160 lines
17 KiB
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160 lines
17 KiB
XML
<document id="10835BF1457FC28B3BDF77B95B371B8B" ID-DOI="http://dx.doi.org/10.3897/subtbiol.19.9513" ID-GBIF-Dataset="09e0e283-8cca-446d-9985-dcbd94157a8c" ID-Pensoft-Pub="1314-2615--23" ID-ZooBank="47000C3AA8C44BC6BB4C063C46D20BD6" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2615--23" ModsDocOrigin="Subterranean Biology " ModsDocTitle="A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1553126978140" checkinUser="pensoft" docAuthor="Watts, Chris H. S., Hendrich, Lars & Balke, Michael" docDate="2016" docId="C6105444DA9071B9F0133A816B19AC51" docLanguage="en" docName="SubterBiol 19: 23-29" docOrigin="Subterranean Biology 19" docSource="http://dx.doi.org/10.3897/subtbiol.19.9513" docTitle="Exocelina saltusholmesensis Watts, Hendrich & Balke, 2016, sp. n." docType="treatment" docUuid="90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="26" masterDocId="FFBA6A219A2AFF8BFFFBFFE1FF81FFC3" masterDocTitle="A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="29" masterPageNumber="23" pageNumber="24" updateTime="1701317740128" updateUser="plazi">
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<mods:titleInfo id="65FA31DD84FA0209F6E34C4FA31A9742">
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<mods:title id="14C34F52FD1EFD15FF8147DE801F2A7B">A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:name id="55883BE6D0ADE5414108234D07DCC86A" type="personal">
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<mods:roleTerm id="C39F26CFC3EDF729CECF7FD531B136B8">Author</mods:roleTerm>
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<mods:namePart id="850D2931A398CA51C732D7F39B63AA1A">Watts, Chris H. S.</mods:namePart>
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<mods:namePart id="51F3C36F4F9908620A6C6E15137A89C7">Hendrich, Lars</mods:namePart>
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<mods:name id="B0D5B479DBA3525B2803D81992ACE873" type="personal">
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<mods:roleTerm id="0E06BB3204B6D7CB3C05524937724A2E">Author</mods:roleTerm>
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<mods:namePart id="E39C718043D164C2F1A17285AE34ED42">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="AC989F7C70564FAFE804D946FB0287BC">text</mods:typeOfResource>
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<mods:title id="3D6EE7D04E847CF4310B8200B758C9AA">Subterranean Biology</mods:title>
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<mods:part id="1F0FD42004AD80B57CE781DC86E889FA">
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<mods:date id="D71E6557AC6FF4EAC39E6E796B5A19D5">2016</mods:date>
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<mods:number id="CFBC708FBF8D69580379E52714C403F3">19</mods:number>
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<mods:url id="1014BC9CB8C4E99751AB609541755614">http://dx.doi.org/10.3897/subtbiol.19.9513</mods:url>
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<mods:classification id="0959E5789AD1BA957A99662F88D65F3D">journal article</mods:classification>
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<mods:identifier id="DAAC71E2FCD4A9127D28BC23AA581429" type="DOI">http://dx.doi.org/10.3897/subtbiol.19.9513</mods:identifier>
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<mods:identifier id="30F0C690FF057ABE05805BF16E376F00" type="Pensoft-Pub">1314-2615--23</mods:identifier>
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<treatment id="C6105444DA9071B9F0133A816B19AC51" ID-GBIF-Taxon="154474303" LSID="urn:lsid:zoobank.org:act:90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" httpUri="http://treatment.plazi.org/id/C6105444DA9071B9F0133A816B19AC51" lastPageId="3" lastPageNumber="26" pageId="1" pageNumber="24">
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<subSubSection id="005298E1203CFBD5BF7244707472D58C" pageId="1" pageNumber="24" type="nomenclature">
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<paragraph id="62DEC8ED43061B752C88398D211F4D4A" pageId="1" pageNumber="24">
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<taxonomicName id="BAA24595B9ECD6AD7D0F6B2EE78A516F" ID-CoL="8LXT8" LSID="http://zoobank.org/90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina saltusholmesensis" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="saltusholmesensis">Exocelina saltusholmesensis</taxonomicName>
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<taxonomicNameLabel id="5C726019D630F90286D013742A8DAA0C" pageId="1" pageNumber="24">sp. n.</taxonomicNameLabel>
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Figs 1, 3
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</paragraph>
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</subSubSection>
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||
<subSubSection id="DB5595B4C3B758F5B88828C5164B582C" pageId="1" pageNumber="24" type="type locality">
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<paragraph id="1359A681518DB674E30E71AF8B0910F0" pageId="1" pageNumber="24">Type locality.</paragraph>
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<paragraph id="B5D25E3239A5F54242BB61FD088711D7" pageId="1" pageNumber="24">Small seasonal creek [-12.400714° 130.931297°], Holmes Jungle Reserve, Darwin, Northern Territory, Australia.</paragraph>
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</subSubSection>
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<subSubSection id="BE3E9FE49DADB5C52CE3A243C7E98CD8" pageId="1" pageNumber="24" type="type material">
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<paragraph id="1FC41706325E019CFF8B408DAF4A4FD3" pageId="1" pageNumber="24">Type material.</paragraph>
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<paragraph id="53920EC2BCBA3F451473DE8B8D9D346A" pageId="1" pageNumber="24">
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Holotype, female (South Australian Museum Adelaide, SAMA): "Australia, Northern Territory, Darwin, Holmes Jungle Reserve, -12.400714° 130.931297°, 25. March 2004, C.H.S. Watts (SAMA database no. 25-012926)". The beetle was originally labelled as "
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<taxonomicName id="C54B894A832E3824D760F6B173ABC970" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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sp.nov." by C.H.S. Watts and has now a red, printed holotype label attached in addition.
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</paragraph>
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<paragraph id="A33D5CE63059296C45355D22DDAF9ABA" pageId="1" pageNumber="24">European nucleotide archive (genbank) accession number for the 3' end of the mitochondrial cytochrome c oxidase 1 gene: www.ebi.ac.uk/ena/data/view/LN994839</paragraph>
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</subSubSection>
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<subSubSection id="617201DB4696E634BF050ECB1B8D492C" lastPageId="3" lastPageNumber="26" pageId="1" pageNumber="24" type="description">
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<paragraph id="02720759CEF6ACD76532DB88C69AAA57" pageId="1" pageNumber="24">Description.</paragraph>
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<paragraph id="A14C1A05A3D02E76596CD855D649D29E" pageId="1" pageNumber="24">Size and shape: Beetle small (length of beetle 3.7 mm, greatest width 1.8 mm, length of pronotum 0.6 mm, width of pronotum at base 1.7 mm), comparably parallel-sided (Fig. 1C).</paragraph>
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<paragraph id="3704F52B72A567D7D163F46F73623FD7" pageId="1" pageNumber="24">Coloration. Mainly testaceous and slightly translucent with lighter band on elytral base and darker area in middle basal area of pronotum (Fig. 1C).</paragraph>
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<paragraph id="31C7778A1FD4776E71D03FF2521BE4D3" pageId="2" pageNumber="25">
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<pageBreakToken id="82F3BD0FBA6053A3A6BC61F50AC6679A" pageId="2" pageNumber="25" start="start">Surface</pageBreakToken>
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sculpture. Head (frons), pronotum and elytra striolate; with distinct mircoreticulation and coarser punctation. Metacoxal plates and ventrites 1-4 with few strioles, otherwise with fine mircoreticulation and few punctures. The sixth (apical) ventrite broadly rounded.
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</paragraph>
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<paragraph id="80F9628A2CCAD648F63203143E18A75C" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
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Structures. Eyes small in comparison to epigean
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<taxonomicName id="3A04584F17FDD2418A3231709200DF17" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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, length of eye c. 0.16 mm, distance between eyes c. 0.86 mm (Figs 1C, 3B). Pronotum with faint lateral
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<pageBreakToken id="DA97D8355CA2108C42D3FC19CE85EE2A" pageId="3" pageNumber="26" start="start">bead</pageBreakToken>
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that does not reach anterior corner. Prosternal process lanceolate, gently rounded ventrally; lateral wings of metaventrite narrow. Flight wings vestigial.
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</paragraph>
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<paragraph id="8D09C1C5CB770BDF53B5FC8E9CCC3416" pageId="3" pageNumber="26">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="9B8418760A41354568E5BCB88CBAAD59" pageId="3" pageNumber="26" type="etymology">
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<paragraph id="5E4C6F462CAFE51CB4661B1DD93D9227" pageId="3" pageNumber="26">Etymology.</paragraph>
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<paragraph id="EEF0EE9DEA11DD16A9FCC249CBB13921" pageId="3" pageNumber="26">The name is an adjective is the nominative singular derived from the locality name - Holmes Jungle. Latin: saltus = woodland.</paragraph>
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</subSubSection>
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<subSubSection id="BCC41BA08AD37151B86AEAF610C2CEE1" pageId="3" pageNumber="26" type="differential diagnosis">
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<paragraph id="7A305673680A63A022143FAC4DE35241" pageId="3" pageNumber="26">Differential diagnosis.</paragraph>
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<paragraph id="3FD9B474AA18660B71EA26D7445D6A50" pageId="3" pageNumber="26">
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The species is sister to a clade containing
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<taxonomicName id="02991B69335E798349ADF84B0EEAEB01" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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(Lea, 1899) and
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<taxonomicName id="C1767F27DF8551562CDB969DBA68FB86" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(Sharp, 1882) established based on DNA sequence data (
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<bibRefCitation id="5A5A480EE3CCB35ED254F5D1AC169400" author="Toussaint, EF" journalOrPublisher="Scientific Reports" pageId="4" pageNumber="27" pagination=": e 16016" title="Mosaic patterns of diversification dynamics following the colonization of Melanesian islands." url="10.1038/srep16016" volume="5" year="2015">Toussaint et al. 2015</bibRefCitation>
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), but both these species are larger (> 5mm), and have larger eyes (Figs 1
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<normalizedToken id="B7E8E32BD985C1D85ABBB8A2087E1E9E" originalValue="A–C">A-C</normalizedToken>
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, 3A, B). Both are epigean species, distributed along the eastcoast of Queensland (
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<taxonomicName id="FE2080A2ABFF9C0F2329E162F9C8C481" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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) and the Adelaide Hills in South Australia (
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<taxonomicName id="CB83EDE69FECB41588680F2C52C40146" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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) (Hendrich et al. in prep.). They are also fully winged.
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</paragraph>
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</subSubSection>
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<subSubSection id="FEC9B0F5BB61AAA0A3DFD4C75B2908BA" pageId="3" pageNumber="26" type="habitat">
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<paragraph id="2FB1B08C826221467D7D770BA1AB9891" pageId="3" pageNumber="26">Habitat.</paragraph>
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<paragraph id="95AE10F152F2B645683C046418FE8C1A" pageId="3" pageNumber="26">
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A small pool in the bed of a small ephemeral creek through eucalypt woodland. A small eyed, wingless
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<taxonomicName id="ACC56C2F33DD016F561890DEBBC0D2C5" family="Hydraenidae" lsidName="" pageId="3" pageNumber="26" rank="family">Hydraenidae</taxonomicName>
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(
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<taxonomicName id="9B6090E8863BAF096F8109B3239A9A2B" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydraena wattsi" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="wattsi">Hydraena wattsi</taxonomicName>
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Perkins, 2007) was collected at the same time (
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<bibRefCitation id="D1EF56B7D8A6EE90D7A4ED988A68F57C" author="Perkins, PD" journalOrPublisher="Zootaxa" pageId="4" pageNumber="27" pagination="1 - 139" title="A revision of the Australian species of the water beetle genus Hydraena Kugelmann (Coleoptera: Hydraenidae)." volume="1489" year="2007">Perkins 2007</bibRefCitation>
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). During the dry season (
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<normalizedToken id="5DDCE9F48AEF651676EB6FA44FC3A652" originalValue="June–September">June-September</normalizedToken>
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) this creek dries out. However, we assume that subsurface water remains in the watercourse during this season providing an interstitial habitat for these flightless, low vagility species.
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</paragraph>
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<caption id="15F84E1056C2BF138B3043BC6606D0A4" pageId="3" pageNumber="26">
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<paragraph id="E06878A91FB0B77918B773B54600C628" pageId="3" pageNumber="26">
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Figure 1. Habitus of
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<taxonomicName id="59ABD32AB41F0D650EDE49E70451D358" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="C506A285EEBE31FCBF5BE75540494A15" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(A),
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<taxonomicName id="1718BBB7512078B68BEB589CEB93AD84" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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(B),
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<taxonomicName id="7BE4AF54FA6F489C274A67E30D9F083D" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina saltusholmesensis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="saltusholmesensis">Exocelina saltusholmesensis</taxonomicName>
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sp. n. (C).
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</paragraph>
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</caption>
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<caption id="40CF763ED12DC34741A573BF8D82B8C7" pageId="3" pageNumber="26">
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<paragraph id="B663A259229A93BEE1F7DE57FD40E142" pageId="3" pageNumber="26">
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Figure 2. Habitus of
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<taxonomicName id="FCAABAF8DF41F44ADA29DDC7BFBB60AA" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="48DB99FB39ADDCCF448F11A8460B1D27" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina australis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="australis">Exocelina australis</taxonomicName>
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(A),
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<taxonomicName id="12C3350564FA8481EB6069DD1B16A8E6" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina abdita" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="abdita">Exocelina abdita</taxonomicName>
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(B).
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</paragraph>
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</caption>
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<caption id="22A28D33B44F11766BE9253E45486E3D" pageId="3" pageNumber="26">
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<paragraph id="6EB30AA77D921E8A63F0B077A0A6A795" pageId="3" pageNumber="26">
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Figure 3. Right lateral aspect of head of
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<taxonomicName id="A443F0AB4C8CE14CD1764AFF05B93377" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="AD6A411FE76A3AE56211030ADC723A89" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(A), E.
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<taxonomicName id="8286357225BC75D51F19BB960087D44E" lsidName="saltusholmesensis" pageId="3" pageNumber="26" rank="species" species="saltusholmesensis">saltusholmesensis</taxonomicName>
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sp. n. (B),
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<taxonomicName id="69E477ACCB348C5FA59AEEE5B0A545A1" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina abdita" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="abdita">Exocelina abdita</taxonomicName>
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(C).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |