3B7600BA457" ID-GBIF-Occurrence="4876020301" pageId="2" pageNumber="405">
1 heteromorphic
<specimenCount id="62B4FD171264FFE5057BE3B76401A7B2" box="[1358,1379,470,496]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE5055CE3B764ECA7B2" box="[1385,1422,470,497]" count="5" pageId="2" pageNumber="405" type="female">5 ♀</specimenCount>
f (CHNUFPI 4065);
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50144E39B6209A456" ID-GBIF-Occurrence="4876020303" box="[369,875,506,532]" pageId="2" pageNumber="405">
same municipality and collector as
<typeStatus id="AB09883C1264FFE50337E39B6205A456" box="[770,871,506,532]" pageId="2" pageNumber="405" type="holotype">holotype</typeStatus>
,
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50346E39B61BDA41E" ID-GBIF-Occurrence="4876020308" latitude="-6.761667" longitude="-43.05472" pageId="2" pageNumber="405">
<location id="716D60451264FFE50346E39B6421A456" LSID="urn:lsid:plazi:treatment:FC1B87881264FFE800A2E2F662ABA75F:716D60451264FFE50346E39B6421A456" box="[883,1347,506,532]" country="Brazil" latitude="-6.761667" longLatPrecision="21" longitude="-43.05472" name="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" stateProvince="Piaui">Fazenda do Colégio Técnico de Floriano</location>
, gallery forest along small water stream (
<geoCoordinate id="118650591264FFE50221E07F63EFA47B" box="[532,653,542,569]" degrees="6" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="405" precision="15" seconds="42" value="-6.761667">6°45’42”S</geoCoordinate>
,
<geoCoordinate id="118650591264FFE502A9E07F6242A47A" box="[668,800,542,568]" degrees="43" direction="west" minutes="3" orientation="longitude" pageId="2" pageNumber="405" precision="15" seconds="17" value="-43.05472">43°3’17”W</geoCoordinate>
),
<date id="000C105E1264FFE50302E07F62D2A47A" box="[823,944,542,568]" pageId="2" pageNumber="405" value="2012-02-13">13/II/2012</date>
,
<specimenCount id="62B4FD171264FFE5038BE07F6284A47A" box="[958,998,542,568]" count="4" pageId="2" pageNumber="405" type="female">4 ♀</specimenCount>
f (CHNUFPI 0676),
<specimenCount id="62B4FD171264FFE504EEE07F6461A47A" box="[1243,1283,542,568]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 0677),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE500D2E0236238A41E" ID-GBIF-Occurrence="4876020323" box="[231,858,578,604]" pageId="2" pageNumber="405">
<date id="000C105E1264FFE500D2E0236035A41E" box="[231,343,578,604]" pageId="2" pageNumber="405" value="2013-01-30">30/I/2013</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE5021FE0236323A41E" box="[554,577,578,604]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
,
<specimenCount id="62B4FD171264FFE5027BE0236311A41E" box="[590,627,578,604]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 0678),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50357E02363D2A4C2" ID-GBIF-Occurrence="4876020305" pageId="2" pageNumber="405">
<date id="000C105E1264FFE50357E02362B9A41E" box="[866,987,578,604]" pageId="2" pageNumber="405" value="2012-02-13">13/II/2012</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE5049BE02365A1A41E" box="[1198,1219,578,604]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
(CHNUFPI 0679),
<specimenCount id="62B4FD171264FFE500A2E00761DEA4C2" box="[151,188,614,640]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 0680),
<specimenCount id="62B4FD171264FFE50192E00760AEA4C2" box="[423,460,614,640]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 0681),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50282E0076558A4C2" ID-GBIF-Occurrence="4876020318" box="[695,1082,614,640]" pageId="2" pageNumber="405">
<date id="000C105E1264FFE50282E0076245A4C2" box="[695,807,614,640]" pageId="2" pageNumber="405" value="2013-01-30">30/I/2013</date>
,
<specimenCount id="62B4FD171264FFE50304E0076234A4C2" box="[817,854,614,640]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 2699),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50474E00760C7A4E6" ID-GBIF-Occurrence="4876020316" collectingDate="2013-01-30" pageId="2" pageNumber="405">
<date id="000C105E1264FFE50474E00765D3A4C2" box="[1089,1201,614,640]" pageId="2" pageNumber="405" value="2013-01-30">
<collectingDate id="1048E9B61264FFE50474E00765D3A4C2" box="[1089,1201,614,640]" pageId="2" pageNumber="405" value="2013-01-30">30/I/2013</collectingDate>
</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE505B5E00764F5A4C2" box="[1408,1431,614,640]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
,
<specimenCount id="62B4FD171264FFE500A2E0EB6185A4E6" box="[151,231,650,676]" count="1" pageId="2" pageNumber="405" type="generic">1 imm.</specimenCount>
(
<specimenCode id="24149EE51264FFE500C0E0EB60FBA4E6" box="[245,409,650,676]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="405">UFMG 29800</specimenCode>
),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50199E0EB62DFA4E6" ID-GBIF-Occurrence="4876020311" box="[428,957,650,676]" collectingDate="2013-01-30" pageId="2" pageNumber="405">
<date id="000C105E1264FFE50199E0EB637EA4E6" box="[428,540,650,676]" pageId="2" pageNumber="405" value="2013-01-30">
<collectingDate id="1048E9B61264FFE50199E0EB637EA4E6" box="[428,540,650,676]" pageId="2" pageNumber="405" value="2013-01-30">30/I/2013</collectingDate>
</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE502D8E0EB6260A4E6" box="[749,770,650,676]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
(
<specimenCode id="24149EE51264FFE50325E0EB62D7A4E6" box="[784,949,650,676]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="405">UFMG 29793</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="740D369E1264FFE500F2E0CC6283A5E3" blockId="2.[151,1437,325,1648]" pageId="2" pageNumber="405">
<emphasis id="46C6EA8C1264FFE500F2E0CC6377A485" bold="true" box="[199,533,685,711]" pageId="2" pageNumber="405">Other examined specimens.</emphasis>
<materialsCitation id="C4DA3CC31264FFE5022BE0CC6271A553" ID-GBIF-Occurrence="4876020313" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" collectorName="I. S. S. Silva &amp; T.C. Santos" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<emphasis id="46C6EA8C1264FFE5022BE0CC6380A48A" bold="true" box="[542,738,685,712]" pageId="2" pageNumber="405">
<collectingCountry id="0CA5760E1264FFE5022BE0CC63E9A485" box="[542,651,685,711]" name="Brazil" pageId="2" pageNumber="405">BRAZIL</collectingCountry>
.
<collectingRegion id="B676F87C1264FFE502AFE0CC63BFA48A" box="[666,733,685,712]" country="Brazil" name="Piaui" pageId="2" pageNumber="405">Piauí</collectingRegion>
:
</emphasis>
<emphasis id="46C6EA8C1264FFE502D9E0CF6232A48A" box="[748,848,686,712]" italics="true" pageId="2" pageNumber="405">
<collectingCounty id="9D6C4E121264FFE502D9E0CF6232A48A" box="[748,848,686,712]" pageId="2" pageNumber="405">Floriano</collectingCounty>
</emphasis>
,
<collectingMunicipality id="9469ACE41264FFE50355E0CF645AA48A" box="[864,1336,686,712]" pageId="2" pageNumber="405">Fazenda do Colégio Técnico de Floriano</collectingMunicipality>
, riverine forest along the
<location id="716D60451264FFE5017BE0B36097A4AE" LSID="urn:lsid:plazi:treatment:FC1B87881264FFE800A2E2F662ABA75F:716D60451264FFE5017BE0B36097A4AE" box="[334,501,722,748]" country="Brazil" county="Floriano" latitude="-6.759222" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" name="Parnaiba River" pageId="2" pageNumber="405" stateProvince="Piaui">Parnaíba River</location>
margins,
<specimenCount id="62B4FD171264FFE50250E0B363D5A4AE" box="[613,695,722,748]" count="109" pageId="2" pageNumber="405" type="generic">
<quantity id="B34A9B7B1264FFE50250E0B363D1A4AE" box="[613,691,722,748]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="405" unit="m" value="109.0">
<elevation id="FF9FD1AD1264FFE50250E0B363D1A4AE" box="[613,691,722,748]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="405" unit="m" value="109.0">109 m</elevation>
</quantity>
.
</specimenCount>
a.s.l. (
<geoCoordinate id="118650591264FFE502C2E0B362E4A4AF" box="[759,902,722,749]" degrees="6" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="405" precision="1" seconds="33.2" value="-6.759222">6°45’33.2”S</geoCoordinate>
,
<geoCoordinate id="118650591264FFE503A7E0B3654EA4AE" box="[914,1068,722,748]" degrees="43" direction="west" minutes="3" orientation="longitude" pageId="2" pageNumber="405" precision="1" seconds="17.9" value="-43.05497">43°3’17.9”W</geoCoordinate>
),
<collectorName id="D94753481264FFE50475E0B365DDA4AE" box="[1088,1215,722,748]" pageId="2" pageNumber="405">I.S.S. Silva</collectorName>
&amp;
<collectionCode id="12A3AE5B1264FFE504D6E0B36595A4AE" box="[1251,1271,722,748]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="405">T</collectionCode>
.
<collectorName id="D94753481264FFE504CFE0B36407A4AE" box="[1274,1381,722,748]" pageId="2" pageNumber="405">C. Santos</collectorName>
leg.,
<date id="000C105E1264FFE500A2E0976072A552" box="[151,272,758,784]" pageId="2" pageNumber="405" value="2020-02-20">
<collectingDate id="1048E9B61264FFE500A2E0976072A552" box="[151,272,758,784]" pageId="2" pageNumber="405" value="2020-02-20">20/II/2020</collectingDate>
</date>
, 1 homeomorphic
<specimenCount id="62B4FD171264FFE501D3E0976099A552" box="[486,507,758,784]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE50231E0976348A552" box="[516,554,758,784]" count="2" pageId="2" pageNumber="405" type="female">2 ♀</specimenCount>
f (CHNUFPI 4058),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50329E0976005A577" ID-GBIF-Occurrence="4876020309" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" collectorName="L. S. Carvalho et al." country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<collectorName id="D94753481264FFE50329E09762D9A552" box="[796,955,758,784]" pageId="2" pageNumber="405">L.S. Carvalho</collectorName>
<emphasis id="46C6EA8C1264FFE503F6E0966294A552" box="[963,1014,758,784]" italics="true" pageId="2" pageNumber="405">et al</emphasis>
. leg.,
<date id="000C105E1264FFE5040AE09765D5A552" box="[1087,1207,758,784]" pageId="2" pageNumber="405" value="2021-02-10">
<collectingDate id="1048E9B61264FFE5040AE09765D5A552" box="[1087,1207,758,784]" pageId="2" pageNumber="405" value="2021-02-10">10/II/2021</collectingDate>
</date>
, 1 heteromorphic
<specimenCount id="62B4FD171264FFE505B2E09764FEA552" box="[1415,1436,758,784]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
(CHNUFPI 4059),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5015BE17B631AA576" ID-GBIF-Occurrence="4876020319" box="[366,632,794,820]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<specimenCount id="62B4FD171264FFE5015BE17B60F1A576" box="[366,403,794,820]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4060),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5024AE17B6005A51A" ID-GBIF-Occurrence="4876020312" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" collectorName="L. S. Carvalho &amp; R. F. Ferreira" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<collectorName id="D94753481264FFE5024AE17B627EA576" box="[639,796,794,820]" pageId="2" pageNumber="405">L.S. Carvalho</collectorName>
&amp;
<collectionCode id="12A3AE5B1264FFE5030AE17B6236A576" box="[831,852,794,820]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="405">R</collectionCode>
.
<collectorName id="D94753481264FFE5036DE17B62AFA576" box="[856,973,794,820]" pageId="2" pageNumber="405">F. Ferreira</collectorName>
leg.,
<date id="000C105E1264FFE5043FE17B65E1A576" box="[1034,1155,794,820]" pageId="2" pageNumber="405" value="2020-02-20">
<collectingDate id="1048E9B61264FFE5043FE17B65E1A576" box="[1034,1155,794,820]" pageId="2" pageNumber="405" value="2020-02-20">20/II/2020</collectingDate>
</date>
, 1 heteromorphic
<specimenCount id="62B4FD171264FFE50578E17B6400A576" box="[1357,1378,794,820]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE5055CE17B64ECA576" box="[1385,1422,794,820]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4063),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5015BE15F6557A51A" ID-GBIF-Occurrence="4876020321" box="[366,1077,830,856]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
1 heteromorphic
<specimenCount id="62B4FD171264FFE50218E15F6320A51A" box="[557,578,830,856]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
1 homeomorphic
<specimenCount id="62B4FD171264FFE5033AE15F6246A51A" box="[783,804,830,856]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
<specimenCount id="62B4FD171264FFE5031FE15F622DA51A" box="[810,847,830,856]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4066),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50409E15F64FEA51A" ID-GBIF-Occurrence="4876020320" box="[1084,1436,830,856]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<specimenCount id="62B4FD171264FFE50409E15F6503A51A" box="[1084,1121,830,856]" count="3" pageId="2" pageNumber="405" type="female">3 ♀</specimenCount>
f
<specimenCount id="62B4FD171264FFE50443E15F65A4A51A" box="[1142,1222,830,856]" count="1" pageId="2" pageNumber="405" type="generic">1 imm.</specimenCount>
(CHNUFPI 4067),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE500A2E1036242A53E" ID-GBIF-Occurrence="4876020304" box="[151,800,866,892]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<collectorName id="D94753481264FFE500A2E1036054A53E" box="[151,310,866,892]" pageId="2" pageNumber="405">L.S. Carvalho</collectorName>
leg.,
<date id="000C105E1264FFE50142E1036366A53E" box="[375,516,866,892]" pageId="2" pageNumber="405" value="2022-12-20">
<collectingDate id="1048E9B61264FFE50142E1036366A53E" box="[375,516,866,892]" pageId="2" pageNumber="405" value="2022-12-20">20/XII/2022</collectingDate>
</date>
,
<specimenCount id="62B4FD171264FFE50224E1036355A53E" box="[529,567,866,892]" count="1" pageId="2" pageNumber="405" type="female">1 ♀</specimenCount>
f (CHNUFPI 4210),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE5031CE10364E7A53E" ID-GBIF-Occurrence="4876020310" box="[809,1413,866,892]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" collectorName="L. S. Carvalho et al." country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
same collector,
<date id="000C105E1264FFE503E9E103650BA53E" box="[988,1129,866,892]" pageId="2" pageNumber="405" value="2022-12-20">
<collectingDate id="1048E9B61264FFE503E9E103650BA53E" box="[988,1129,866,892]" pageId="2" pageNumber="405" value="2022-12-20">20/XII/2022</collectingDate>
</date>
,
<specimenCount id="62B4FD171264FFE50443E10365FFA53E" box="[1142,1181,866,892]" count="2" pageId="2" pageNumber="405" type="female">2 ♀</specimenCount>
f (CHNUFPI 4212),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE505BBE103634CA5E2" ID-GBIF-Occurrence="4876020315" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
1 heteromorphic
<specimenCount id="62B4FD171264FFE50174E1E76034A5E2" box="[321,342,902,928]" count="1" pageId="2" pageNumber="405" type="male">♂</specimenCount>
(CHNUFPI 4239),
</materialsCitation>
<materialsCitation id="C4DA3CC31264FFE50200E1E762BFA5E3" ID-GBIF-Occurrence="4876020314" box="[565,989,902,929]" collectingDate="2020-02-20" collectingDateMax="2022-12-20" collectingDateMin="2020-02-20" collectionCode="T, R" country="Brazil" county="Floriano" elevation="109" latitude="-6.759222" location="Parnaiba River" longLatPrecision="1" longitude="-43.05497" municipality="Fazenda do Colegio Tecnico de Floriano" pageId="2" pageNumber="405" specimenCount="128" specimenCount-female="11" specimenCount-male="7" stateProvince="Piaui">
<specimenCount id="62B4FD171264FFE50200E1E7633DA5E2" box="[565,607,902,928]" count="1" pageId="2" pageNumber="405" type="male">1 ♂</specimenCount>
heteromorphic (CHNUFPI 4245)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3CA865151264FFE500F2E1CB6544A5AA" pageId="2" pageNumber="405" type="etymology">
<paragraph id="740D369E1264FFE500F2E1CB6544A5AA" blockId="2.[151,1437,325,1648]" pageId="2" pageNumber="405">
<emphasis id="46C6EA8C1264FFE500F2E1CB602CA586" bold="true" box="[199,334,938,964]" pageId="2" pageNumber="405">Etymology.</emphasis>
The specific name is a noun in apposition taken from the acronym of the Universidade Federal do
<collectingRegion id="B676F87C1264FFE500A2E1AF61B6A5AA" box="[151,212,974,1000]" country="Brazil" name="Piaui" pageId="2" pageNumber="405">Piauí</collectingRegion>
. The university owns the type-locality area and supported the present study.
</paragraph>
</subSubSection>
<subSubSection id="3CA865151264FFE500F2E19062F8A3FE" pageId="2" pageNumber="405" type="diagnosis">
<paragraph id="740D369E1264FFE500F2E19062F8A3FE" blockId="2.[151,1437,325,1648]" pageId="2" pageNumber="405">
<emphasis id="46C6EA8C1264FFE500F2E1906020A249" bold="true" box="[199,322,1009,1035]" pageId="2" pageNumber="405">Diagnosis.</emphasis>
The flagellum of males of
<taxonomicName id="B3B24D1D1264FFE502B5E1936258A24E" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[640,826,1010,1036]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1264FFE502B5E1936258A24E" box="[640,826,1010,1036]" italics="true" pageId="2" pageNumber="405">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1264FFE50376E19362FCA24E" bold="true" box="[835,926,1010,1036]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="5DF557F71264FFE50376E19362FCA24E" box="[835,926,1010,1036]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is subquadrate, as in
<taxonomicName id="B3B24D1D1264FFE504AEE192642BA24E" authorityName="Santos, Dias, Brescovit &amp; Santos" authorityYear="2008" box="[1179,1353,1010,1036]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="linsduarteae">
<emphasis id="46C6EA8C1264FFE504AEE192642BA24E" box="[1179,1353,1010,1036]" italics="true" pageId="2" pageNumber="405">R. linsduarteae</emphasis>
</taxonomicName>
and
<taxonomicName id="B3B24D1D1264FFE505B1E1926195A272" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="potiguar">
<emphasis id="46C6EA8C1264FFE505B1E1926195A272" italics="true" pageId="2" pageNumber="405">R. potiguar</emphasis>
</taxonomicName>
, but differs in the
<emphasis id="46C6EA8C1264FFE501F1E6776096A272" box="[452,500,1046,1072]" italics="true" pageId="2" pageNumber="405">dm1</emphasis>
setae located on the basal third of the bulb (
<figureCitation id="EC892A1B1264FFE503EBE6776502A272" box="[990,1120,1046,1072]" captionStart="FIGURES 20–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[237,1349,239,1415]" captionTargetId="figure-56@6.[228,1359,234,1415]" captionTargetPageId="6" captionText="FIGURES 20–26. Rowlandius ufpi sp. nov., heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547614" httpUri="https://zenodo.org/record/11547614/files/figure.png" pageId="2" pageNumber="405">Figs 22–24</figureCitation>
), whereas in
<taxonomicName id="B3B24D1D1264FFE504C7E67664FEA272" authorityName="Santos, Dias, Brescovit &amp; Santos" authorityYear="2008" box="[1266,1436,1046,1072]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="linsduarteae">
<emphasis id="46C6EA8C1264FFE504C7E67664FEA272" box="[1266,1436,1046,1072]" italics="true" pageId="2" pageNumber="405">R. linsduarteae</emphasis>
</taxonomicName>
and
<taxonomicName id="B3B24D1D1264FFE500F3E65A6027A216" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[198,325,1083,1108]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="potiguar">
<emphasis id="46C6EA8C1264FFE500F3E65A6027A216" box="[198,325,1083,1108]" italics="true" pageId="2" pageNumber="405">R. potiguar</emphasis>
</taxonomicName>
it is located on the pedicel. It also resembles
<taxonomicName id="B3B24D1D1264FFE50309E65A62D6A216" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[828,948,1082,1108]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ubajara">
<emphasis id="46C6EA8C1264FFE50309E65A62D6A216" box="[828,948,1082,1108]" italics="true" pageId="2" pageNumber="405">R. ubajara</emphasis>
</taxonomicName>
by the rounded sides, but differs in the long postero-median dorsal protuberance with the
<emphasis id="46C6EA8C1264FFE502A2E63F63AAA23A" box="[663,712,1118,1144]" italics="true" pageId="2" pageNumber="405">dm4</emphasis>
setae at the posterior end (
<figureCitation id="EC892A1B1264FFE503CCE63F6519A23A" box="[1017,1147,1118,1144]" captionStart="FIGURES 15–19" captionStartId="5.[151,264,1577,1601]" captionTargetBox="[151,1436,423,1551]" captionTargetId="figure-148@5.[151,1436,414,1551]" captionTargetPageId="5" captionText="FIGURES 15–19. Rowlandius ufpi sp. nov., heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19)." figureDoi="http://doi.org/10.5281/zenodo.11547612" httpUri="https://zenodo.org/record/11547612/files/figure.png" pageId="2" pageNumber="405">Figs 17–19</figureCitation>
), whereas it is short in
<taxonomicName id="B3B24D1D1264FFE505B1E63E6192A2DE" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ubajara">
<emphasis id="46C6EA8C1264FFE505B1E63E6192A2DE" italics="true" pageId="2" pageNumber="405">R. ubajara</emphasis>
</taxonomicName>
. The flagellum of
<taxonomicName id="B3B24D1D1264FFE50188E6E26365A2DE" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[445,519,1154,1180]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1264FFE50188E6E26365A2DE" box="[445,519,1154,1180]" italics="true" pageId="2" pageNumber="405">R. ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1264FFE50238E6E36304A2DE" bold="true" box="[525,614,1154,1180]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="5DF557F71264FFE50238E6E36304A2DE" box="[525,614,1154,1180]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to that of
<taxonomicName id="B3B24D1D1264FFE50377E6E262A3A2DE" authorityName="Giupponi, Miranda &amp; Villarreal" authorityYear="2016" box="[834,961,1154,1180]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="pedrosoi">
<emphasis id="46C6EA8C1264FFE50377E6E262A3A2DE" box="[834,961,1154,1180]" italics="true" pageId="2" pageNumber="405">R. pedrosoi</emphasis>
</taxonomicName>
by the protuberance in the postero-median region, differing from the latter by the postero-median protuberance connected to the dorso-lateral depressions, whereas in
<taxonomicName id="B3B24D1D1264FFE50122E6AA60FAA2A6" authorityName="Giupponi, Miranda &amp; Villarreal" authorityYear="2016" box="[279,408,1226,1252]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="pedrosoi">
<emphasis id="46C6EA8C1264FFE50122E6AA60FAA2A6" box="[279,408,1226,1252]" italics="true" pageId="2" pageNumber="405">R. pedrosoi</emphasis>
</taxonomicName>
the protuberance is distal to the depressions. Additionally, the male flagellum of
<taxonomicName id="B3B24D1D1264FFE5051DE6AA6416A2A6" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[1320,1396,1226,1252]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1264FFE5051DE6AA6416A2A6" box="[1320,1396,1226,1252]" italics="true" pageId="2" pageNumber="405">R. ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1264FFE5054EE6AB61AAA34A" bold="true" pageId="2" pageNumber="405">
<taxonomicNameLabel id="5DF557F71264FFE5054EE6AB61AAA34A" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is longer than the length of the pedicel, with the
<emphasis id="46C6EA8C1264FFE502D7E68F6261A34A" box="[738,771,1262,1288]" italics="true" pageId="2" pageNumber="405">vl2</emphasis>
setae proximal to the
<emphasis id="46C6EA8C1264FFE503C3E68F6578A34A" box="[1014,1050,1262,1288]" italics="true" pageId="2" pageNumber="405">dl3</emphasis>
level and
<emphasis id="46C6EA8C1264FFE504BEE68F65CFA34A" box="[1163,1197,1262,1288]" italics="true" pageId="2" pageNumber="405">vl1</emphasis>
posterior to
<emphasis id="46C6EA8C1264FFE5050DE68F6405A34A" box="[1336,1383,1262,1288]" italics="true" pageId="2" pageNumber="405">vm4</emphasis>
, and
<emphasis id="46C6EA8C1264FFE500A2E77361A4A36E" box="[151,198,1298,1324]" italics="true" pageId="2" pageNumber="405">vm5</emphasis>
distant from
<emphasis id="46C6EA8C1264FFE5016BE77360E3A36E" box="[350,385,1298,1324]" italics="true" pageId="2" pageNumber="405">vl1</emphasis>
. The female spermathecae is similar to that of
<taxonomicName id="B3B24D1D1264FFE503A0E772656CA36E" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[917,1038,1298,1324]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ubajara">
<emphasis id="46C6EA8C1264FFE503A0E772656CA36E" box="[917,1038,1298,1324]" italics="true" pageId="2" pageNumber="405">R. ubajara</emphasis>
</taxonomicName>
by the long and thin median lobes, and similar to
<taxonomicName id="B3B24D1D1264FFE50102E75660D4A312" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[311,438,1335,1360]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="potiguar">
<emphasis id="46C6EA8C1264FFE50102E75660D4A312" box="[311,438,1335,1360]" italics="true" pageId="2" pageNumber="405">R. potiguar</emphasis>
</taxonomicName>
,
<taxonomicName id="B3B24D1D1264FFE501F1E7566326A312" authorityName="Giupponi, Miranda &amp; Villarreal" authorityYear="2016" box="[452,580,1334,1360]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="pedrosoi">
<emphasis id="46C6EA8C1264FFE501F1E7566326A312" box="[452,580,1334,1360]" italics="true" pageId="2" pageNumber="405">R. pedrosoi</emphasis>
</taxonomicName>
and
<taxonomicName id="B3B24D1D1264FFE5024FE7566268A312" authorityName="Santos, Dias, Brescovit &amp; Santos" authorityYear="2008" box="[634,778,1334,1360]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="linsduarte">
<emphasis id="46C6EA8C1264FFE5024FE7566268A312" box="[634,778,1334,1360]" italics="true" pageId="2" pageNumber="405">R. linsduarte</emphasis>
</taxonomicName>
by the developed bulbs of the lateral lobes. The females of
<taxonomicName id="B3B24D1D1264FFE500A2E73A6180A336" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[151,226,1370,1396]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1264FFE500A2E73A6180A336" box="[151,226,1370,1396]" italics="true" pageId="2" pageNumber="405">R. ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1264FFE500DCE73B6020A336" bold="true" box="[233,322,1370,1396]" pageId="2" pageNumber="405">
<taxonomicNameLabel id="5DF557F71264FFE500DCE73B6020A336" box="[233,322,1370,1396]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="EC892A1B1264FFE50164E73B60B3A337" box="[337,465,1370,1397]" captionStart-0="FIGURES 43–46" captionStart-1="FIGURES 47–50" captionStartId-0="10.[151,264,1884,1908]" captionStartId-1="11.[151,264,1226,1250]" captionTargetBox-0="[169,1420,313,1854]" captionTargetBox-1="[151,1436,342,1200]" captionTargetId-0="figure-85@10.[151,1436,306,1858]" captionTargetId-1="figure-93@11.[151,1436,342,1200]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 43–46. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4057, flagellum (43–45): dorsal (43), ventral (44) and lateral (45) views; and internal genitalia (46). Scale bars: 0.5 mm (43–45), 0.1 mm (46). Abbreviations: APA, anterior portion of the chitinized arch; B, bulb; G, gonopod; LL, lateral lobes; ML, median lobes; PPA, posterior portion of the chitinized arch; S, stalk." captionText-1="FIGURES 47–50. Rowlandius ufpi sp. nov. female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11547620" figureDoi-1="http://doi.org/10.5281/zenodo.11547622" httpUri-0="https://zenodo.org/record/11547620/files/figure.png" httpUri-1="https://zenodo.org/record/11547622/files/figure.png" pageId="2" pageNumber="405">Figs 46–50</figureCitation>
) can be differentiated by the combination of: (1) highly developed lateral lobes, which are attached to the chitinized arch by very thin stalks; (2) elongated and very thin median lobes; and (3) anterior portion of the chitinized arch with an obtuse (&gt;150º) v-shaped angle.
</paragraph>
</subSubSection>
<subSubSection id="3CA865151264FFEC00F2E7A46355A703" lastPageId="11" lastPageNumber="414" pageId="2" pageNumber="405" type="description">
<paragraph id="740D369E1264FFE500F2E7A460B8A032" blockId="2.[151,1437,325,1648]" pageId="2" pageNumber="405">
<emphasis id="46C6EA8C1264FFE500F2E7A4603BA39D" bold="true" box="[199,345,1477,1503]" pageId="2" pageNumber="405">Description.</emphasis>
<emphasis id="46C6EA8C1264FFE50154E7A762ECA3A2" box="[353,910,1478,1504]" italics="true" pageId="2" pageNumber="405">
Heteromorphic male (
<typeStatus id="AB09883C1264FFE5026EE7A763DDA3A2" box="[603,703,1478,1504]" pageId="2" pageNumber="405" type="holotype">holotype</typeStatus>
, CHNUFPI 4056)
</emphasis>
. Coloration (
<figureCitation id="EC892A1B1264FFE50413E7A76508A3A2" box="[1062,1130,1478,1504]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="2" pageNumber="405">Fig. 1</figureCitation>
): general pattern greenish-brown. Prosoma is light brown; pedipalp and chelicerae reddish brown. Venter pale green. Legs: coxae I–IV brownish; trochanter I light red and II–IV greenish; femur I–IV greenish brown; patella I slightly reddish and II–IV greenish brown; tibia I reddish and II–IV light brown; tarsus I reddish and II–IV light brown. The opisthosoma and the flagellum are dark green.
</paragraph>
<caption id="20CD66161265FFE400A2E7B96228A056" ID-DOI="http://doi.org/10.5281/zenodo.11547605" ID-Zenodo-Dep="11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="3" pageNumber="406" startId="3.[151,264,1496,1520]" targetBox="[151,1436,179,1470]" targetPageId="3" targetType="figure">
<paragraph id="740D369E1265FFE400A2E7B96228A056" blockId="3.[151,1436,1496,1556]" pageId="3" pageNumber="406">
<emphasis id="46C6EA8C1265FFE400A2E7B96054A3B2" bold="true" box="[151,310,1496,1520]" pageId="3" pageNumber="406">FIGURES 1–6</emphasis>
.
<taxonomicName id="B3B24D1D1265FFE40176E7B96088A3B2" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[323,490,1496,1520]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1265FFE40176E7B96088A3B2" box="[323,490,1496,1520]" italics="true" pageId="3" pageNumber="406">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1265FFE401C7E7B96321A3B2" bold="true" box="[498,579,1496,1520]" pageId="3" pageNumber="406">
<taxonomicNameLabel id="5DF557F71265FFE401C7E7B96321A3B2" box="[498,579,1496,1520]" pageId="3" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3).
</paragraph>
</caption>
<paragraph id="740D369E1265FFE400F2E4066226A0AF" blockId="3.[151,1437,1638,1773]" pageId="3" pageNumber="406">
<emphasis id="46C6EA8C1265FFE400F2E4066049A0C2" box="[199,299,1639,1664]" italics="true" pageId="3" pageNumber="406">Prosoma</emphasis>
(
<figureCitation id="EC892A1B1265FFE4010EE40760E1A0C3" box="[315,387,1638,1665]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="3" pageNumber="406">Figs 1</figureCitation>
,
<figureCitation id="EC892A1B1265FFE401A7E406608AA0C3" box="[402,488,1639,1665]" captionStart="FIGURES 7–14" captionStartId="4.[151,264,1887,1911]" captionTargetBox="[151,1436,179,1861]" captionTargetId="figure-23@4.[151,1436,179,1861]" captionTargetPageId="4" captionText="FIGURES 7–14. Rowlandius ufpi sp. nov. heteromorphic male, holotype (CHNUFPI 4056), dorsal (7–8), ventral (9–10) and lateral (11–12) habitus; prolateral (13) and retrolateral (14) views of the right pedipalp. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547607" httpUri="https://zenodo.org/record/11547607/files/figure.png" pageId="3" pageNumber="406">7, 9, 11</figureCitation>
). 1.5 long, 0.6 wide. Dorsal region: anterior process of propeltidium with 1+1 setae followed by 2 pairs of dorso-submedian setae; the tip of the process is bent downwards. Propeltidium 2 times longer than wide; eyespot present; metapeltidium entire, much wider than long and totally bristle-less. Ventral region: sternum anteriorly with 13 setae and posteriorly with 6 setae.
</paragraph>
<caption id="20CD66161262FFE300A2E53E65BAA1D8" ID-DOI="http://doi.org/10.5281/zenodo.11547607" ID-Zenodo-Dep="11547607" httpUri="https://zenodo.org/record/11547607/files/figure.png" pageId="4" pageNumber="407" startId="4.[151,264,1887,1911]" targetBox="[151,1436,179,1861]" targetPageId="4" targetType="figure">
<paragraph id="740D369E1262FFE300A2E53E65BAA1D8" blockId="4.[151,1437,1887,1947]" pageId="4" pageNumber="407">
<emphasis id="46C6EA8C1262FFE300A2E53E6023A135" bold="true" box="[151,321,1887,1911]" pageId="4" pageNumber="407">FIGURES 7–14</emphasis>
.
<taxonomicName id="B3B24D1D1262FFE3017AE53E6097A135" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[335,501,1887,1911]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1262FFE3017AE53E6097A135" box="[335,501,1887,1911]" italics="true" pageId="4" pageNumber="407">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1262FFE301C8E53E632DA135" bold="true" box="[509,591,1887,1911]" pageId="4" pageNumber="407">
<taxonomicNameLabel id="5DF557F71262FFE301C8E53E632DA135" box="[509,591,1887,1911]" pageId="4" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
heteromorphic male, holotype (CHNUFPI 4056), dorsal (7–8), ventral (9–10) and lateral (11–12) habitus; prolateral (13) and retrolateral (14) views of the right pedipalp. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="740D369E1263FFE200F2E2F663DBA7CB" blockId="5.[151,1437,150,393]" pageId="5" pageNumber="408">
<emphasis id="46C6EA8C1263FFE200F2E2F66020A6F3" box="[199,322,151,177]" italics="true" pageId="5" pageNumber="408">Chelicerae</emphasis>
(
<figureCitation id="EC892A1B1263FFE20165E2F660B3A6F3" box="[336,465,150,177]" captionStart="FIGURES 15–19" captionStartId="5.[151,264,1577,1601]" captionTargetBox="[151,1436,423,1551]" captionTargetId="figure-148@5.[151,1436,414,1551]" captionTargetPageId="5" captionText="FIGURES 15–19. Rowlandius ufpi sp. nov., heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19)." figureDoi="http://doi.org/10.5281/zenodo.11547612" httpUri="https://zenodo.org/record/11547612/files/figure.png" pageId="5" pageNumber="408">Figs 15–16</figureCitation>
). Movable finger sharp and curved distally, guard tooth present, serrula composed of 12 hyaline teeth, and a row of 15 long setae. Fixed finger with one basal tooth, followed by five small teeth of similar size and one larger distal tooth, curved tooth with acute apex. Setation: G1 with three spatulate setae; G2 composed of five feathered setae, and long setae, almost the length of the moving finger; G3 with four regular, cylindrical setae; G4 consisting of two smooth and short setae with thin apex; G5 (A) with seven similar setae; G5 (B) with six setae along the ventral border; G6 with one smooth setae, longer than half of movable finger length; G7 with six setae. Setal group formula: 3–5–4–2–(7+6)–1–6.
</paragraph>
<caption id="20CD66161263FFE200A2E4486408A027" ID-DOI="http://doi.org/10.5281/zenodo.11547612" ID-Zenodo-Dep="11547612" httpUri="https://zenodo.org/record/11547612/files/figure.png" pageId="5" pageNumber="408" startId="5.[151,264,1577,1601]" targetBox="[151,1436,423,1551]" targetPageId="5" targetType="figure">
<paragraph id="740D369E1263FFE200A2E4486408A027" blockId="5.[151,1436,1577,1637]" pageId="5" pageNumber="408">
<emphasis id="46C6EA8C1263FFE200A2E448602EA003" bold="true" box="[151,332,1577,1601]" pageId="5" pageNumber="408">FIGURES 15–19</emphasis>
.
<taxonomicName id="B3B24D1D1263FFE2016DE448609EA003" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[344,508,1577,1601]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="5" pageNumber="408" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1263FFE2016DE448609EA003" box="[344,508,1577,1601]" italics="true" pageId="5" pageNumber="408">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1263FFE20234E4486333A003" bold="true" box="[513,593,1577,1601]" pageId="5" pageNumber="408">
<taxonomicNameLabel id="5DF557F71263FFE20234E4486333A003" box="[513,593,1577,1601]" pageId="5" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19).
</paragraph>
</caption>
<paragraph id="740D369E1263FFE100F2E4D6650CA697" blockId="5.[151,1437,1719,2034]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="409" pageId="5" pageNumber="408">
<emphasis id="46C6EA8C1263FFE200F2E4D6604EA093" box="[199,300,1719,1745]" italics="true" pageId="5" pageNumber="408">Pedipalp</emphasis>
(
<figureCitation id="EC892A1B1263FFE2010FE4D660D9A090" box="[314,443,1719,1746]" captionStart="FIGURES 7–14" captionStartId="4.[151,264,1887,1911]" captionTargetBox="[151,1436,179,1861]" captionTargetId="figure-23@4.[151,1436,179,1861]" captionTargetPageId="4" captionText="FIGURES 7–14. Rowlandius ufpi sp. nov. heteromorphic male, holotype (CHNUFPI 4056), dorsal (7–8), ventral (9–10) and lateral (11–12) habitus; prolateral (13) and retrolateral (14) views of the right pedipalp. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547607" httpUri="https://zenodo.org/record/11547607/files/figure.png" pageId="5" pageNumber="408">Figs 13–14</figureCitation>
,
<figureCitation id="EC892A1B1263FFE201F3E4D9636CA090" box="[454,526,1720,1746]" captionStart="FIGURES 20–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[237,1349,239,1415]" captionTargetId="figure-56@6.[228,1359,234,1415]" captionTargetPageId="6" captionText="FIGURES 20–26. Rowlandius ufpi sp. nov., heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547614" httpUri="https://zenodo.org/record/11547614/files/figure.png" pageId="5" pageNumber="408">20, 21</figureCitation>
). Setation described in detail for homeomorphic male. Trochanter: Subcylindrical in retrolateral view, longer than wide, with apical portion curved upwards; with 12 ventral setae, a ventral row of large setae with an intermediate row of small setae. Femur: subcylindrical, curved in the basal portion, club-shaped, 9.2 times longer than wide and with distal portion three times longer than basal part; the femur is longer than the total length of the prosoma (pro-, meso- and metapeltidium together) and longer than the patella; with a dorsal row of setae. Patella: subcylindrical, club-shaped, with distal portion twice as wide as the basal part; with more setae than the femur, with two dorsal and two ventral rows; 3.3 times longer than high. Tibia: cylindrical, straight, with slightly wider distal portion; shorter than half the length of the femur; base as high as patella, with at least two ventral, one lateral, two dorsal setae; tibia has the largest number of setae on the pedipalps. Tarsus: conical, approximately half the length of tibia, with numerous setae (dorsal and lateral smooth, and ventral plumose). Pedipalp tarsal spurs asymmetrical, as long as half for the length of tarsal claw; tarsal claw sharp and curved.
</paragraph>
<caption id="20CD66161260FFE100A2E7C36501A043" ID-DOI="http://doi.org/10.5281/zenodo.11547614" ID-Zenodo-Dep="11547614" httpUri="https://zenodo.org/record/11547614/files/figure.png" pageId="6" pageNumber="409" startId="6.[151,264,1442,1466]" targetBox="[237,1349,239,1415]" targetPageId="6" targetType="figure">
<paragraph id="740D369E1260FFE100A2E7C36501A043" blockId="6.[151,1436,1442,1538]" pageId="6" pageNumber="409">
<emphasis id="46C6EA8C1260FFE100A2E7C3602FA3F8" bold="true" box="[151,333,1442,1466]" pageId="6" pageNumber="409">FIGURES 20–26</emphasis>
.
<taxonomicName id="B3B24D1D1260FFE1016CE7C3609CA3F8" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[345,510,1442,1466]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1260FFE1016CE7C3609CA3F8" box="[345,510,1442,1466]" italics="true" pageId="6" pageNumber="409">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1260FFE10231E7C36336A3F8" bold="true" box="[516,596,1442,1466]" pageId="6" pageNumber="409">
<taxonomicNameLabel id="5DF557F71260FFE10231E7C36336A3F8" box="[516,596,1442,1466]" pageId="6" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="740D369E1260FFE100F2E42D645CA0B5" blockId="6.[151,1437,1612,2034]" pageId="6" pageNumber="409">
<emphasis id="46C6EA8C1260FFE100F2E42D6039A024" box="[199,347,1612,1638]" italics="true" pageId="6" pageNumber="409">Opisthosoma</emphasis>
(
<figureCitation id="EC892A1B1260FFE1015EE42D6369A024" box="[363,523,1612,1638]" captionStart="FIGURES 7–14" captionStartId="4.[151,264,1887,1911]" captionTargetBox="[151,1436,179,1861]" captionTargetId="figure-23@4.[151,1436,179,1861]" captionTargetPageId="4" captionText="FIGURES 7–14. Rowlandius ufpi sp. nov. heteromorphic male, holotype (CHNUFPI 4056), dorsal (7–8), ventral (9–10) and lateral (11–12) habitus; prolateral (13) and retrolateral (14) views of the right pedipalp. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547607" httpUri="https://zenodo.org/record/11547607/files/figure.png" pageId="6" pageNumber="409">Figs 8, 10, 12</figureCitation>
). Setae: Tergites I–IX each with a dorsal pair of large Dm setae. Tergites XI–XII telescopic. Segment VIII with a pair of dorso-submedian setae and a pair of Dl1 and a pair of Dl2 setae; IX with pairs Dl1 and Dl2, Vm1, Vm2 and pair Vl1 and Vl2 present. Segments X–XII same as IX. Segment XII with the medial Vm1 and Vm2 shorter than Vll; with postero-dorsal opisthosomal process short, subtriangular.;Sternites I and II with numerous small setae; sternites III–VII with medium setae; sternites VIII–XII with large setae.
</paragraph>
<paragraph id="740D369E1260FFE100F2E56165D5A1B0" blockId="6.[151,1437,1612,2034]" pageId="6" pageNumber="409">
<emphasis id="46C6EA8C1260FFE100F2E5616058A158" box="[199,314,1792,1818]" italics="true" pageId="6" pageNumber="409">Flagellum</emphasis>
(
<figureCitation id="EC892A1B1260FFE10178E56160B0A158" box="[333,466,1792,1818]" captionStart="FIGURES 15–19" captionStartId="5.[151,264,1577,1601]" captionTargetBox="[151,1436,423,1551]" captionTargetId="figure-148@5.[151,1436,414,1551]" captionTargetPageId="5" captionText="FIGURES 15–19. Rowlandius ufpi sp. nov., heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19)." figureDoi="http://doi.org/10.5281/zenodo.11547612" httpUri="https://zenodo.org/record/11547612/files/figure.png" pageId="6" pageNumber="409">Figs 17–19</figureCitation>
,
<figureCitation id="EC892A1B1260FFE101D7E561634CA158" box="[482,558,1792,1818]" captionStart="FIGURES 20–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[237,1349,239,1415]" captionTargetId="figure-56@6.[228,1359,234,1415]" captionTargetPageId="6" captionText="FIGURES 20–26. Rowlandius ufpi sp. nov., heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547614" httpUri="https://zenodo.org/record/11547614/files/figure.png" pageId="6" pageNumber="409">22–24</figureCitation>
). Setation Dm1, Dm4; Dl1, Dl3; Vm1, Vm2, Vm4, Vm5; Vl1, Vl2. Shape subquadrate with rounded and dorsoventrally flattened sides, with the posterior region slightly tapered. With three projections: a pair located in the dorso-submedian region (between the setae Dl1 and Vm5), without setae, separated by a depression and a projection positioned in the posteromedian region, connected to the lateral ones, with the setae Dm4 at its apex. Setation: Dm4 proximal of the posterior margin on a protuberance; Vm1 at the same level as Dm1; Vm4 proximal to Vl1 level; Dl1 at same level to Vl1; Vm5 proximal to Dm4; Vl2 proximal to Dl3 level. One pair of dorsolateral microsetae between Dm1 and Dl1; six lateral microsetae between Dl1 and Dl3.
</paragraph>
<paragraph id="740D369E1261FFE000F2E2F66589A727" blockId="7.[151,1437,150,501]" pageId="7" pageNumber="410">
<emphasis id="46C6EA8C1261FFE000F2E2F66399A6F2" box="[199,763,150,177]" italics="true" pageId="7" pageNumber="410">
Homeomorphic male (
<typeStatus id="AB09883C1261FFE001F4E2F6634BA6F3" box="[449,553,151,177]" pageId="7" pageNumber="410" type="paratype">paratype</typeStatus>
, CHNUFPI 0678).
</emphasis>
Chelicerae reddish and flagellum light brownish. Pedipalps: trochanter, femur, patella and tibia light reddish-brown; tarsus dark reddish-brown. Legs: coxae I–IV, sternum light brown; trochanters I reddish and II–IV greenish-brown; femur I reddish and II–IV greenish-brown; patellae I reddish and II–IV light greenish-brown; tibiae II–IV light brownish-brown, except for tibia I that is reddish; tarsus I light reddish and II–IV greenish-brown. Prosoma light brownish with propeltidium reddish-brown, meso and metapeltidium yellowish-brown, opisthosomal tergites greenish-brown and sternite light brownish.
</paragraph>
<paragraph id="740D369E1261FFE000F2E30E605CA7B7" blockId="7.[151,1437,150,501]" pageId="7" pageNumber="410">
<emphasis id="46C6EA8C1261FFE000F2E30E6049A7CA" box="[199,299,367,392]" italics="true" pageId="7" pageNumber="410">Prosoma</emphasis>
(
<figureCitation id="EC892A1B1261FFE0010EE30E60E6A7CB" box="[315,388,367,393]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="7" pageNumber="410">Figs 2</figureCitation>
,
<figureCitation id="EC892A1B1261FFE001A4E30E60B9A7CB" box="[401,475,367,393]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1939,1963]" captionTargetBox="[289,1299,522,1913]" captionTargetId="figure-157@7.[289,1299,522,1913]" captionTargetPageId="7" captionText="FIGURES 27–35. Rowlandius ufpi sp. nov. homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547616" httpUri="https://zenodo.org/record/11547616/files/figure.png" pageId="7" pageNumber="410">27, 29</figureCitation>
). 1.37 long, 0.62 wide. Anterior process of propeltidium with 2 setae (one behind the other) followed by 2 pairs of dorso-submedian setae transversally oriented. Propeltidium 1.8 times longer than high. metapeltidium entire. Chelicerae (
<figureCitation id="EC892A1B1261FFE0022DE3D663F9A793" box="[536,667,438,465]" captionStart="FIGURES 15–19" captionStartId="5.[151,264,1577,1601]" captionTargetBox="[151,1436,423,1551]" captionTargetId="figure-148@5.[151,1436,414,1551]" captionTargetPageId="5" captionText="FIGURES 15–19. Rowlandius ufpi sp. nov., heteromorphic male, holotype (CHNUFPI 4056), chelicerae (15, 16) and flagellum (17–19), prolateral (15, 16), dorsal (17), ventral (18) and lateral (19) views. Scale bars: 0.1 mm (15–16), 0.5 mm (17–19)." figureDoi="http://doi.org/10.5281/zenodo.11547612" httpUri="https://zenodo.org/record/11547612/files/figure.png" pageId="7" pageNumber="410">Figs 15–16</figureCitation>
), opisthosoma (
<figureCitation id="EC892A1B1261FFE0037BE3D662FAA793" box="[846,920,439,465]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="7" pageNumber="410">Figs 2</figureCitation>
,
<figureCitation id="EC892A1B1261FFE00393E3D6628CA793" box="[934,1006,439,465]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1939,1963]" captionTargetBox="[289,1299,522,1913]" captionTargetId="figure-157@7.[289,1299,522,1913]" captionTargetPageId="7" captionText="FIGURES 27–35. Rowlandius ufpi sp. nov. homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547616" httpUri="https://zenodo.org/record/11547616/files/figure.png" pageId="7" pageNumber="410">28, 30</figureCitation>
) and flagellum (
<figureCitation id="EC892A1B1261FFE0049EE3D6644EA793" box="[1195,1324,438,465]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1939,1963]" captionTargetBox="[289,1299,522,1913]" captionTargetId="figure-157@7.[289,1299,522,1913]" captionTargetPageId="7" captionText="FIGURES 27–35. Rowlandius ufpi sp. nov. homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547616" httpUri="https://zenodo.org/record/11547616/files/figure.png" pageId="7" pageNumber="410">Figs 33–35</figureCitation>
) as in the male
<typeStatus id="AB09883C1261FFE000E0E3BA6059A7B7" box="[213,315,475,501]" pageId="7" pageNumber="410" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="20CD66161261FFE000A2E5F26003A1B1" ID-DOI="http://doi.org/10.5281/zenodo.11547616" ID-Zenodo-Dep="11547616" httpUri="https://zenodo.org/record/11547616/files/figure.png" pageId="7" pageNumber="410" startId="7.[151,264,1939,1963]" targetBox="[289,1299,522,1913]" targetPageId="7" targetType="figure">
<paragraph id="740D369E1261FFE000A2E5F26003A1B1" blockId="7.[151,1437,1939,2035]" pageId="7" pageNumber="410">
<emphasis id="46C6EA8C1261FFE000A2E5F2602CA1E9" bold="true" box="[151,334,1939,1963]" pageId="7" pageNumber="410">FIGURES 27–35</emphasis>
.
<taxonomicName id="B3B24D1D1261FFE0016EE5F26363A1E9" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[347,513,1939,1963]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="410" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1261FFE0016EE5F26363A1E9" box="[347,513,1939,1963]" italics="true" pageId="7" pageNumber="410">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1261FFE0023DE5F2633BA1E9" bold="true" box="[520,601,1939,1963]" pageId="7" pageNumber="410">
<taxonomicNameLabel id="5DF557F71261FFE0023DE5F2633BA1E9" box="[520,601,1939,1963]" pageId="7" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="740D369E126EFFEF00F2E2F6604FA7B7" blockId="8.[151,1437,150,609]" pageId="8" pageNumber="411">
<emphasis id="46C6EA8C126EFFEF00F2E2F6604EA6F3" box="[199,300,151,177]" italics="true" pageId="8" pageNumber="411">Pedipalp</emphasis>
(
<figureCitation id="EC892A1B126EFFEF0108E2F660A3A6F3" box="[317,449,150,177]" captionStart="FIGURES 20–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[237,1349,239,1415]" captionTargetId="figure-56@6.[228,1359,234,1415]" captionTargetPageId="6" captionText="FIGURES 20–26. Rowlandius ufpi sp. nov., heteromorphic male (20–24) holotype (CHNUFPI 4056) and homeomorphic male (25–26) paratype (CHNUFPI 0678), right pedipalp (20–21; 25–26) and flagellum (22–24), schematic drawings in retrolateral (20, 25), prolateral (21, 26), ventral (22), dorsal (23) and lateral (24) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547614" httpUri="https://zenodo.org/record/11547614/files/figure.png" pageId="8" pageNumber="411">Figs 25–26</figureCitation>
,
<figureCitation id="EC892A1B126EFFEF01FAE2F6637BA6F3" box="[463,537,151,177]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1939,1963]" captionTargetBox="[289,1299,522,1913]" captionTargetId="figure-157@7.[289,1299,522,1913]" captionTargetPageId="7" captionText="FIGURES 27–35. Rowlandius ufpi sp. nov. homeomorphic male, paratype (CHNUFPI 0678), habitus (27–30), right pedipalp (31–32) and flagellum (33–35), in dorsal (27–28, 33) and ventral (29–30, 34), retrolateral (31, 35) and prolateral (32) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547616" httpUri="https://zenodo.org/record/11547616/files/figure.png" pageId="8" pageNumber="411">31–32</figureCitation>
). Dorsal and lateral setae of tarsus smooth, ventral setae plumose. Trochanter: with ten ventral setae; prolaterally with an acute mesal spur (MS). Femur: subcylindrical, 2.3 times longer than high; dorsal edge with a row of eight setae; retrolaterally with Fe1, Fv1, Fv2, Fe2; prolaterally with Fvr with setae 1, 2 and 3, and with Fd1, Fd2. Patella: cylindrical, straight, 3.3 times longer than high, with 1 row of dorsal setae, 1 row on lateral and 2 rows on ventral; prolaterally with Pm1, Pm3, Pm4 and Pm5, surrounded by small pores; retrolaterally with Pe2, Pe3 and Pe5. Tibia: cylindrical, straight, 3 times longer than high, base as high as patella, with at least 2 ventral, 1 lateral, 2 dorsal setae; Ter with setae 4, 5 and 6; Tir with setae 1, 2, 3 and 4; Tmr with setae 2, 3 and 4; one other Tm1 setae; Tarsus: conical, with asymmetrical spurs (prolateral and retrolateral tarsal spurs: pTS/rTS); approximately half the length of tibia, with numerous setae. Tarsal claw sharp and curved, smaller than half the tarsus length.
</paragraph>
<paragraph id="740D369E126EFFEF00F2E39E6302A423" blockId="8.[151,1437,150,609]" pageId="8" pageNumber="411">
<emphasis id="46C6EA8C126EFFEF00F2E39E6308A45A" box="[199,618,510,537]" italics="true" pageId="8" pageNumber="411">
Female (
<typeStatus id="AB09883C126EFFEF011BE39E60F4A45B" box="[302,406,511,537]" pageId="8" pageNumber="411" type="paratype">paratype</typeStatus>
, CHNUFPI 4064)
</emphasis>
. Coloration and setation as in the male
<typeStatus id="AB09883C126EFFEF0471E39E65C8A45B" box="[1092,1194,511,537]" pageId="8" pageNumber="411" type="holotype">holotype</typeStatus>
. Propeltidium longer than wide (
<tableCitation id="39300325126EFFEF0126E0426005A47F" box="[275,359,547,573]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Measurements (in mm) of Rowlandius ufpi sp. nov." httpUri="http://table.plazi.org/id/20CD6616126FFFEE00A2E2F6625AA6F2" pageId="8" pageNumber="411" tableUuid="20CD6616126FFFEE00A2E2F6625AA6F2">Table 1</tableCitation>
). Prosoma (
<figureCitation id="EC892A1B126EFFEF01C5E042635AA47F" box="[496,568,547,573]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="8" pageNumber="411">Figs 3</figureCitation>
,
<figureCitation id="EC892A1B126EFFEF0270E04263ECA47F" box="[581,654,547,573]" captionStart="FIGURES 36–42" captionStartId="8.[151,264,1911,1935]" captionTargetBox="[151,1436,630,1885]" captionTargetId="figure-246@8.[151,1436,630,1885]" captionTargetPageId="8" captionText="FIGURES 36–42. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4064), habitus (36–39) and flagellum (40–42), in dorsal (36–37, 40) and ventral (38–39, 41) and retrolateral (42) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547618" httpUri="https://zenodo.org/record/11547618/files/figure.png" pageId="8" pageNumber="411">36, 38</figureCitation>
), chelicerae and opisthosoma (
<figureCitation id="EC892A1B126EFFEF03D3E042654CA47F" box="[998,1070,547,573]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="8" pageNumber="411">Figs 3</figureCitation>
,
<figureCitation id="EC892A1B126EFFEF040EE04265E0A47F" box="[1083,1154,547,573]" captionStart="FIGURES 36–42" captionStartId="8.[151,264,1911,1935]" captionTargetBox="[151,1436,630,1885]" captionTargetId="figure-246@8.[151,1436,630,1885]" captionTargetPageId="8" captionText="FIGURES 36–42. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4064), habitus (36–39) and flagellum (40–42), in dorsal (36–37, 40) and ventral (38–39, 41) and retrolateral (42) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547618" httpUri="https://zenodo.org/record/11547618/files/figure.png" pageId="8" pageNumber="411">37, 39</figureCitation>
) as in the male
<typeStatus id="AB09883C126EFFEF0506E04264FBA47F" box="[1331,1433,547,573]" pageId="8" pageNumber="411" type="holotype">holotype</typeStatus>
. Pedipalp similar to homeomorphic male.
</paragraph>
<caption id="20CD6616126EFFEF00A2E516654CA1F1" ID-DOI="http://doi.org/10.5281/zenodo.11547618" ID-Zenodo-Dep="11547618" httpUri="https://zenodo.org/record/11547618/files/figure.png" pageId="8" pageNumber="411" startId="8.[151,264,1911,1935]" targetBox="[151,1436,630,1885]" targetPageId="8" targetType="figure">
<paragraph id="740D369E126EFFEF00A2E516654CA1F1" blockId="8.[151,1437,1911,1971]" pageId="8" pageNumber="411">
<emphasis id="46C6EA8C126EFFEF00A2E5166032A1CD" bold="true" box="[151,336,1911,1935]" pageId="8" pageNumber="411">FIGURES 36–42</emphasis>
.
<taxonomicName id="B3B24D1D126EFFEF016AE5166364A1CD" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[351,518,1911,1935]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="411" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126EFFEF016AE5166364A1CD" box="[351,518,1911,1935]" italics="true" pageId="8" pageNumber="411">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126EFFEF023AE5166300A1CD" bold="true" box="[527,610,1911,1935]" pageId="8" pageNumber="411">
<taxonomicNameLabel id="5DF557F7126EFFEF023AE5166300A1CD" box="[527,610,1911,1935]" pageId="8" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, paratype (CHNUFPI 4064), habitus (36–39) and flagellum (40–42), in dorsal (36–37, 40) and ventral (38–39, 41) and retrolateral (42) views. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="20CD6616126FFFEE00A2E2F6625AA6F2" ID-Table-UUID="20CD6616126FFFEE00A2E2F6625AA6F2" box="[151,824,151,176]" httpUri="http://table.plazi.org/id/20CD6616126FFFEE00A2E2F6625AA6F2" pageId="9" pageNumber="412" startId="9.[151,239,151,175]" targetBox="[159,1388,191,1991]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="740D369E126FFFEE00A2E2F6625AA6F2" blockId="9.[151,824,151,176]" box="[151,824,151,176]" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE00A2E2F66068A6ED" bold="true" box="[151,266,151,175]" pageId="9" pageNumber="412">TABLE 1.</emphasis>
Measurements (in mm) of
<taxonomicName id="B3B24D1D126FFFEE0218E2F663B9A6F2" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[557,731,151,176]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="9" pageNumber="412" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126FFFEE0218E2F663B9A6F2" box="[557,731,151,176]" italics="true" pageId="9" pageNumber="412">Rowlandius ufpi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF557F7126FFFEE02D7E2F9625AA6F2" box="[738,824,152,176]" pageId="9" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="740D369E126FFFEE00AAE2DE6409A185" pageId="9" pageNumber="412">
<table id="06B2C43E126F001800AAE2DE640EA185" box="[159,1388,191,1991]" gridcols="6" gridrows="50" pageId="9" pageNumber="412">
<tr id="CA8234DC126F001800AAE2DE640EA697" box="[159,1388,191,213]" gridrow="0" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE2DE600CA697" box="[159,366,191,213]" gridcol="0" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE00AAE2DE619AA696" bold="true" box="[159,248,191,212]" pageId="9" pageNumber="412">Measure</emphasis>
</th>
<th id="89535DA0126F00180192E2DE6338A697" box="[423,602,191,213]" gridcol="1" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE0198E2DE6336A697" bold="true" box="[429,596,191,213]" pageId="9" pageNumber="412">Male-α hOlOtYpe</emphasis>
</th>
<th id="89535DA0126F001802A4E2DE6227A697" box="[657,837,191,213]" gridcol="2" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE02A1E2DE6223A696" bold="true" box="[660,833,191,212]" pageId="9" pageNumber="412">Female paratype</emphasis>
</th>
<th id="89535DA0126F001803BBE2DE628DA697" box="[910,1007,191,213]" gridcol="3" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE03A0E2DE628AA697" bold="true" box="[917,1000,191,213]" pageId="9" pageNumber="412">MaleS-α</emphasis>
</th>
<th id="89535DA0126F0018047EE2DE65C9A697" box="[1099,1195,191,213]" gridcol="4" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE047AE2DE65C5A696" bold="true" box="[1103,1191,191,213]" pageId="9" pageNumber="412">MaleS- β</emphasis>
</th>
<th id="89535DA0126F0018053EE2DE640EA697" box="[1291,1388,191,213]" gridcol="5" gridrow="0" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE0524E2DE6407A696" bold="true" box="[1297,1381,191,212]" pageId="9" pageNumber="412">Females</emphasis>
</th>
</tr>
<tr id="CA8234DC126F001800AAE283640EA6BB" box="[159,1388,226,249]" gridrow="1" pageId="9" pageNumber="412" rowspan-0="1">
<td id="89535DA0126F00180192E2836338A6BB" box="[423,602,226,249]" gridcol="1" gridrow="1" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE0192E2836338A6BA" bold="true" box="[423,602,226,248]" pageId="9" pageNumber="412">(CHNUFPI 4056)</emphasis>
</td>
<td id="89535DA0126F001802A4E2836227A6BB" box="[657,837,226,249]" gridcol="2" gridrow="1" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE02A4E2836227A6BA" bold="true" box="[657,837,226,248]" pageId="9" pageNumber="412">(CHNUFPI 4064)</emphasis>
</td>
<td id="89535DA0126F001803BBE283628DA6BB" box="[910,1007,226,249]" gridcol="3" gridrow="1" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE03A8E2826282A6BB" bold="true" box="[925,992,227,249]" pageId="9" pageNumber="412">(n = 5)</emphasis>
</td>
<td id="89535DA0126F0018047EE28365C9A6BB" box="[1099,1195,226,249]" gridcol="4" gridrow="1" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE046CE28265FEA6BB" bold="true" box="[1113,1180,227,249]" pageId="9" pageNumber="412">(n = 5)</emphasis>
</td>
<td id="89535DA0126F0018053EE283640EA6BB" box="[1291,1388,226,249]" gridcol="5" gridrow="1" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE052FE282643EA6BB" bold="true" box="[1306,1372,227,249]" pageId="9" pageNumber="412">(n = 5)</emphasis>
</td>
</tr>
<tr id="CA8234DC126F001800AAE366640EA75F" box="[159,1388,263,285]" gridrow="2" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE366600CA75F" box="[159,366,263,285]" gridcol="0" gridrow="2" pageId="9" pageNumber="412">
<emphasis id="46C6EA8C126FFFEE00AAE366607BA75E" bold="true" box="[159,281,263,284]" pageId="9" pageNumber="412">Total length</emphasis>
</th>
<td id="89535DA0126F00180192E3666338A75F" box="[423,602,263,285]" gridcol="1" gridrow="2" pageId="9" pageNumber="412">3.45</td>
<td id="89535DA0126F001802A4E3666227A75F" box="[657,837,263,285]" gridcol="2" gridrow="2" pageId="9" pageNumber="412">4.6</td>
<td id="89535DA0126F001803BBE366628DA75F" box="[910,1007,263,285]" gridcol="3" gridrow="2" pageId="9" pageNumber="412">2.54–3.6</td>
<td id="89535DA0126F0018047EE36665C9A75F" box="[1099,1195,263,285]" gridcol="4" gridrow="2" pageId="9" pageNumber="412">3.02–3.6</td>
<td id="89535DA0126F0018053EE366640EA75F" box="[1291,1388,263,285]" gridcol="5" gridrow="2" pageId="9" pageNumber="412">3.44–5</td>
</tr>
<tr id="CA8234DC126F001800AAE34D640EA700" box="[159,1388,300,322]" gridrow="3" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE34D600CA700" box="[159,366,300,322]" gridcol="0" gridrow="3" pageId="9" pageNumber="412">Prosoma length</th>
<td id="89535DA0126F00180192E34D6338A700" box="[423,602,300,322]" gridcol="1" gridrow="3" pageId="9" pageNumber="412">1.5</td>
<td id="89535DA0126F001802A4E34D6227A700" box="[657,837,300,322]" gridcol="2" gridrow="3" pageId="9" pageNumber="412">1.3</td>
<td id="89535DA0126F001803BBE34D628DA700" box="[910,1007,300,322]" gridcol="3" gridrow="3" pageId="9" pageNumber="412">1.14–1.67</td>
<td id="89535DA0126F0018047EE34D65C9A700" box="[1099,1195,300,322]" gridcol="4" gridrow="3" pageId="9" pageNumber="412">1.33–1.5</td>
<td id="89535DA0126F0018053EE34D640EA700" box="[1291,1388,300,322]" gridcol="5" gridrow="3" pageId="9" pageNumber="412">1.47–1.9</td>
</tr>
<tr id="CA8234DC126F001800AAE331640EA724" box="[159,1388,336,358]" gridrow="4" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE331600CA724" box="[159,366,336,358]" gridcol="0" gridrow="4" pageId="9" pageNumber="412">Propeltidium length</th>
<td id="89535DA0126F00180192E3316338A724" box="[423,602,336,358]" gridcol="1" gridrow="4" pageId="9" pageNumber="412">1.25</td>
<td id="89535DA0126F001802A4E3316227A724" box="[657,837,336,358]" gridcol="2" gridrow="4" pageId="9" pageNumber="412">1.25</td>
<td id="89535DA0126F001803BBE331628DA724" box="[910,1007,336,358]" gridcol="3" gridrow="4" pageId="9" pageNumber="412">0.96–1.35</td>
<td id="89535DA0126F0018047EE33165C9A724" box="[1099,1195,336,358]" gridcol="4" gridrow="4" pageId="9" pageNumber="412">1.15–1.2</td>
<td id="89535DA0126F0018053EE331640EA724" box="[1291,1388,336,358]" gridcol="5" gridrow="4" pageId="9" pageNumber="412">1.17–1.37</td>
</tr>
<tr id="CA8234DC126F001800AAE315640EA7C8" box="[159,1388,372,394]" gridrow="5" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE315600CA7C8" box="[159,366,372,394]" gridcol="0" gridrow="5" pageId="9" pageNumber="412">Propeltidium width</th>
<td id="89535DA0126F00180192E3156338A7C8" box="[423,602,372,394]" gridcol="1" gridrow="5" pageId="9" pageNumber="412">0.6</td>
<td id="89535DA0126F001802A4E3156227A7C8" box="[657,837,372,394]" gridcol="2" gridrow="5" pageId="9" pageNumber="412">0.67</td>
<td id="89535DA0126F001803BBE315628DA7C8" box="[910,1007,372,394]" gridcol="3" gridrow="5" pageId="9" pageNumber="412">0.54–0.75</td>
<td id="89535DA0126F0018047EE31565C9A7C8" box="[1099,1195,372,394]" gridcol="4" gridrow="5" pageId="9" pageNumber="412">0.6–0.62</td>
<td id="89535DA0126F0018053EE315640EA7C8" box="[1291,1388,372,394]" gridcol="5" gridrow="5" pageId="9" pageNumber="412">0.75–0.82</td>
</tr>
<tr id="CA8234DC126F001800AAE3F9640EA7ED" box="[159,1388,408,431]" gridrow="6" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE3F9600CA7ED" box="[159,366,408,431]" gridcol="0" gridrow="6" pageId="9" pageNumber="412">Metapeltidium length</th>
<td id="89535DA0126F00180192E3F96338A7ED" box="[423,602,408,431]" gridcol="1" gridrow="6" pageId="9" pageNumber="412">0.02</td>
<td id="89535DA0126F001802A4E3F96227A7ED" box="[657,837,408,431]" gridcol="2" gridrow="6" pageId="9" pageNumber="412">0.25</td>
<td id="89535DA0126F001803BBE3F9628DA7ED" box="[910,1007,408,431]" gridcol="3" gridrow="6" pageId="9" pageNumber="412">0.02–0.25</td>
<td id="89535DA0126F0018047EE3F965C9A7ED" box="[1099,1195,408,431]" gridcol="4" gridrow="6" pageId="9" pageNumber="412">0.01–0.2</td>
<td id="89535DA0126F0018053EE3F9640EA7ED" box="[1291,1388,408,431]" gridcol="5" gridrow="6" pageId="9" pageNumber="412">0.02–0.37</td>
</tr>
<tr id="CA8234DC126F001800AAE3DC640EA791" box="[159,1388,445,467]" gridrow="7" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE3DC600CA791" box="[159,366,445,467]" gridcol="0" gridrow="7" pageId="9" pageNumber="412">Metapeltidium width</th>
<td id="89535DA0126F00180192E3DC6338A791" box="[423,602,445,467]" gridcol="1" gridrow="7" pageId="9" pageNumber="412">0.65</td>
<td id="89535DA0126F001802A4E3DC6227A791" box="[657,837,445,467]" gridcol="2" gridrow="7" pageId="9" pageNumber="412">0.7</td>
<td id="89535DA0126F001803BBE3DC628DA791" box="[910,1007,445,467]" gridcol="3" gridrow="7" pageId="9" pageNumber="412">0.6–0.8</td>
<td id="89535DA0126F0018047EE3DC65C9A791" box="[1099,1195,445,467]" gridcol="4" gridrow="7" pageId="9" pageNumber="412">0.62–0.75</td>
<td id="89535DA0126F0018053EE3DC640EA791" box="[1291,1388,445,467]" gridcol="5" gridrow="7" pageId="9" pageNumber="412">0.7–0.8</td>
</tr>
<tr id="CA8234DC126F001800AAE380640EA7B5" box="[159,1388,481,503]" gridrow="8" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE380600CA7B5" box="[159,366,481,503]" gridcol="0" gridrow="8" pageId="9" pageNumber="412">Opisthosoma length</th>
<td id="89535DA0126F00180192E3806338A7B5" box="[423,602,481,503]" gridcol="1" gridrow="8" pageId="9" pageNumber="412">1.95</td>
<td id="89535DA0126F001802A4E3806227A7B5" box="[657,837,481,503]" gridcol="2" gridrow="8" pageId="9" pageNumber="412">3.3</td>
<td id="89535DA0126F001803BBE380628DA7B5" box="[910,1007,481,503]" gridcol="3" gridrow="8" pageId="9" pageNumber="412">1.4–3.3</td>
<td id="89535DA0126F0018047EE38065C9A7B5" box="[1099,1195,481,503]" gridcol="4" gridrow="8" pageId="9" pageNumber="412">1.67–2.1</td>
<td id="89535DA0126F0018053EE380640EA7B5" box="[1291,1388,481,503]" gridcol="5" gridrow="8" pageId="9" pageNumber="412">1.95–3.1</td>
</tr>
<tr id="CA8234DC126F001800AAE064640EA459" box="[159,1388,517,539]" gridrow="9" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE064600CA459" box="[159,366,517,539]" gridcol="0" gridrow="9" pageId="9" pageNumber="412">Opisthosoma width</th>
<td id="89535DA0126F00180192E0646338A459" box="[423,602,517,539]" gridcol="1" gridrow="9" pageId="9" pageNumber="412">0.87</td>
<td id="89535DA0126F001802A4E0646227A459" box="[657,837,517,539]" gridcol="2" gridrow="9" pageId="9" pageNumber="412">1.1</td>
<td id="89535DA0126F001803BBE064628DA459" box="[910,1007,517,539]" gridcol="3" gridrow="9" pageId="9" pageNumber="412">0.74–1.1</td>
<td id="89535DA0126F0018047EE06465C9A459" box="[1099,1195,517,539]" gridcol="4" gridrow="9" pageId="9" pageNumber="412">0.77–1.05</td>
<td id="89535DA0126F0018053EE064640EA459" box="[1291,1388,517,539]" gridcol="5" gridrow="9" pageId="9" pageNumber="412">0.87–1.2</td>
</tr>
<tr id="CA8234DC126F001800AAE04B640EA402" box="[159,1388,554,576]" gridrow="10" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE04B600CA402" box="[159,366,554,576]" gridcol="0" gridrow="10" pageId="9" pageNumber="412">Flagellum length</th>
<td id="89535DA0126F00180192E04B6338A402" box="[423,602,554,576]" gridcol="1" gridrow="10" pageId="9" pageNumber="412">0.42</td>
<td id="89535DA0126F001802A4E04B6227A402" box="[657,837,554,576]" gridcol="2" gridrow="10" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001803BBE04B628DA402" box="[910,1007,554,576]" gridcol="3" gridrow="10" pageId="9" pageNumber="412">0.42–0.47</td>
<td id="89535DA0126F0018047EE04B65C9A402" box="[1099,1195,554,576]" gridcol="4" gridrow="10" pageId="9" pageNumber="412">0.42–0.45</td>
<td id="89535DA0126F0018053EE04B640EA402" box="[1291,1388,554,576]" gridcol="5" gridrow="10" pageId="9" pageNumber="412">0.32–0.37</td>
</tr>
<tr id="CA8234DC126F001800AAE02F640EA426" box="[159,1388,590,612]" gridrow="11" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE02F600CA426" box="[159,366,590,612]" gridcol="0" gridrow="11" pageId="9" pageNumber="412">Flagellum width</th>
<td id="89535DA0126F00180192E02F6338A426" box="[423,602,590,612]" gridcol="1" gridrow="11" pageId="9" pageNumber="412">0.32</td>
<td id="89535DA0126F001802A4E02F6227A426" box="[657,837,590,612]" gridcol="2" gridrow="11" pageId="9" pageNumber="412">0.01</td>
<td id="89535DA0126F001803BBE02F628DA426" box="[910,1007,590,612]" gridcol="3" gridrow="11" pageId="9" pageNumber="412">0.01–0.32</td>
<td id="89535DA0126F0018047EE02F65C9A426" box="[1099,1195,590,612]" gridcol="4" gridrow="11" pageId="9" pageNumber="412">0.3–0.32</td>
<td id="89535DA0126F0018053EE02F640EA426" box="[1291,1388,590,612]" gridcol="5" gridrow="11" pageId="9" pageNumber="412">0.01–0.07</td>
</tr>
<tr id="CA8234DC126F001800AAE013640EA4CA" box="[159,1388,626,648]" gridrow="12" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE013640EA4CA" box="[159,1388,626,648]" colspan="6" colspanRight="5" gridcol="0" gridrow="12" pageId="9" pageNumber="412">Pedipalp length</th>
</tr>
<tr id="CA8234DC126F001800AAE0F6640EA4EF" box="[159,1388,663,685]" gridrow="13" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE0F6600CA4EF" box="[159,366,663,685]" gridcol="0" gridrow="13" pageId="9" pageNumber="412">trochanter</th>
<td id="89535DA0126F00180192E0F66338A4EF" box="[423,602,663,685]" gridcol="1" gridrow="13" pageId="9" pageNumber="412">0.83</td>
<td id="89535DA0126F001802A4E0F66227A4EF" box="[657,837,663,685]" gridcol="2" gridrow="13" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE0F6628DA4EF" box="[910,1007,663,685]" gridcol="3" gridrow="13" pageId="9" pageNumber="412">0.37–0.85</td>
<td id="89535DA0126F0018047EE0F665C9A4EF" box="[1099,1195,663,685]" gridcol="4" gridrow="13" pageId="9" pageNumber="412">0.35–0.48</td>
<td id="89535DA0126F0018053EE0F6640EA4EF" box="[1291,1388,663,685]" gridcol="5" gridrow="13" pageId="9" pageNumber="412">0.42–0.62</td>
</tr>
<tr id="CA8234DC126F001800AAE0DA640EA493" box="[159,1388,699,721]" gridrow="14" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE0DA600CA493" box="[159,366,699,721]" gridcol="0" gridrow="14" pageId="9" pageNumber="412">femur</th>
<td id="89535DA0126F00180192E0DA6338A493" box="[423,602,699,721]" gridcol="1" gridrow="14" pageId="9" pageNumber="412">1.65</td>
<td id="89535DA0126F001802A4E0DA6227A493" box="[657,837,699,721]" gridcol="2" gridrow="14" pageId="9" pageNumber="412">0.4</td>
<td id="89535DA0126F001803BBE0DA628DA493" box="[910,1007,699,721]" gridcol="3" gridrow="14" pageId="9" pageNumber="412">0.87–1.7</td>
<td id="89535DA0126F0018047EE0DA65C9A493" box="[1099,1195,699,721]" gridcol="4" gridrow="14" pageId="9" pageNumber="412">0.47–0.6</td>
<td id="89535DA0126F0018053EE0DA640EA493" box="[1291,1388,699,721]" gridcol="5" gridrow="14" pageId="9" pageNumber="412">0.5–0.75</td>
</tr>
<tr id="CA8234DC126F001800AAE0BE640EA4B7" box="[159,1388,735,757]" gridrow="15" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE0BE600CA4B7" box="[159,366,735,757]" gridcol="0" gridrow="15" pageId="9" pageNumber="412">patella</th>
<td id="89535DA0126F00180192E0BE6338A4B7" box="[423,602,735,757]" gridcol="1" gridrow="15" pageId="9" pageNumber="412">1.55</td>
<td id="89535DA0126F001802A4E0BE6227A4B7" box="[657,837,735,757]" gridcol="2" gridrow="15" pageId="9" pageNumber="412">0.5</td>
<td id="89535DA0126F001803BBE0BE628DA4B7" box="[910,1007,735,757]" gridcol="3" gridrow="15" pageId="9" pageNumber="412">1.2–1.95</td>
<td id="89535DA0126F0018047EE0BE65C9A4B7" box="[1099,1195,735,757]" gridcol="4" gridrow="15" pageId="9" pageNumber="412">0.6–0.75</td>
<td id="89535DA0126F0018053EE0BE640EA4B7" box="[1291,1388,735,757]" gridcol="5" gridrow="15" pageId="9" pageNumber="412">0.47–0.65</td>
</tr>
<tr id="CA8234DC126F001800AAE162640EA558" box="[159,1388,771,794]" gridrow="16" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE162600CA558" box="[159,366,771,794]" gridcol="0" gridrow="16" pageId="9" pageNumber="412">tibia</th>
<td id="89535DA0126F00180192E1626338A558" box="[423,602,771,794]" gridcol="1" gridrow="16" pageId="9" pageNumber="412">0.82</td>
<td id="89535DA0126F001802A4E1626227A558" box="[657,837,771,794]" gridcol="2" gridrow="16" pageId="9" pageNumber="412">0.42</td>
<td id="89535DA0126F001803BBE162628DA558" box="[910,1007,771,794]" gridcol="3" gridrow="16" pageId="9" pageNumber="412">0.6–1.8</td>
<td id="89535DA0126F0018047EE16265C9A558" box="[1099,1195,771,794]" gridcol="4" gridrow="16" pageId="9" pageNumber="412">0.37–0.55</td>
<td id="89535DA0126F0018053EE162640EA558" box="[1291,1388,771,794]" gridcol="5" gridrow="16" pageId="9" pageNumber="412">0.45–0.55</td>
</tr>
<tr id="CA8234DC126F001800AAE149640EA57C" box="[159,1388,808,830]" gridrow="17" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE149600CA57C" box="[159,366,808,830]" gridcol="0" gridrow="17" pageId="9" pageNumber="412">tarsus</th>
<td id="89535DA0126F00180192E1496338A57C" box="[423,602,808,830]" gridcol="1" gridrow="17" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001802A4E1496227A57C" box="[657,837,808,830]" gridcol="2" gridrow="17" pageId="9" pageNumber="412">0.27</td>
<td id="89535DA0126F001803BBE149628DA57C" box="[910,1007,808,830]" gridcol="3" gridrow="17" pageId="9" pageNumber="412">0.3–0.47</td>
<td id="89535DA0126F0018047EE14965C9A57C" box="[1099,1195,808,830]" gridcol="4" gridrow="17" pageId="9" pageNumber="412">0.25–0.27</td>
<td id="89535DA0126F0018053EE149640EA57C" box="[1291,1388,808,830]" gridcol="5" gridrow="17" pageId="9" pageNumber="412">0.27–0.35</td>
</tr>
<tr id="CA8234DC126F001800AAE12D640EA520" box="[159,1388,844,866]" gridrow="18" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE12D640EA520" box="[159,1388,844,866]" colspan="6" colspanRight="5" gridcol="0" gridrow="18" pageId="9" pageNumber="412">Leg I length</th>
</tr>
<tr id="CA8234DC126F001800AAE111640EA5C4" box="[159,1388,880,902]" gridrow="19" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE111600CA5C4" box="[159,366,880,902]" gridcol="0" gridrow="19" pageId="9" pageNumber="412">coxa</th>
<td id="89535DA0126F00180192E1116338A5C4" box="[423,602,880,902]" gridcol="1" gridrow="19" pageId="9" pageNumber="412">0.55</td>
<td id="89535DA0126F001802A4E1116227A5C4" box="[657,837,880,902]" gridcol="2" gridrow="19" pageId="9" pageNumber="412">0.55</td>
<td id="89535DA0126F001803BBE111628DA5C4" box="[910,1007,880,902]" gridcol="3" gridrow="19" pageId="9" pageNumber="412">0.5–0.58</td>
<td id="89535DA0126F0018047EE11165C9A5C4" box="[1099,1195,880,902]" gridcol="4" gridrow="19" pageId="9" pageNumber="412">0.52–0.55</td>
<td id="89535DA0126F0018053EE111640EA5C4" box="[1291,1388,880,902]" gridcol="5" gridrow="19" pageId="9" pageNumber="412">0.5–0.62</td>
</tr>
<tr id="CA8234DC126F001800AAE1F4640EA5E9" box="[159,1388,917,939]" gridrow="20" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE1F4600CA5E9" box="[159,366,917,939]" gridcol="0" gridrow="20" pageId="9" pageNumber="412">trochanter</th>
<td id="89535DA0126F00180192E1F46338A5E9" box="[423,602,917,939]" gridcol="1" gridrow="20" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001802A4E1F46227A5E9" box="[657,837,917,939]" gridcol="2" gridrow="20" pageId="9" pageNumber="412">0.3</td>
<td id="89535DA0126F001803BBE1F4628DA5E9" box="[910,1007,917,939]" gridcol="3" gridrow="20" pageId="9" pageNumber="412">0.3–0.48</td>
<td id="89535DA0126F0018047EE1F465C9A5E9" box="[1099,1195,917,939]" gridcol="4" gridrow="20" pageId="9" pageNumber="412">0.3–0.37</td>
<td id="89535DA0126F0018053EE1F4640EA5E9" box="[1291,1388,917,939]" gridcol="5" gridrow="20" pageId="9" pageNumber="412">0.25–0.37</td>
</tr>
<tr id="CA8234DC126F001800AAE1D8640EA58D" box="[159,1388,953,975]" gridrow="21" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE1D8600CA58D" box="[159,366,953,975]" gridcol="0" gridrow="21" pageId="9" pageNumber="412">femur</th>
<td id="89535DA0126F00180192E1D86338A58D" box="[423,602,953,975]" gridcol="1" gridrow="21" pageId="9" pageNumber="412">1.25</td>
<td id="89535DA0126F001802A4E1D86227A58D" box="[657,837,953,975]" gridcol="2" gridrow="21" pageId="9" pageNumber="412">1.02</td>
<td id="89535DA0126F001803BBE1D8628DA58D" box="[910,1007,953,975]" gridcol="3" gridrow="21" pageId="9" pageNumber="412">1.02–1.35</td>
<td id="89535DA0126F0018047EE1D865C9A58D" box="[1099,1195,953,975]" gridcol="4" gridrow="21" pageId="9" pageNumber="412">1.07–1.12</td>
<td id="89535DA0126F0018053EE1D8640EA58D" box="[1291,1388,953,975]" gridcol="5" gridrow="21" pageId="9" pageNumber="412">1.02–1.22</td>
</tr>
<tr id="CA8234DC126F001800AAE1BC640EA5B1" box="[159,1388,989,1011]" gridrow="22" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE1BC600CA5B1" box="[159,366,989,1011]" gridcol="0" gridrow="22" pageId="9" pageNumber="412">patella</th>
<td id="89535DA0126F00180192E1BC6338A5B1" box="[423,602,989,1011]" gridcol="1" gridrow="22" pageId="9" pageNumber="412">1.45</td>
<td id="89535DA0126F001802A4E1BC6227A5B1" box="[657,837,989,1011]" gridcol="2" gridrow="22" pageId="9" pageNumber="412">1.12</td>
<td id="89535DA0126F001803BBE1BC628DA5B1" box="[910,1007,989,1011]" gridcol="3" gridrow="22" pageId="9" pageNumber="412">1.12–1.57</td>
<td id="89535DA0126F0018047EE1BC65C9A5B1" box="[1099,1195,989,1011]" gridcol="4" gridrow="22" pageId="9" pageNumber="412">1.17–1.5</td>
<td id="89535DA0126F0018053EE1BC640EA5B1" box="[1291,1388,989,1011]" gridcol="5" gridrow="22" pageId="9" pageNumber="412">1–1.47</td>
</tr>
<tr id="CA8234DC126F001800AAE660640EA25A" box="[159,1388,1025,1048]" gridrow="23" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE660600CA25A" box="[159,366,1025,1048]" gridcol="0" gridrow="23" pageId="9" pageNumber="412">tibia</th>
<td id="89535DA0126F00180192E6606338A25A" box="[423,602,1025,1048]" gridcol="1" gridrow="23" pageId="9" pageNumber="412">1.05</td>
<td id="89535DA0126F001802A4E6606227A25A" box="[657,837,1025,1048]" gridcol="2" gridrow="23" pageId="9" pageNumber="412">0.87</td>
<td id="89535DA0126F001803BBE660628DA25A" box="[910,1007,1025,1048]" gridcol="3" gridrow="23" pageId="9" pageNumber="412">0.82–1.06</td>
<td id="89535DA0126F0018047EE66065C9A25A" box="[1099,1195,1025,1048]" gridcol="4" gridrow="23" pageId="9" pageNumber="412">0.92–1.12</td>
<td id="89535DA0126F0018053EE660640EA25A" box="[1291,1388,1025,1048]" gridcol="5" gridrow="23" pageId="9" pageNumber="412">0.95–1.02</td>
</tr>
<tr id="CA8234DC126F001800AAE647640EA27E" box="[159,1388,1062,1084]" gridrow="24" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE647600CA27E" box="[159,366,1062,1084]" gridcol="0" gridrow="24" pageId="9" pageNumber="412">basitarsus</th>
<td id="89535DA0126F00180192E6476338A27E" box="[423,602,1062,1084]" gridcol="1" gridrow="24" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001802A4E6476227A27E" box="[657,837,1062,1084]" gridcol="2" gridrow="24" pageId="9" pageNumber="412">0.32</td>
<td id="89535DA0126F001803BBE647628DA27E" box="[910,1007,1062,1084]" gridcol="3" gridrow="24" pageId="9" pageNumber="412">0.3–0.38</td>
<td id="89535DA0126F0018047EE64765C9A27E" box="[1099,1195,1062,1084]" gridcol="4" gridrow="24" pageId="9" pageNumber="412">0.32–0.37</td>
<td id="89535DA0126F0018053EE647640EA27E" box="[1291,1388,1062,1084]" gridcol="5" gridrow="24" pageId="9" pageNumber="412">0.3–0.35</td>
</tr>
<tr id="CA8234DC126F001800AAE62B640EA222" box="[159,1388,1098,1120]" gridrow="25" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE62B600CA222" box="[159,366,1098,1120]" gridcol="0" gridrow="25" pageId="9" pageNumber="412">telotarsus</th>
<td id="89535DA0126F00180192E62B6338A222" box="[423,602,1098,1120]" gridcol="1" gridrow="25" pageId="9" pageNumber="412">0.67</td>
<td id="89535DA0126F001802A4E62B6227A222" box="[657,837,1098,1120]" gridcol="2" gridrow="25" pageId="9" pageNumber="412">0.52</td>
<td id="89535DA0126F001803BBE62B628DA222" box="[910,1007,1098,1120]" gridcol="3" gridrow="25" pageId="9" pageNumber="412">0.5–0.67</td>
<td id="89535DA0126F0018047EE62B65C9A222" box="[1099,1195,1098,1120]" gridcol="4" gridrow="25" pageId="9" pageNumber="412">0.4–0.87</td>
<td id="89535DA0126F0018053EE62B640EA222" box="[1291,1388,1098,1120]" gridcol="5" gridrow="25" pageId="9" pageNumber="412">0.52–0.6</td>
</tr>
<tr id="CA8234DC126F001800AAE60F640EA2C6" box="[159,1388,1134,1156]" gridrow="26" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE60F640EA2C6" box="[159,1388,1134,1156]" colspan="6" colspanRight="5" gridcol="0" gridrow="26" pageId="9" pageNumber="412">Leg II length</th>
</tr>
<tr id="CA8234DC126F001800AAE6F2640EA2EB" box="[159,1388,1171,1193]" gridrow="27" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE6F2600CA2EB" box="[159,366,1171,1193]" gridcol="0" gridrow="27" pageId="9" pageNumber="412">coxa</th>
<td id="89535DA0126F00180192E6F26338A2EB" box="[423,602,1171,1193]" gridcol="1" gridrow="27" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001802A4E6F26227A2EB" box="[657,837,1171,1193]" gridcol="2" gridrow="27" pageId="9" pageNumber="412">0.5</td>
<td id="89535DA0126F001803BBE6F2628DA2EB" box="[910,1007,1171,1193]" gridcol="3" gridrow="27" pageId="9" pageNumber="412">0.42–0.5</td>
<td id="89535DA0126F0018047EE6F265C9A2EB" box="[1099,1195,1171,1193]" gridcol="4" gridrow="27" pageId="9" pageNumber="412">0.37–0.47</td>
<td id="89535DA0126F0018053EE6F2640EA2EB" box="[1291,1388,1171,1193]" gridcol="5" gridrow="27" pageId="9" pageNumber="412">0.37–0.57</td>
</tr>
<tr id="CA8234DC126F001800AAE6D6640EA28F" box="[159,1388,1207,1229]" gridrow="28" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE6D6600CA28F" box="[159,366,1207,1229]" gridcol="0" gridrow="28" pageId="9" pageNumber="412">trochanter</th>
<td id="89535DA0126F00180192E6D66338A28F" box="[423,602,1207,1229]" gridcol="1" gridrow="28" pageId="9" pageNumber="412">0.02</td>
<td id="89535DA0126F001802A4E6D66227A28F" box="[657,837,1207,1229]" gridcol="2" gridrow="28" pageId="9" pageNumber="412">0.25</td>
<td id="89535DA0126F001803BBE6D6628DA28F" box="[910,1007,1207,1229]" gridcol="3" gridrow="28" pageId="9" pageNumber="412">0.02–0.25</td>
<td id="89535DA0126F0018047EE6D665C9A28F" box="[1099,1195,1207,1229]" gridcol="4" gridrow="28" pageId="9" pageNumber="412">0.01–0.25</td>
<td id="89535DA0126F0018053EE6D6640EA28F" box="[1291,1388,1207,1229]" gridcol="5" gridrow="28" pageId="9" pageNumber="412">0.01–0.25</td>
</tr>
<tr id="CA8234DC126F001800AAE6BA640EA2B3" box="[159,1388,1243,1265]" gridrow="29" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE6BA600CA2B3" box="[159,366,1243,1265]" gridcol="0" gridrow="29" pageId="9" pageNumber="412">femur</th>
<td id="89535DA0126F00180192E6BA6338A2B3" box="[423,602,1243,1265]" gridcol="1" gridrow="29" pageId="9" pageNumber="412">0.9</td>
<td id="89535DA0126F001802A4E6BA6227A2B3" box="[657,837,1243,1265]" gridcol="2" gridrow="29" pageId="9" pageNumber="412">0.75</td>
<td id="89535DA0126F001803BBE6BA628DA2B3" box="[910,1007,1243,1265]" gridcol="3" gridrow="29" pageId="9" pageNumber="412">0.7–0.95</td>
<td id="89535DA0126F0018047EE6BA65C9A2B3" box="[1099,1195,1243,1265]" gridcol="4" gridrow="29" pageId="9" pageNumber="412">0.62–0.75</td>
<td id="89535DA0126F0018053EE6BA640EA2B3" box="[1291,1388,1243,1265]" gridcol="5" gridrow="29" pageId="9" pageNumber="412">0.75–0.9</td>
</tr>
<tr id="CA8234DC126F001800AAE761640EA354" box="[159,1388,1280,1302]" gridrow="30" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE761600CA354" box="[159,366,1280,1302]" gridcol="0" gridrow="30" pageId="9" pageNumber="412">patella</th>
<td id="89535DA0126F00180192E7616338A354" box="[423,602,1280,1302]" gridcol="1" gridrow="30" pageId="9" pageNumber="412">0.57</td>
<td id="89535DA0126F001802A4E7616227A354" box="[657,837,1280,1302]" gridcol="2" gridrow="30" pageId="9" pageNumber="412">0.4</td>
<td id="89535DA0126F001803BBE761628DA354" box="[910,1007,1280,1302]" gridcol="3" gridrow="30" pageId="9" pageNumber="412">0.36–0.59</td>
<td id="89535DA0126F0018047EE76165C9A354" box="[1099,1195,1280,1302]" gridcol="4" gridrow="30" pageId="9" pageNumber="412">0.37–0.42</td>
<td id="89535DA0126F0018053EE761640EA354" box="[1291,1388,1280,1302]" gridcol="5" gridrow="30" pageId="9" pageNumber="412">0.35–0.5</td>
</tr>
<tr id="CA8234DC126F001800AAE745640EA378" box="[159,1388,1316,1338]" gridrow="31" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE745600CA378" box="[159,366,1316,1338]" gridcol="0" gridrow="31" pageId="9" pageNumber="412">tibia</th>
<td id="89535DA0126F00180192E7456338A378" box="[423,602,1316,1338]" gridcol="1" gridrow="31" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001802A4E7456227A378" box="[657,837,1316,1338]" gridcol="2" gridrow="31" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001803BBE745628DA378" box="[910,1007,1316,1338]" gridcol="3" gridrow="31" pageId="9" pageNumber="412">0.4–0.56</td>
<td id="89535DA0126F0018047EE74565C9A378" box="[1099,1195,1316,1338]" gridcol="4" gridrow="31" pageId="9" pageNumber="412">0.42–0.5</td>
<td id="89535DA0126F0018053EE745640EA378" box="[1291,1388,1316,1338]" gridcol="5" gridrow="31" pageId="9" pageNumber="412">0.45–0.57</td>
</tr>
<tr id="CA8234DC126F001800AAE729640EA31C" box="[159,1388,1352,1374]" gridrow="32" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE729600CA31C" box="[159,366,1352,1374]" gridcol="0" gridrow="32" pageId="9" pageNumber="412">basitarsus</th>
<td id="89535DA0126F00180192E7296338A31C" box="[423,602,1352,1374]" gridcol="1" gridrow="32" pageId="9" pageNumber="412">0.5</td>
<td id="89535DA0126F001802A4E7296227A31C" box="[657,837,1352,1374]" gridcol="2" gridrow="32" pageId="9" pageNumber="412">0.45</td>
<td id="89535DA0126F001803BBE729628DA31C" box="[910,1007,1352,1374]" gridcol="3" gridrow="32" pageId="9" pageNumber="412">0.36–0.5</td>
<td id="89535DA0126F0018047EE72965C9A31C" box="[1099,1195,1352,1374]" gridcol="4" gridrow="32" pageId="9" pageNumber="412">0.45–0.5</td>
<td id="89535DA0126F0018053EE729640EA31C" box="[1291,1388,1352,1374]" gridcol="5" gridrow="32" pageId="9" pageNumber="412">0.42–0.5</td>
</tr>
<tr id="CA8234DC126F001800AAE70D640EA3C1" box="[159,1388,1388,1411]" gridrow="33" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE70D600CA3C1" box="[159,366,1388,1411]" gridcol="0" gridrow="33" pageId="9" pageNumber="412">telotarsus</th>
<td id="89535DA0126F00180192E70D6338A3C1" box="[423,602,1388,1411]" gridcol="1" gridrow="33" pageId="9" pageNumber="412">0.47</td>
<td id="89535DA0126F001802A4E70D6227A3C1" box="[657,837,1388,1411]" gridcol="2" gridrow="33" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE70D628DA3C1" box="[910,1007,1388,1411]" gridcol="3" gridrow="33" pageId="9" pageNumber="412">0.37–0.47</td>
<td id="89535DA0126F0018047EE70D65C9A3C1" box="[1099,1195,1388,1411]" gridcol="4" gridrow="33" pageId="9" pageNumber="412">0.37–0.52</td>
<td id="89535DA0126F0018053EE70D640EA3C1" box="[1291,1388,1388,1411]" gridcol="5" gridrow="33" pageId="9" pageNumber="412">0.37–0.47</td>
</tr>
<tr id="CA8234DC126F001800AAE7F0640EA3E5" box="[159,1388,1425,1447]" gridrow="34" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE7F0640EA3E5" box="[159,1388,1425,1447]" colspan="6" colspanRight="5" gridcol="0" gridrow="34" pageId="9" pageNumber="412">Leg III length</th>
</tr>
<tr id="CA8234DC126F001800AAE7D4640EA389" box="[159,1388,1461,1483]" gridrow="35" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE7D4600CA389" box="[159,366,1461,1483]" gridcol="0" gridrow="35" pageId="9" pageNumber="412">coxa</th>
<td id="89535DA0126F00180192E7D46338A389" box="[423,602,1461,1483]" gridcol="1" gridrow="35" pageId="9" pageNumber="412">0.45</td>
<td id="89535DA0126F001802A4E7D46227A389" box="[657,837,1461,1483]" gridcol="2" gridrow="35" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE7D4628DA389" box="[910,1007,1461,1483]" gridcol="3" gridrow="35" pageId="9" pageNumber="412">0.3–0.45</td>
<td id="89535DA0126F0018047EE7D465C9A389" box="[1099,1195,1461,1483]" gridcol="4" gridrow="35" pageId="9" pageNumber="412">0.3–0.42</td>
<td id="89535DA0126F0018053EE7D4640EA389" box="[1291,1388,1461,1483]" gridcol="5" gridrow="35" pageId="9" pageNumber="412">0.35–0.57</td>
</tr>
<tr id="CA8234DC126F001800AAE7B8640EA3AD" box="[159,1388,1497,1519]" gridrow="36" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE7B8600CA3AD" box="[159,366,1497,1519]" gridcol="0" gridrow="36" pageId="9" pageNumber="412">trochanter</th>
<td id="89535DA0126F00180192E7B86338A3AD" box="[423,602,1497,1519]" gridcol="1" gridrow="36" pageId="9" pageNumber="412">0.02</td>
<td id="89535DA0126F001802A4E7B86227A3AD" box="[657,837,1497,1519]" gridcol="2" gridrow="36" pageId="9" pageNumber="412">0.02</td>
<td id="89535DA0126F001803BBE7B8628DA3AD" box="[910,1007,1497,1519]" gridcol="3" gridrow="36" pageId="9" pageNumber="412">0.02–0.25</td>
<td id="89535DA0126F0018047EE7B865C9A3AD" box="[1099,1195,1497,1519]" gridcol="4" gridrow="36" pageId="9" pageNumber="412">0.01–0.25</td>
<td id="89535DA0126F0018053EE7B8640EA3AD" box="[1291,1388,1497,1519]" gridcol="5" gridrow="36" pageId="9" pageNumber="412">0.02–0.3</td>
</tr>
<tr id="CA8234DC126F001800AAE79F640EA056" box="[159,1388,1534,1556]" gridrow="37" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE79F600CA056" box="[159,366,1534,1556]" gridcol="0" gridrow="37" pageId="9" pageNumber="412">femur</th>
<td id="89535DA0126F00180192E79F6338A056" box="[423,602,1534,1556]" gridcol="1" gridrow="37" pageId="9" pageNumber="412">0.65</td>
<td id="89535DA0126F001802A4E79F6227A056" box="[657,837,1534,1556]" gridcol="2" gridrow="37" pageId="9" pageNumber="412">0.7</td>
<td id="89535DA0126F001803BBE79F628DA056" box="[910,1007,1534,1556]" gridcol="3" gridrow="37" pageId="9" pageNumber="412">0.6–0.75</td>
<td id="89535DA0126F0018047EE79F65C9A056" box="[1099,1195,1534,1556]" gridcol="4" gridrow="37" pageId="9" pageNumber="412">0.5–0.67</td>
<td id="89535DA0126F0018053EE79F640EA056" box="[1291,1388,1534,1556]" gridcol="5" gridrow="37" pageId="9" pageNumber="412">0.5–0.8</td>
</tr>
<tr id="CA8234DC126F001800AAE443640EA07A" box="[159,1388,1570,1592]" gridrow="38" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE443600CA07A" box="[159,366,1570,1592]" gridcol="0" gridrow="38" pageId="9" pageNumber="412">patella</th>
<td id="89535DA0126F00180192E4436338A07A" box="[423,602,1570,1592]" gridcol="1" gridrow="38" pageId="9" pageNumber="412">0.35</td>
<td id="89535DA0126F001802A4E4436227A07A" box="[657,837,1570,1592]" gridcol="2" gridrow="38" pageId="9" pageNumber="412">0.32</td>
<td id="89535DA0126F001803BBE443628DA07A" box="[910,1007,1570,1592]" gridcol="3" gridrow="38" pageId="9" pageNumber="412">0.26–0.38</td>
<td id="89535DA0126F0018047EE44365C9A07A" box="[1099,1195,1570,1592]" gridcol="4" gridrow="38" pageId="9" pageNumber="412">0.3–0.45</td>
<td id="89535DA0126F0018053EE443640EA07A" box="[1291,1388,1570,1592]" gridcol="5" gridrow="38" pageId="9" pageNumber="412">0.02–0.37</td>
</tr>
<tr id="CA8234DC126F001800AAE427640EA01E" box="[159,1388,1606,1628]" gridrow="39" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE427600CA01E" box="[159,366,1606,1628]" gridcol="0" gridrow="39" pageId="9" pageNumber="412">tibia</th>
<td id="89535DA0126F00180192E4276338A01E" box="[423,602,1606,1628]" gridcol="1" gridrow="39" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001802A4E4276227A01E" box="[657,837,1606,1628]" gridcol="2" gridrow="39" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE427628DA01E" box="[910,1007,1606,1628]" gridcol="3" gridrow="39" pageId="9" pageNumber="412">0.3–0.42</td>
<td id="89535DA0126F0018047EE42765C9A01E" box="[1099,1195,1606,1628]" gridcol="4" gridrow="39" pageId="9" pageNumber="412">0.3–0.35</td>
<td id="89535DA0126F0018053EE427640EA01E" box="[1291,1388,1606,1628]" gridcol="5" gridrow="39" pageId="9" pageNumber="412">0.35–0.4</td>
</tr>
<tr id="CA8234DC126F001800AAE40B640EA0C3" box="[159,1388,1642,1665]" gridrow="40" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE40B600CA0C3" box="[159,366,1642,1665]" gridcol="0" gridrow="40" pageId="9" pageNumber="412">basitarsus</th>
<td id="89535DA0126F00180192E40B6338A0C3" box="[423,602,1642,1665]" gridcol="1" gridrow="40" pageId="9" pageNumber="412">0.62</td>
<td id="89535DA0126F001802A4E40B6227A0C3" box="[657,837,1642,1665]" gridcol="2" gridrow="40" pageId="9" pageNumber="412">0.42</td>
<td id="89535DA0126F001803BBE40B628DA0C3" box="[910,1007,1642,1665]" gridcol="3" gridrow="40" pageId="9" pageNumber="412">0.37–0.63</td>
<td id="89535DA0126F0018047EE40B65C9A0C3" box="[1099,1195,1642,1665]" gridcol="4" gridrow="40" pageId="9" pageNumber="412">0.37–0.5</td>
<td id="89535DA0126F0018053EE40B640EA0C3" box="[1291,1388,1642,1665]" gridcol="5" gridrow="40" pageId="9" pageNumber="412">0.27–0.5</td>
</tr>
<tr id="CA8234DC126F001800AAE4EE640EA0E7" box="[159,1388,1679,1701]" gridrow="41" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE4EE600CA0E7" box="[159,366,1679,1701]" gridcol="0" gridrow="41" pageId="9" pageNumber="412">telotarsus</th>
<td id="89535DA0126F00180192E4EE6338A0E7" box="[423,602,1679,1701]" gridcol="1" gridrow="41" pageId="9" pageNumber="412">0.45</td>
<td id="89535DA0126F001802A4E4EE6227A0E7" box="[657,837,1679,1701]" gridcol="2" gridrow="41" pageId="9" pageNumber="412">0.25</td>
<td id="89535DA0126F001803BBE4EE628DA0E7" box="[910,1007,1679,1701]" gridcol="3" gridrow="41" pageId="9" pageNumber="412">0.25–0.46</td>
<td id="89535DA0126F0018047EE4EE65C9A0E7" box="[1099,1195,1679,1701]" gridcol="4" gridrow="41" pageId="9" pageNumber="412">0.27–0.37</td>
<td id="89535DA0126F0018053EE4EE640EA0E7" box="[1291,1388,1679,1701]" gridcol="5" gridrow="41" pageId="9" pageNumber="412">0.25–0.47</td>
</tr>
<tr id="CA8234DC126F001800AAE4D2640EA08B" box="[159,1388,1715,1737]" gridrow="42" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE4D2640EA08B" box="[159,1388,1715,1737]" colspan="6" colspanRight="5" gridcol="0" gridrow="42" pageId="9" pageNumber="412">Leg IV length</th>
</tr>
<tr id="CA8234DC126F001800AAE4B6640EA0AF" box="[159,1388,1751,1773]" gridrow="43" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE4B6600CA0AF" box="[159,366,1751,1773]" gridcol="0" gridrow="43" pageId="9" pageNumber="412">coxa</th>
<td id="89535DA0126F00180192E4B66338A0AF" box="[423,602,1751,1773]" gridcol="1" gridrow="43" pageId="9" pageNumber="412">0.3</td>
<td id="89535DA0126F001802A4E4B66227A0AF" box="[657,837,1751,1773]" gridcol="2" gridrow="43" pageId="9" pageNumber="412">0.35</td>
<td id="89535DA0126F001803BBE4B6628DA0AF" box="[910,1007,1751,1773]" gridcol="3" gridrow="43" pageId="9" pageNumber="412">0.3–0.35</td>
<td id="89535DA0126F0018047EE4B665C9A0AF" box="[1099,1195,1751,1773]" gridcol="4" gridrow="43" pageId="9" pageNumber="412">0.3–0.37</td>
<td id="89535DA0126F0018053EE4B6640EA0AF" box="[1291,1388,1751,1773]" gridcol="5" gridrow="43" pageId="9" pageNumber="412">0.32–0.4</td>
</tr>
<tr id="CA8234DC126F001800AAE49D640EA150" box="[159,1388,1788,1810]" gridrow="44" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE49D600CA150" box="[159,366,1788,1810]" gridcol="0" gridrow="44" pageId="9" pageNumber="412">trochanter</th>
<td id="89535DA0126F00180192E49D6338A150" box="[423,602,1788,1810]" gridcol="1" gridrow="44" pageId="9" pageNumber="412">0.27</td>
<td id="89535DA0126F001802A4E49D6227A150" box="[657,837,1788,1810]" gridcol="2" gridrow="44" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE49D628DA150" box="[910,1007,1788,1810]" gridcol="3" gridrow="44" pageId="9" pageNumber="412">0.25–0.42</td>
<td id="89535DA0126F0018047EE49D65C9A150" box="[1099,1195,1788,1810]" gridcol="4" gridrow="44" pageId="9" pageNumber="412">0.27–0.37</td>
<td id="89535DA0126F0018053EE49D640EA150" box="[1291,1388,1788,1810]" gridcol="5" gridrow="44" pageId="9" pageNumber="412">0.25–0.37</td>
</tr>
<tr id="CA8234DC126F001800AAE541640EA174" box="[159,1388,1824,1846]" gridrow="45" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE541600CA174" box="[159,366,1824,1846]" gridcol="0" gridrow="45" pageId="9" pageNumber="412">femur</th>
<td id="89535DA0126F00180192E5416338A174" box="[423,602,1824,1846]" gridcol="1" gridrow="45" pageId="9" pageNumber="412">1.2</td>
<td id="89535DA0126F001802A4E5416227A174" box="[657,837,1824,1846]" gridcol="2" gridrow="45" pageId="9" pageNumber="412">1.25</td>
<td id="89535DA0126F001803BBE541628DA174" box="[910,1007,1824,1846]" gridcol="3" gridrow="45" pageId="9" pageNumber="412">0.86–1.3</td>
<td id="89535DA0126F0018047EE54165C9A174" box="[1099,1195,1824,1846]" gridcol="4" gridrow="45" pageId="9" pageNumber="412">1–1.17</td>
<td id="89535DA0126F0018053EE541640EA174" box="[1291,1388,1824,1846]" gridcol="5" gridrow="45" pageId="9" pageNumber="412">1.15–1.3</td>
</tr>
<tr id="CA8234DC126F001800AAE525640EA118" box="[159,1388,1860,1882]" gridrow="46" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE525600CA118" box="[159,366,1860,1882]" gridcol="0" gridrow="46" pageId="9" pageNumber="412">patella</th>
<td id="89535DA0126F00180192E5256338A118" box="[423,602,1860,1882]" gridcol="1" gridrow="46" pageId="9" pageNumber="412">0.5</td>
<td id="89535DA0126F001802A4E5256227A118" box="[657,837,1860,1882]" gridcol="2" gridrow="46" pageId="9" pageNumber="412">0.52</td>
<td id="89535DA0126F001803BBE525628DA118" box="[910,1007,1860,1882]" gridcol="3" gridrow="46" pageId="9" pageNumber="412">0.4–0.6</td>
<td id="89535DA0126F0018047EE52565C9A118" box="[1099,1195,1860,1882]" gridcol="4" gridrow="46" pageId="9" pageNumber="412">0.45–0.55</td>
<td id="89535DA0126F0018053EE525640EA118" box="[1291,1388,1860,1882]" gridcol="5" gridrow="46" pageId="9" pageNumber="412">0.45–0.57</td>
</tr>
<tr id="CA8234DC126F001800AAE508640EA13D" box="[159,1388,1897,1919]" gridrow="47" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE508600CA13D" box="[159,366,1897,1919]" gridcol="0" gridrow="47" pageId="9" pageNumber="412">tibia</th>
<td id="89535DA0126F00180192E5086338A13D" box="[423,602,1897,1919]" gridcol="1" gridrow="47" pageId="9" pageNumber="412">0.8</td>
<td id="89535DA0126F001802A4E5086227A13D" box="[657,837,1897,1919]" gridcol="2" gridrow="47" pageId="9" pageNumber="412">0.72</td>
<td id="89535DA0126F001803BBE508628DA13D" box="[910,1007,1897,1919]" gridcol="3" gridrow="47" pageId="9" pageNumber="412">0.6–0.9</td>
<td id="89535DA0126F0018047EE50865C9A13D" box="[1099,1195,1897,1919]" gridcol="4" gridrow="47" pageId="9" pageNumber="412">0.67–0.85</td>
<td id="89535DA0126F0018053EE508640EA13D" box="[1291,1388,1897,1919]" gridcol="5" gridrow="47" pageId="9" pageNumber="412">0.8–0.87</td>
</tr>
<tr id="CA8234DC126F001800AAE5EC640EA1E1" box="[159,1388,1933,1955]" gridrow="48" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE5EC600CA1E1" box="[159,366,1933,1955]" gridcol="0" gridrow="48" pageId="9" pageNumber="412">basitarsus</th>
<td id="89535DA0126F00180192E5EC6338A1E1" box="[423,602,1933,1955]" gridcol="1" gridrow="48" pageId="9" pageNumber="412">0.65</td>
<td id="89535DA0126F001802A4E5EC6227A1E1" box="[657,837,1933,1955]" gridcol="2" gridrow="48" pageId="9" pageNumber="412">0.62</td>
<td id="89535DA0126F001803BBE5EC628DA1E1" box="[910,1007,1933,1955]" gridcol="3" gridrow="48" pageId="9" pageNumber="412">0.62–0.75</td>
<td id="89535DA0126F0018047EE5EC65C9A1E1" box="[1099,1195,1933,1955]" gridcol="4" gridrow="48" pageId="9" pageNumber="412">0.6–0.62</td>
<td id="89535DA0126F0018053EE5EC640EA1E1" box="[1291,1388,1933,1955]" gridcol="5" gridrow="48" pageId="9" pageNumber="412">0.6–0.72</td>
</tr>
<tr id="CA8234DC126F001800AAE5D0640EA185" box="[159,1388,1969,1991]" gridrow="49" pageId="9" pageNumber="412">
<th id="89535DA0126F001800AAE5D0600CA185" box="[159,366,1969,1991]" gridcol="0" gridrow="49" pageId="9" pageNumber="412">telotarsus</th>
<td id="89535DA0126F00180192E5D06338A185" box="[423,602,1969,1991]" gridcol="1" gridrow="49" pageId="9" pageNumber="412">0.4</td>
<td id="89535DA0126F001802A4E5D06227A185" box="[657,837,1969,1991]" gridcol="2" gridrow="49" pageId="9" pageNumber="412">0.37</td>
<td id="89535DA0126F001803BBE5D0628DA185" box="[910,1007,1969,1991]" gridcol="3" gridrow="49" pageId="9" pageNumber="412">0.37–0.45</td>
<td id="89535DA0126F0018047EE5D065C9A185" box="[1099,1195,1969,1991]" gridcol="4" gridrow="49" pageId="9" pageNumber="412">0.37–0.52</td>
<td id="89535DA0126F0018053EE5D0640EA185" box="[1291,1388,1969,1991]" gridcol="5" gridrow="49" pageId="9" pageNumber="412">0.42–0.52</td>
</tr>
</table>
</paragraph>
<paragraph id="740D369E126CFFED00F2E2F661B5A75F" blockId="10.[151,1437,150,285]" pageId="10" pageNumber="413">
<emphasis id="46C6EA8C126CFFED00F2E2F66058A6F3" box="[199,314,151,177]" italics="true" pageId="10" pageNumber="413">Flagellum</emphasis>
<emphasis id="46C6EA8C126CFFED0176E2F7602EA6F2" bold="true" box="[323,332,150,176]" pageId="10" pageNumber="413">(</emphasis>
<figureCitation id="EC892A1B126CFFED0179E2F660AFA6F3" box="[332,461,151,177]" captionStart="FIGURES 36–42" captionStartId="8.[151,264,1911,1935]" captionTargetBox="[151,1436,630,1885]" captionTargetId="figure-246@8.[151,1436,630,1885]" captionTargetPageId="8" captionText="FIGURES 36–42. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4064), habitus (36–39) and flagellum (40–42), in dorsal (36–37, 40) and ventral (38–39, 41) and retrolateral (42) views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11547618" httpUri="https://zenodo.org/record/11547618/files/figure.png" pageId="10" pageNumber="413">Figs 40–42</figureCitation>
,
<figureCitation id="EC892A1B126CFFED01EFE2F76342A6F3" box="[474,544,150,177]" captionStart="FIGURES 43–46" captionStartId="10.[151,264,1884,1908]" captionTargetBox="[169,1420,313,1854]" captionTargetId="figure-85@10.[151,1436,306,1858]" captionTargetPageId="10" captionText="FIGURES 43–46. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4057, flagellum (43–45): dorsal (43), ventral (44) and lateral (45) views; and internal genitalia (46). Scale bars: 0.5 mm (43–45), 0.1 mm (46). Abbreviations: APA, anterior portion of the chitinized arch; B, bulb; G, gonopod; LL, lateral lobes; ML, median lobes; PPA, posterior portion of the chitinized arch; S, stalk." figureDoi="http://doi.org/10.5281/zenodo.11547620" httpUri="https://zenodo.org/record/11547620/files/figure.png" pageId="10" pageNumber="413">43–45</figureCitation>
<emphasis id="46C6EA8C126CFFED0215E2F7634BA6F2" bold="true" box="[544,553,150,176]" pageId="10" pageNumber="413">)</emphasis>
. With 4 flagellomeres and 3 annuli, 4.6 times longer than wide. Flagellomere II with Dm1, Vm1 and Vm2 present. Flagellomere III with Vm4. Flagellomere IV with Vl1 at the same level Vm5 and Dl1; Dl1 between level of Vl1 and Vm5; Dm4 between Dl1 and Dl3; Vl2 slightly proximal to Dl3 and distal to Dm4.
</paragraph>
<caption id="20CD6616126CFFED00A2E53D618BA1A2" ID-DOI="http://doi.org/10.5281/zenodo.11547620" ID-Zenodo-Dep="11547620" httpUri="https://zenodo.org/record/11547620/files/figure.png" pageId="10" pageNumber="413" startId="10.[151,264,1884,1908]" targetBox="[169,1420,313,1854]" targetPageId="10" targetType="figure">
<paragraph id="740D369E126CFFED00A2E53D618BA1A2" blockId="10.[151,1437,1884,2016]" pageId="10" pageNumber="413">
<emphasis id="46C6EA8C126CFFED00A2E53D602EA136" bold="true" box="[151,332,1884,1908]" pageId="10" pageNumber="413">FIGURES 43–46</emphasis>
.
<taxonomicName id="B3B24D1D126CFFED016CE53D609FA136" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[345,509,1884,1908]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="10" pageNumber="413" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126CFFED016CE53D609FA136" box="[345,509,1884,1908]" italics="true" pageId="10" pageNumber="413">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126CFFED0236E53D6330A136" bold="true" box="[515,594,1884,1908]" pageId="10" pageNumber="413">
<taxonomicNameLabel id="5DF557F7126CFFED0236E53D6330A136" box="[515,594,1884,1908]" pageId="10" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, paratype (CHNUFPI 4057, flagellum (43–45): dorsal (43), ventral (44) and lateral (45) views; and internal genitalia (46). Scale bars: 0.5 mm (43–45), 0.1 mm (46). Abbreviations: APA, anterior portion of the chitinized arch; B, bulb; G, gonopod; LL, lateral lobes; ML, median lobes; PPA, posterior portion of the chitinized arch; S, stalk.
</paragraph>
</caption>
<paragraph id="740D369E126DFFEC00F2E2F66355A703" blockId="11.[151,1437,150,321]" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00F2E2F66004A6F3" box="[199,358,151,177]" italics="true" pageId="11" pageNumber="414">Spermathecae</emphasis>
(
<figureCitation id="EC892A1B126DFFEC014DE2F6609CA6F3" box="[376,510,150,177]" captionStart-0="FIGURES 43–46" captionStart-1="FIGURES 47–50" captionStartId-0="10.[151,264,1884,1908]" captionStartId-1="11.[151,264,1226,1250]" captionTargetBox-0="[169,1420,313,1854]" captionTargetBox-1="[151,1436,342,1200]" captionTargetId-0="figure-85@10.[151,1436,306,1858]" captionTargetId-1="figure-93@11.[151,1436,342,1200]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 43–46. Rowlandius ufpi sp. nov. female, paratype (CHNUFPI 4057, flagellum (43–45): dorsal (43), ventral (44) and lateral (45) views; and internal genitalia (46). Scale bars: 0.5 mm (43–45), 0.1 mm (46). Abbreviations: APA, anterior portion of the chitinized arch; B, bulb; G, gonopod; LL, lateral lobes; ML, median lobes; PPA, posterior portion of the chitinized arch; S, stalk." captionText-1="FIGURES 47–50. Rowlandius ufpi sp. nov. female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11547620" figureDoi-1="http://doi.org/10.5281/zenodo.11547622" httpUri-0="https://zenodo.org/record/11547620/files/figure.png" httpUri-1="https://zenodo.org/record/11547622/files/figure.png" pageId="11" pageNumber="414">Figs 46–50</figureCitation>
). Chitinized arch evident; closed and formed by anterior and posterior portions; anterior portion with an obtuse (&gt;150°) v-shaped angle; posterior portion with a gonopod medially; pointed laterally. One pair of lateral lobes (LL) with long, thin stalks whose bases are separated by a distance greater than their length; with the bulb globose and oriented towards the middle; the median lobes (ML) short, with a thin base. Gonopod present, hyaline, directed posteriorly.
</paragraph>
</subSubSection>
<caption id="20CD6616126DFFEC00A2E6AB62A7A344" ID-DOI="http://doi.org/10.5281/zenodo.11547622" ID-Zenodo-Dep="11547622" httpUri="https://zenodo.org/record/11547622/files/figure.png" pageId="11" pageNumber="414" startId="11.[151,264,1226,1250]" targetBox="[151,1436,342,1200]" targetPageId="11" targetType="figure">
<paragraph id="740D369E126DFFEC00A2E6AB62A7A344" blockId="11.[151,1437,1226,1287]" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00A2E6AB6032A2A0" bold="true" box="[151,336,1226,1250]" pageId="11" pageNumber="414">FIGURES 47–50</emphasis>
.
<taxonomicName id="B3B24D1D126DFFEC0155E6AB636AA2A1" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[352,520,1226,1251]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="11" pageNumber="414" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126DFFEC0155E6AB636AA2A1" box="[352,520,1226,1251]" italics="true" pageId="11" pageNumber="414">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126DFFEC0227E6AA6307A2A1" bold="true" box="[530,613,1227,1251]" pageId="11" pageNumber="414">
<taxonomicNameLabel id="5DF557F7126DFFEC0227E6AA6307A2A1" box="[530,613,1227,1251]" pageId="11" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="3CA86515126DFFEC00F2E73962C9A3D5" pageId="11" pageNumber="414" type="distribution">
<paragraph id="740D369E126DFFEC00F2E73962C9A3D5" blockId="11.[151,1437,1368,2043]" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00F2E7396003A330" bold="true" box="[199,353,1368,1394]" pageId="11" pageNumber="414">Distribution.</emphasis>
This species is known only from the
<typeStatus id="AB09883C126DFFEC032DE738622AA331" box="[792,840,1369,1395]" pageId="11" pageNumber="414">type</typeStatus>
locality (
<figureCitation id="EC892A1B126DFFEC0382E738655FA331" box="[951,1085,1368,1395]" captionStart-0="FIGURES 51–54" captionStart-1="FIGURES 55–57" captionStart-2="FIGURE 58" captionStartId-0="12.[151,264,1112,1136]" captionStartId-1="13.[151,264,1467,1491]" captionStartId-2="14.[151,250,1079,1103]" captionTargetBox-0="[152,1436,234,1086]" captionTargetBox-1="[151,1436,187,1443]" captionTargetBox-2="[176,1413,187,1050]" captionTargetId-0="figure-52@12.[152,1436,234,1086]" captionTargetId-1="figure-19@13.[151,1436,181,1443]" captionTargetId-2="figure-23@14.[171,1417,181,1055]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURES 51–54. Environments at the type-locality of Rowlandius ufpi sp. nov.. 51–52: Riparian zone, where the examined specimens were found. 53–54: Phytophysiognomies where no schizomid was observed, savanna (Cerrado stricto sensu; 53) and seasonally dry tropical forest (54)." captionText-1="FIGURES 55–57. Sites sampled in the search for Rowlandius ufpi sp. nov. specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where R. ufpi sp. nov. specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where R. ufpi sp. nov., no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth." captionText-2="FIGURE 58. Known geographic distribution of described schizomid species from continental South America, including the Rowlandius ufpi sp. nov. (red circle), the remaining Rowlandius taxa (yellow squares) and other taxa (blue triangles)." figureDoi-0="http://doi.org/10.5281/zenodo.11547626" figureDoi-1="http://doi.org/10.5281/zenodo.11547628" figureDoi-2="http://doi.org/10.5281/zenodo.11547630" httpUri-0="https://zenodo.org/record/11547626/files/figure.png" httpUri-1="https://zenodo.org/record/11547628/files/figure.png" httpUri-2="https://zenodo.org/record/11547630/files/figure.png" pageId="11" pageNumber="414">Figs 51–58</figureCitation>
). This record fills a gap in the geographic distribution of
<taxonomicName id="B3B24D1D126DFFEC01F4E71C6327A3D5" authorityName="Reddell &amp; Cokendolpher" authorityYear="1995" box="[449,581,1405,1431]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="11" pageNumber="414" phylum="Arthropoda" rank="genus">
<emphasis id="46C6EA8C126DFFEC01F4E71C6327A3D5" box="[449,581,1405,1431]" italics="true" pageId="11" pageNumber="414">Rowlandius</emphasis>
</taxonomicName>
in South America (see
<figureCitation id="EC892A1B126DFFEC037EE71C62FDA3D5" box="[843,927,1404,1431]" captionStart="FIGURE 58" captionStartId="14.[151,250,1079,1103]" captionTargetBox="[176,1413,187,1050]" captionTargetId="figure-23@14.[171,1417,181,1055]" captionTargetPageId="14" captionText="FIGURE 58. Known geographic distribution of described schizomid species from continental South America, including the Rowlandius ufpi sp. nov. (red circle), the remaining Rowlandius taxa (yellow squares) and other taxa (blue triangles)." figureDoi="http://doi.org/10.5281/zenodo.11547630" httpUri="https://zenodo.org/record/11547630/files/figure.png" pageId="11" pageNumber="414">Fig. 58</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3CA86515126DFFE800F2E7C162ABA75F" lastPageId="15" lastPageNumber="418" pageId="11" pageNumber="414" type="description">
<paragraph id="740D369E126DFFEC00F2E7C16579A099" blockId="11.[151,1437,1368,2043]" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00F2E7C1605DA3F8" bold="true" box="[199,319,1440,1466]" pageId="11" pageNumber="414">Variation.</emphasis>
Number of individuals measured (
<tableCitation id="39300325126DFFEC02F6E7C0627AA3F9" box="[707,792,1441,1467]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Measurements (in mm) of Rowlandius ufpi sp. nov." httpUri="http://table.plazi.org/id/20CD6616126FFFEE00A2E2F6625AA6F2" pageId="11" pageNumber="414" tableUuid="20CD6616126FFFEE00A2E2F6625AA6F2">Table 1</tableCitation>
): heteromorphic male (N=5), homeomorphic male (N=5) and female (N=5). Total length: heteromorphic male
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, homeomorphic male
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, female
<quantity id="B34A9B7B126DFFEC0568E7A461BEA041" metricMagnitude="-3" metricUnit="m" metricValue="4.22" metricValueMax="5.0" metricValueMin="3.44" pageId="11" pageNumber="414" unit="mm" value="4.22" valueMax="5.0" valueMin="3.44">3.44– 5 mm</quantity>
. Length of pedipalpal femur: heteromorphic male
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, homeomorphic male
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, female
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. Length of pedipalpal patella: heteromorphic male
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, homeomorphic male
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, female
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(
<tableCitation id="39300325126DFFEC01AEE450608DA009" box="[411,495,1585,1611]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Measurements (in mm) of Rowlandius ufpi sp. nov." httpUri="http://table.plazi.org/id/20CD6616126FFFEE00A2E2F6625AA6F2" pageId="11" pageNumber="414" tableUuid="20CD6616126FFFEE00A2E2F6625AA6F2">Table 1</tableCitation>
). No relevant variation was observed in the pedipalp and flagellum setation of male specimens. The female spermathecae revealed a slight variation in the orientation of the median lobes, and in the position and shape of the lateral lobe bulbs (
<figureCitation id="EC892A1B126DFFEC02BDE418626EA0D1" box="[648,780,1656,1683]" captionStart="FIGURES 47–50" captionStartId="11.[151,264,1226,1250]" captionTargetBox="[151,1436,342,1200]" captionTargetId="figure-93@11.[151,1436,342,1200]" captionTargetPageId="11" captionText="FIGURES 47–50. Rowlandius ufpi sp. nov. female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11547622" httpUri="https://zenodo.org/record/11547622/files/figure.png" pageId="11" pageNumber="414">Figs 47–50</figureCitation>
). The orientation of the bulbs can be different due to piece preparation, owing to the very thin stalk to which the bulb is attached. The gonopod might be challenging to observe owing to its weak sclerotization and destructive effects of specimen preparation.
</paragraph>
<paragraph id="740D369E126DFFEC00F2E4846244A129" blockId="11.[151,1437,1368,2043]" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00F2E48460EAA0BC" bold="true" box="[199,392,1764,1791]" pageId="11" pageNumber="414">Natural history.</emphasis>
In
<date id="000C105E126DFFEC0186E484633FA0BD" box="[435,605,1765,1791]" pageId="11" pageNumber="414" value="2020-02">February 2020</date>
, a female carrying her offspring on her opisthosoma was collected. The hatchling cluster presented 17 postembryos attached to the ventral side of the female opisthosoma (
<figureCitation id="EC892A1B126DFFEC04DCE5686432A161" box="[1257,1360,1801,1827]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="11" pageNumber="414">Figs 4–6</figureCitation>
). After collection, the clutch detached from the opisthosoma, but the female did not grab it back. All the embryos were parallel to each other and in the same direction (
<figureCitation id="EC892A1B126DFFEC0287E530627BA129" box="[690,793,1873,1899]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1496,1520]" captionTargetBox="[151,1436,179,1470]" captionTargetId="figure-19@3.[151,1436,179,1470]" captionTargetPageId="3" captionText="FIGURES 1–6. Rowlandius ufpi sp. nov., habitus of heteromorphic (1) and homeomorphic male (2), female (3) and embryo clutch (4–6). Photos: L.S. Carvalho (1, 4–6) and P.H. Martins (2–3)." figureDoi="http://doi.org/10.5281/zenodo.11547605" httpUri="https://zenodo.org/record/11547605/files/figure.png" pageId="11" pageNumber="414">Figs 4–6</figureCitation>
).
</paragraph>
<paragraph id="740D369E126DFFEB00F2E515640DA697" blockId="11.[151,1437,1368,2043]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="415" pageId="11" pageNumber="414">
<emphasis id="46C6EA8C126DFFEC00F2E515609AA1CD" bold="true" box="[199,504,1908,1935]" pageId="11" pageNumber="414">Conservation assessment.</emphasis>
This species has been under observation for ten years in its type-locality, in gallery and riverine forests within the Cerrado (savanna) domain (or Cerrado province sensu
<bibRefCitation id="10234B6F126DFFEC046CE5F8659CA1F1" author="Morrone, J. J." box="[1113,1278,1945,1971]" pageId="11" pageNumber="414" refId="ref8082" refString="Morrone, J. J. (2017) Neotropical Biogeography: Regionalization and Evolution. CRC Press, Boca Raton, Florida, 282 pp. https: // doi. org / 10.1201 / b 21824" type="book" year="2017">Morrone 2017</bibRefCitation>
), northeastern
<collectingCountry id="0CA5760E126DFFEC00A2E5DC61BDA195" box="[151,223,1981,2007]" name="Brazil" pageId="11" pageNumber="414">Brazil</collectingCountry>
. The type-locality comprises a mosaic of approx. 200 hectares of pastures, corn, and banana crops, goat and cattle livestock, and secondary and primary patches of several natural phytophysiognomies. The vegetation patches are all typical of the Cerrado domain, including savannas (‘Cerrado
<emphasis id="46C6EA8C126AFFEB03DCE2F66517A6F2" box="[1001,1141,151,176]" italics="true" pageId="12" pageNumber="415">strictu sensu</emphasis>
’), evergreen broad-leaved woodland (‘Cerradão’), seasonally dry tropical forests, riverine and gallery forests following the watercourses.
</paragraph>
<caption id="20CD6616126AFFEB00A2E6396096A2FA" ID-DOI="http://doi.org/10.5281/zenodo.11547626" ID-Zenodo-Dep="11547626" httpUri="https://zenodo.org/record/11547626/files/figure.png" pageId="12" pageNumber="415" startId="12.[151,264,1112,1136]" targetBox="[152,1436,234,1086]" targetPageId="12" targetType="figure">
<paragraph id="740D369E126AFFEB00A2E6396096A2FA" blockId="12.[151,1437,1112,1208]" pageId="12" pageNumber="415">
<emphasis id="46C6EA8C126AFFEB00A2E6396037A232" bold="true" box="[151,341,1112,1136]" pageId="12" pageNumber="415">FIGURES 51–54.</emphasis>
Environments at the type-locality of
<taxonomicName id="B3B24D1D126AFFEB02E3E639621EA232" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[726,892,1112,1136]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="12" pageNumber="415" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126AFFEB02E3E639621EA232" box="[726,892,1112,1136]" italics="true" pageId="12" pageNumber="415">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126AFFEB03B1E63962B4A232" bold="true" box="[900,982,1112,1136]" pageId="12" pageNumber="415">
<taxonomicNameLabel id="5DF557F7126AFFEB03B1E63962B4A232" box="[900,982,1112,1136]" pageId="12" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. 51–52: Riparian zone, where the examined specimens were found. 53–54: Phytophysiognomies where no schizomid was observed, savanna (Cerrado
<emphasis id="46C6EA8C126AFFEB04F0E61C6421A2D6" box="[1221,1347,1149,1172]" italics="true" pageId="12" pageNumber="415">stricto sensu</emphasis>
; 53) and seasonally dry tropical forest (54).
</paragraph>
</caption>
<paragraph id="740D369E126AFFEB00F2E76B65E1A062" blockId="12.[151,1437,1290,1821]" pageId="12" pageNumber="415">
The individuals collected in 2012 and 2013 were found in very dense leaf litter of a gallery forest, situated very close (&lt;
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) to a small temporary stream. However, sampling events conducted by the same collectors, during the dry season in 2012, 2013 and 2014 failed to find any individuals in the same site. We hypothesize that the individuals were buried in the ground at that time, as a strategy to protect themselves from desiccation. This hypothesis contrasts with a study carried out with
<taxonomicName id="B3B24D1D126AFFEB01D8E7FB63B8A3F6" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[493,730,1434,1460]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="12" pageNumber="415" phylum="Arthropoda" rank="species" species="potiguar">
<emphasis id="46C6EA8C126AFFEB01D8E7FB63B8A3F6" box="[493,730,1434,1460]" italics="true" pageId="12" pageNumber="415">Rowlandius potiguar</emphasis>
</taxonomicName>
in captivity (
<bibRefCitation id="10234B6F126AFFEB0343E7FB65F8A3F7" author="Oliveira, M. P. A. &amp; Ferreira, R. L." box="[886,1178,1434,1461]" pageId="12" pageNumber="415" pagination="91913" refId="ref8120" refString="Oliveira, M. P. A. &amp; Ferreira, R. L. (2014) Aspects of the behavior and activity rhythms of Rowlandius potiguar (Schizomida: Hubbardiidae). PLoS ONE, 9 (3), e 91913. https: // doi. org / 10.1371 / journal. pone. 0091913" type="journal article" year="2014">Oliveira &amp; Ferreira 2014</bibRefCitation>
). The authors reported that individuals have the ability to dig their own nests, a behavior suggested to be rare in natural conditions due to the high availability of shelters (
<bibRefCitation id="10234B6F126AFFEB0231E7836246A3BF" author="Oliveira, M. P. A. &amp; Ferreira, R. L." box="[516,804,1506,1533]" pageId="12" pageNumber="415" pagination="91913" refId="ref8120" refString="Oliveira, M. P. A. &amp; Ferreira, R. L. (2014) Aspects of the behavior and activity rhythms of Rowlandius potiguar (Schizomida: Hubbardiidae). PLoS ONE, 9 (3), e 91913. https: // doi. org / 10.1371 / journal. pone. 0091913" type="journal article" year="2014">Oliveira &amp; Ferreira 2014</bibRefCitation>
). Future behavioral studies should address the effects of shelter availability, humidity, and substrate granulometry as triggers for digging behavior.
</paragraph>
<paragraph id="740D369E126AFFEB00F2E44B634AA15F" blockId="12.[151,1437,1290,1821]" pageId="12" pageNumber="415">
Around
<date id="000C105E126AFFEB0111E44B60BFA007" box="[292,477,1578,1605]" pageId="12" pageNumber="415" value="2017-11">November 2017</date>
, the sampling site was severely damaged due to slash-and-burn agricultural practices (compare
<figureCitation id="EC892A1B126AFFEB013EE42F60DCA02B" box="[267,446,1614,1641]" captionStart="FIGURES 55–57" captionStartId="13.[151,264,1467,1491]" captionTargetBox="[151,1436,187,1443]" captionTargetId="figure-19@13.[151,1436,181,1443]" captionTargetPageId="13" captionText="FIGURES 55–57. Sites sampled in the search for Rowlandius ufpi sp. nov. specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where R. ufpi sp. nov. specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where R. ufpi sp. nov., no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth." figureDoi="http://doi.org/10.5281/zenodo.11547628" httpUri="https://zenodo.org/record/11547628/files/figure.png" pageId="12" pageNumber="415">Figs 56 and 57</figureCitation>
). The secondary gallery forest was subsequently replaced by a banana crop upstream in the gallery course, and the exact same site where the schizomids were collected was highly disturbed. Despite additional collecting attempts in
<date id="000C105E126AFFEB0238E4F763EBA0F3" box="[525,649,1686,1713]" pageId="12" pageNumber="415" value="2018-04">April 2018</date>
and on several occasions in 2019, both at the original site and other locations nearby (mostly in gallery forests;
<figureCitation id="EC892A1B126AFFEB0242E4DB63AAA097" box="[631,712,1722,1749]" captionStart="FIGURES 55–57" captionStartId="13.[151,264,1467,1491]" captionTargetBox="[151,1436,187,1443]" captionTargetId="figure-19@13.[151,1436,181,1443]" captionTargetPageId="13" captionText="FIGURES 55–57. Sites sampled in the search for Rowlandius ufpi sp. nov. specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where R. ufpi sp. nov. specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where R. ufpi sp. nov., no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth." figureDoi="http://doi.org/10.5281/zenodo.11547628" httpUri="https://zenodo.org/record/11547628/files/figure.png" pageId="12" pageNumber="415">Fig. 55</figureCitation>
), no additional individuals were found. In 2020, schizomids were found again, in a very limited area of about 100 x
<quantity id="B34A9B7B126AFFEB02F5E4BE6398A0BB" box="[704,762,1759,1785]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="415" unit="m" value="30.0">30 m</quantity>
, within the riverine forest of the Parnaíba River, approx.
<quantity id="B34A9B7B126AFFEB0547E4BF61CFA15F" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="415" unit="m" value="250.0">250 m</quantity>
from the initial site (
<figureCitation id="EC892A1B126AFFEB01ACE563637EA15F" box="[409,540,1794,1821]" captionStart="FIGURES 55–57" captionStartId="13.[151,264,1467,1491]" captionTargetBox="[151,1436,187,1443]" captionTargetId="figure-19@13.[151,1436,181,1443]" captionTargetPageId="13" captionText="FIGURES 55–57. Sites sampled in the search for Rowlandius ufpi sp. nov. specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where R. ufpi sp. nov. specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where R. ufpi sp. nov., no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth." figureDoi="http://doi.org/10.5281/zenodo.11547628" httpUri="https://zenodo.org/record/11547628/files/figure.png" pageId="12" pageNumber="415">Figs 55–57</figureCitation>
).
</paragraph>
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<paragraph id="740D369E126BFFEA00A2E7DA62F5A0C5" blockId="13.[151,1436,1467,1672]" pageId="13" pageNumber="416">
<emphasis id="46C6EA8C126BFFEA00A2E7DA6036A396" bold="true" box="[151,340,1467,1492]" pageId="13" pageNumber="416">FIGURES 55–57.</emphasis>
Sites sampled in the search for
<taxonomicName id="B3B24D1D126BFFEA02ACE7DA625CA396" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[665,830,1467,1492]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="13" pageNumber="416" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126BFFEA02ACE7DA625CA396" box="[665,830,1467,1492]" italics="true" pageId="13" pageNumber="416">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126BFFEA0370E7DD62F4A396" bold="true" box="[837,918,1468,1492]" pageId="13" pageNumber="416">
<taxonomicNameLabel id="5DF557F7126BFFEA0370E7DD62F4A396" box="[837,918,1468,1492]" pageId="13" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens. 55: Overview of the area including the type-locality (yellow square), the 2012–2013 sampling site (yellow triangle) and searched sites where no specimen was found (white circles). 56: Google Earth image of the type-locality in December 2013, depicting the preserved riparian forest in both sampling sites where
<taxonomicName id="B3B24D1D126BFFEA0140E44960DBA002" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[373,441,1576,1600]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="13" pageNumber="416" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126BFFEA0140E44960DBA002" box="[373,441,1576,1600]" italics="true" pageId="13" pageNumber="416">R. ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126BFFEA01F4E4496370A002" bold="true" box="[449,530,1576,1600]" pageId="13" pageNumber="416">
<taxonomicNameLabel id="5DF557F7126BFFEA01F4E4496370A002" box="[449,530,1576,1600]" pageId="13" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were observed. 57: Google Earth image from August 2020, after the riparian forest being cleared and burnt in the 2012–2013 sampling site, where
<taxonomicName id="B3B24D1D126BFFEA0352E42D62C9A026" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[871,939,1612,1636]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="13" pageNumber="416" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C126BFFEA0352E42D62C9A026" box="[871,939,1612,1636]" italics="true" pageId="13" pageNumber="416">R. ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C126BFFEA0386E42D6567A026" bold="true" box="[947,1029,1612,1636]" pageId="13" pageNumber="416">
<taxonomicNameLabel id="5DF557F7126BFFEA0386E42D6567A026" box="[947,1029,1612,1636]" pageId="13" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, no longer occurs. Image sources: Esri, Maxar, Earthstar Geographics, the GIS User Community and Google Earth.
</paragraph>
</caption>
<paragraph id="740D369E126BFFEA00F2E4BB6017A1B2" blockId="13.[151,1436,1754,2032]" pageId="13" pageNumber="416">
The available data suggests that this species should be considered as critically endangered, in accordance with the criteria B1B2ab (iii+iv) of the IUCN Red List (
<bibRefCitation id="10234B6F126BFFEA02E7E49F623EA15A" author="IUCN" box="[722,860,1790,1816]" pageId="13" pageNumber="416" refId="ref7599" refString="IUCN. 2001. IUCN Red List Categories and Criteria: Version 3.1. Second Edition. IUCN, Gland, and Cambridge. Available from: https: // www. iucnredlist. org / resources / categories-and-criteria (acessed 1 April 2024)" type="url" year="2001">IUCN 2001</bibRefCitation>
). The reasons include: (1) the estimated occurrence extension of this species is less than
<quantity id="B34A9B7B126BFFEA020DE5436316A17C" box="[568,628,1826,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="416" unit="km" value="1.0">1 km</quantity>
²; (2) the species has only one known population; (3) decreasing habitat quality due to human activities; (4) absence of the species within conservation units; and (5) possibility of extinction of the species based on a single slash-and-burn, fire or flooding event. Additionally, further samplings should be carried out upstream and downstream the Parnaíba River, looking for relatively well-preserved riverine and gallery forests, in order to search for additional locations for this species and provide further data for a reassessment of its conservation status.
</paragraph>
<caption id="20CD66161268FFE900A2E656645FA231" ID-DOI="http://doi.org/10.5281/zenodo.11547630" ID-Zenodo-Dep="11547630" httpUri="https://zenodo.org/record/11547630/files/figure.png" pageId="14" pageNumber="417" startId="14.[151,250,1079,1103]" targetBox="[176,1413,187,1050]" targetPageId="14" targetType="figure">
<paragraph id="740D369E1268FFE900A2E656645FA231" blockId="14.[151,1436,1079,1139]" pageId="14" pageNumber="417">
<emphasis id="46C6EA8C1268FFE900A2E6566040A20D" bold="true" box="[151,290,1079,1103]" pageId="14" pageNumber="417">FIGURE 58.</emphasis>
Known geographic distribution of described schizomid species from continental South America, including the
<taxonomicName id="B3B24D1D1268FFE900A2E63A605EA231" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[151,316,1115,1139]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1268FFE900A2E63A605EA231" box="[151,316,1115,1139]" italics="true" pageId="14" pageNumber="417">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1268FFE90176E63A60F1A231" bold="true" box="[323,403,1115,1139]" pageId="14" pageNumber="417">
<taxonomicNameLabel id="5DF557F71268FFE90176E63A60F1A231" box="[323,403,1115,1139]" pageId="14" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red circle), the remaining
<taxonomicName id="B3B24D1D1268FFE9029CE63A6243A231" authorityName="Reddell &amp; Cokendolpher" authorityYear="1995" box="[681,801,1115,1139]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="46C6EA8C1268FFE9029CE63A6243A231" box="[681,801,1115,1139]" italics="true" pageId="14" pageNumber="417">Rowlandius</emphasis>
</taxonomicName>
taxa (yellow squares) and other taxa (blue triangles).
</paragraph>
</caption>
<paragraph id="740D369E1268FFE900F2E6A46087A061" blockId="14.[151,1437,1221,2040]" pageId="14" pageNumber="417">
The conservation status proposed for
<taxonomicName id="B3B24D1D1268FFE9025DE6A4627DA29D" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[616,799,1221,1247]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1268FFE9025DE6A4627DA29D" box="[616,799,1221,1247]" italics="true" pageId="14" pageNumber="417">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1268FFE90311E6A4621FA29D" bold="true" box="[804,893,1221,1247]" pageId="14" pageNumber="417">
<taxonomicNameLabel id="5DF557F71268FFE90311E6A4621FA29D" box="[804,893,1221,1247]" pageId="14" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is not unprecedented. Several other
<taxonomicName id="B3B24D1D1268FFE9052DE6A464FEA29D" authorityName="Petrunkevitch" authorityYear="1945" box="[1304,1436,1221,1247]" class="Arachnida" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="order">Schizomida</taxonomicName>
species have been suggested to be under differing threat categories (see
<bibRefCitation id="10234B6F1268FFE903E7E688653CA341" author="Teruel, R." box="[978,1118,1257,1283]" pageId="14" pageNumber="417" pagination="38 - 40" refId="ref8780" refString="Teruel, R. (2012) Estatus de conservacion del orden Schizomida (Arthropoda: Arachnida) en Cuba Oriental. Revista Iberica de Aracnologia, 21, 38 - 40." type="journal article" year="2012">Teruel 2012</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE9045CE68B6459A341" author="Armas, L. F. &amp; Diaz, R. B. &amp; Barroso, A. A." box="[1129,1339,1257,1284]" pageId="14" pageNumber="417" pagination="534 - 546" refId="ref7034" refString="Armas, L. F., Diaz, R. B. &amp; Barroso, A. A. (2017) Redescription of the monotypic genus Cubacanthozomus (Schizomida: Hubbardiidae) and conservation status of its type-species. Zootaxa, 4323 (4), 534 - 546. https: // doi. org / 10.11646 / zootaxa. 4323.4.6" type="journal article" year="2017">
Armas
<emphasis id="46C6EA8C1268FFE90488E68B6592A341" box="[1213,1264,1257,1283]" italics="true" pageId="14" pageNumber="417">et al</emphasis>
. 2017
</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE9057CE6886048A365" author="Ruiz, G. R. S. &amp; Valente, R. M." pageId="14" pageNumber="417" pagination="81 - 95" refId="ref8418" refString="Ruiz, G. R. S. &amp; Valente, R. M. (2017) The first schizomid from a dry forest in South America (Arachnida: Schizomida). Zootaxa, 4311 (1), 81 - 95. https: // doi. org / 10.11646 / zootaxa. 4311.1.5" type="journal article" year="2017">Ruiz &amp; Valente 2017</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE90106E76C6310A365" author="Castillo-Figueroa, D. &amp; Castillo-Avila, C. &amp; Moreno-Gonzalez, J. A. &amp; Posada, J. M." box="[307,626,1293,1319]" pageId="14" pageNumber="417" refId="ref7263" refString="Castillo-Figueroa, D., Castillo-Avila, C., Moreno-Gonzalez, J. A. &amp; Posada, J. M. (2024) Habitat of two threatened Schizomida in the tropical Andes of South America. Journal of Insect Conservation. [published online] https: // doi. org / 10.1007 / s 10841 - 024 - 00565 - 4" type="book" year="2024">
Castillo-Figueroa
<emphasis id="46C6EA8C1268FFE901C9E76F6353A365" box="[508,561,1293,1319]" italics="true" pageId="14" pageNumber="417">et al.</emphasis>
2024
</bibRefCitation>
). Recently, schizomids have been shown to be associated to environmental characteristics and highly sensitive to environmental changes (
<bibRefCitation id="10234B6F1268FFE9037BE75065F7A309" author="Castillo-Figueroa, D. &amp; Castillo-Avila, C. &amp; Moreno-Gonzalez, J. A. &amp; Posada, J. M." box="[846,1173,1329,1355]" pageId="14" pageNumber="417" refId="ref7263" refString="Castillo-Figueroa, D., Castillo-Avila, C., Moreno-Gonzalez, J. A. &amp; Posada, J. M. (2024) Habitat of two threatened Schizomida in the tropical Andes of South America. Journal of Insect Conservation. [published online] https: // doi. org / 10.1007 / s 10841 - 024 - 00565 - 4" type="book" year="2024">
Castillo-Figueroa
<emphasis id="46C6EA8C1268FFE9042EE7536530A309" box="[1051,1106,1329,1355]" italics="true" pageId="14" pageNumber="417">et al.</emphasis>
2024
</bibRefCitation>
). However, no previous study provided robust evidence of the effects of habitat quality decrease on schizomid populations.Herein, we showed that habitat degradation could lead to local extinction of schizomids. This allowed two conclusions: (1) epigean schizomids from savanna biomes might not be found outside ideal conditions provided by forested environments; and (2) these species require a good habitat quality to be preserved. Further studies should evaluate the effect of recovering degraded riverine and gallery forests to evaluate the reestablishment of schizomid populations, at previously reported localities.
</paragraph>
<paragraph id="740D369E1268FFE900F2E44C62F8A1ED" blockId="14.[151,1437,1221,2040]" pageId="14" pageNumber="417">
<taxonomicName id="B3B24D1D1268FFE900F2E44C601FA005" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[199,381,1581,1607]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1268FFE900F2E44C601FA005" box="[199,381,1581,1607]" italics="true" pageId="14" pageNumber="417">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1268FFE901B6E44C60BEA005" bold="true" box="[387,476,1581,1607]" pageId="14" pageNumber="417">
<taxonomicNameLabel id="5DF557F71268FFE901B6E44C60BEA005" box="[387,476,1581,1607]" pageId="14" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
stands out among most Brazilian schizomids because it has not been collected inside a conservation unit, neither inside caves, nor in forested biomes (e.g., Amazon and Atlantic forests). On the contraire, the new species was found in the leaf litter within a savanna biome. Its conservation status might be shared in some degree with other South American species, considering their usually narrow distribution range and the fact that several species are known only as a byproduct of other taxa sampling (e.g.,
<bibRefCitation id="10234B6F1268FFE903EEE4DC65AEA095" author="Pinto-da-Rocha, R." box="[987,1228,1725,1751]" pageId="14" pageNumber="417" pagination="265 - 267" refId="ref8341" refString="Pinto-da-Rocha, R. (1996) Surazomus chavin new species, first Schizomida (Hubbardiidae, Hubbardiinae) described from Peru. Journal of Arachnology, 24, 265 - 267." type="journal article" year="1996">Pinto-da-Rocha 1996</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE904E2E4DC6026A0B9" author="Bonaldo, A. &amp; Pinto-da-Rocha, R." pageId="14" pageNumber="417" pagination="323 - 326" refId="ref7098" refString="Bonaldo, A. &amp; Pinto-da-Rocha, R. (2007) A new species of Surazomus (Arachnida, Schizomida) from Brazilian Oriental Amazonia. Revista Brasileira de Zoologia, 24 (2), 323 - 326. https: // doi. org / 10.1590 / S 0101 - 81752007000200009" type="journal article" year="2007">Bonaldo &amp; Pinto-da-Rocha 2007</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE90165E4806379A0B9" author="Santos, A. J. &amp; Dias, S. C. &amp; Brescovit, A. D. &amp; Santos, P. P." box="[336,539,1761,1787]" pageId="14" pageNumber="417" pagination="53 - 60" refId="ref8625" refString="Santos, A. J., Dias, S. C., Brescovit, A. D. &amp; Santos, P. P. (2008) The arachnid order Schizomida in the Brazilian Atlantic Forest: a new species of Rowlandius and new records of Stenochrus portoricensis (Schizomida: Hubbardiidae). Zootaxa, 1850 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 1850.1.4" type="journal article" year="2008">
Santos
<emphasis id="46C6EA8C1268FFE90197E48360B8A0B9" box="[418,474,1761,1787]" italics="true" pageId="14" pageNumber="417">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="10234B6F1268FFE90212E4806277A0B9" author="Ruiz, G. R. S. &amp; Valente, R. M." box="[551,789,1761,1787]" pageId="14" pageNumber="417" pagination="81 - 95" refId="ref8418" refString="Ruiz, G. R. S. &amp; Valente, R. M. (2017) The first schizomid from a dry forest in South America (Arachnida: Schizomida). Zootaxa, 4311 (1), 81 - 95. https: // doi. org / 10.11646 / zootaxa. 4311.1.5" type="journal article" year="2017">Ruiz &amp; Valente 2017</bibRefCitation>
,
<bibRefCitation id="10234B6F1268FFE90315E480623FA0B9" author="Ruiz, G. R. S. &amp; Valente, R. M." box="[800,861,1761,1787]" pageId="14" pageNumber="417" pagination="213268" refId="ref8475" refString="Ruiz, G. R. S. &amp; Valente, R. M. (2019) Description of a new species of Surazomus (Arachnida: Schizomida), with comments on homology of male flagellum and mating march anchorage in the genus. PLoS ONE, 14 (3), e 213268. https: // doi. org / 10.1371 / journal. pone. 0213268" type="journal article" year="2019">2019</bibRefCitation>
). The fact that all our recent field work has resulted in the discovery of an undescribed species, rather than new records of species previously known (or introduced species), suggest that its narrow distribution pattern is not necessarily a result of sampling deficiency. This implies that more (potentially endangered) species await discovery, and highlights the relevance of long-term sampling for properly describing schizomid species. In fact, the first heteromorphic male was collected only eight years after the first homeomorphic male and female of the newly described species.
</paragraph>
<paragraph id="740D369E1268FFE800F2E5D862ABA75F" blockId="14.[151,1437,1221,2040]" lastBlockId="15.[151,1436,151,285]" lastPageId="15" lastPageNumber="418" pageId="14" pageNumber="417">
<emphasis id="46C6EA8C1268FFE900F2E5D86064A191" bold="true" box="[199,262,1977,2003]" pageId="14" pageNumber="417">Note.</emphasis>
<taxonomicName id="B3B24D1D1268FFE90127E5D860ACA191" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[274,462,1977,2003]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1268FFE90127E5D860ACA191" box="[274,462,1977,2003]" italics="true" pageId="14" pageNumber="417">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1268FFE901ECE5D8635AA191" bold="true" box="[473,568,1977,2003]" pageId="14" pageNumber="417">
<taxonomicNameLabel id="5DF557F71268FFE901ECE5D8635AA191" box="[473,568,1977,2003]" pageId="14" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="B3B24D1D1268FFE902C3E5D8657EA191" authorityName="Rowland &amp; Reddell" authorityYear="1979" box="[758,1052,1977,2003]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="dumitrescoae">
<emphasis id="46C6EA8C1268FFE902C3E5D8657EA191" box="[758,1052,1977,2003]" italics="true" pageId="14" pageNumber="417">Rowlandius dumitrescoae</emphasis>
</taxonomicName>
species group. The diagnosis of this species group was revised by
<bibRefCitation id="10234B6F1268FFE9022BE5BC627DA1B5" author="Giupponi, A. P. L. &amp; Miranda, G. S. &amp; Villarreal, O. M." box="[542,799,2013,2039]" pageId="14" pageNumber="417" pagination="13 - 34" refId="ref7686" refString="Giupponi, A. P. L., Miranda, G. S. &amp; Villarreal, O. M. (2016) Rowlandius dumitrescoae species group: New diagnosis, key and description of new cave-dwelling species from Brazil (Schizomida, Hubbardiidae). ZooKeys, 632, 13 - 34. https: // doi. org / 10.3897 / zookeys. 632.9337" type="journal article" year="2016">
Giupponi
<emphasis id="46C6EA8C1268FFE902A5E5BF63A8A1B5" box="[656,714,2013,2039]" italics="true" pageId="14" pageNumber="417">et al.</emphasis>
(2016)
</bibRefCitation>
, who considered the absence of a gonopod a distinctive character. However, these authors missed the presence of a gonopod described and illustrated for
<taxonomicName id="B3B24D1D1269FFE804D2E2F66405A6F2" authorityName="Santos, Ferreira &amp; Buzatto" authorityYear="2013" box="[1255,1383,151,176]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="15" pageNumber="418" phylum="Arthropoda" rank="species" species="potiguar">
<emphasis id="46C6EA8C1269FFE804D2E2F66405A6F2" box="[1255,1383,151,176]" italics="true" pageId="15" pageNumber="418">R. potiguar</emphasis>
</taxonomicName>
(see
<bibRefCitation id="10234B6F1269FFE800A2E2DA6006A697" author="Santos, A. J. &amp; Ferreira, R. L. &amp; Buzatto, B. A." box="[151,356,187,213]" pageId="15" pageNumber="418" pagination="63616" refId="ref8705" refString="Santos, A. J., Ferreira, R. L. &amp; Buzatto, B. A. (2013) Two new cave-dwelling species of the short-tailed whipscorpion genus Rowlandius (Arachnida: Schizomida: Hubbardiidae) from Northeastern Brazil, with comments on male dimorphism. PLoS ONE, 8 (5), e 63616. https: // doi. org / 10.1371 / journal. pone. 0063616" type="journal article" year="2013">
Santos
<emphasis id="46C6EA8C1269FFE800DCE2DA6043A697" box="[233,289,187,213]" italics="true" pageId="15" pageNumber="418">et al.</emphasis>
2013
</bibRefCitation>
: figs 4G, 6F). The presence of a gonopod in
<taxonomicName id="B3B24D1D1269FFE80363E2DA656FA697" authority="Silva &amp; Santos &amp; Carvalho, 2024" authorityName="Silva &amp; Santos &amp; Carvalho" authorityYear="2024" box="[854,1037,187,213]" class="Arachnida" family="Hubbardiidae" genus="Rowlandius" kingdom="Animalia" order="Schizomida" pageId="15" pageNumber="418" phylum="Arthropoda" rank="species" species="ufpi" status="sp. nov.">
<emphasis id="46C6EA8C1269FFE80363E2DA656FA697" box="[854,1037,187,213]" italics="true" pageId="15" pageNumber="418">Rowlandius ufpi</emphasis>
</taxonomicName>
<emphasis id="46C6EA8C1269FFE80421E2DA650FA697" bold="true" box="[1044,1133,187,213]" pageId="15" pageNumber="418">
<taxonomicNameLabel id="5DF557F71269FFE80421E2DA650FA697" box="[1044,1133,187,213]" pageId="15" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
suggests that this structure might be present in other species of the group, for which it has not been reported. This absence might be explained by difficulties in visualizing this hyaline structure (compare
<figureCitation id="EC892A1B1269FFE8030EE36262DCA75F" box="[827,958,258,285]" captionStart="FIGURES 47–50" captionStartId="11.[151,264,1226,1250]" captionTargetBox="[151,1436,342,1200]" captionTargetId="figure-93@11.[151,1436,342,1200]" captionTargetPageId="11" captionText="FIGURES 47–50. Rowlandius ufpi sp. nov. female, paratypes, variation in the internal genitalia. 47: CHNUFPI 4064; 48: CHNUFPI 4057; 49: CHNUFPI 4211; 50: CHNUFPI 4065. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11547622" httpUri="https://zenodo.org/record/11547622/files/figure.png" pageId="15" pageNumber="418">Figs 47–50</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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