treatments-xml/data/C3/76/87/C37687EFEC0A225FAD86F8CDFAB21684.xml

<document id="4CCECC4A08724AB39EA651244BF3839F" ID-CLB-Dataset="8469" ID-DOI="10.24349/acarologia/20184236" ID-GBIF-Dataset="3c0084e1-d3c9-4633-8e39-1513f2148596" ID-ISSN="2107-7207" ID-Zenodo-Dep="4487493" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612204954064" checkinUser="felipe" docAuthor="Pešić, Vladimir &amp; Smit, Harry" docDate="2018" docId="C37687EFEC0A225FAD86F8CDFAB21684" docLanguage="en" docName="Acarologia.58.1.165-185.pdf" docOrigin="Acarologia 58 (1)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Atractides (Atractides) alaarchaensis Pešić &amp; Smit 2018, n. sp." docType="treatment" docUuid="51EA2777-8191-4C77-876B-0ACE5B269136" docUuidSource="ZooBank" docVersion="7" lastPageNumber="180" masterDocId="3F4FFF97EC01224FAC6DFF85FF96111D" masterDocTitle="A checklist of the water mites of Central Asia with description of six new species (Acari, Hydrachnidia) from Kyrgyzstan" masterLastPageNumber="185" masterPageNumber="165" pageNumber="175" updateTime="1698925910332" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C117CAD403D4BA04D7B05587CB0E81DE">A checklist of the water mites of Central Asia with description of six new species (Acari, Hydrachnidia) from Kyrgyzstan</mods:title>
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<mods:namePart id="25BE1AFE9EDF3B7F3C37885F5AF26B79">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="FC4E61029C349141A86E5099661CB88D">Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro.</mods:affiliation>
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<mods:namePart id="9E5DA854772356296CA8D887F12A425C">Smit, Harry</mods:namePart>
<mods:affiliation id="3AABCF80E589791CCB90B3469CE7B9E3">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands.</mods:affiliation>
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<treatment id="C37687EFEC0A225FAD86F8CDFAB21684" ID-DOI="http://doi.org/10.5281/zenodo.4501599" ID-GBIF-Taxon="177210028" ID-Zenodo-Dep="4501599" LSID="urn:lsid:zoobank.org:act:51EA2777-8191-4C77-876B-0ACE5B269136" httpUri="http://treatment.plazi.org/id/C37687EFEC0A225FAD86F8CDFAB21684" lastPageId="16" lastPageNumber="180" pageId="11" pageNumber="175">
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<paragraph id="4B6036F9EC0A2244AD86F8CDFB641679" blockId="11.[491,1266,1864,1892]" box="[491,1266,1864,1892]" pageId="11" pageNumber="175">
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<taxonomicName id="8CDF4D7AEC0A2244AD86F8CDFC641679" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[491,1010,1864,1892]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="175" phylum="Arthropoda" rank="species" species="alaarchaensis" status="sp. nov." subGenus="Atractides">
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(
<emphasis id="79ABEAEBEC0A2244AEE0F8CDFC8B1679" bold="true" box="[653,797,1864,1892]" italics="true" pageId="11" pageNumber="175">Atractides</emphasis>
) 
<emphasis id="79ABEAEBEC0A2244AF42F8CDFC641679" bold="true" box="[815,1010,1864,1892]" italics="true" pageId="11" pageNumber="175">alaarchaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62985790EC0A2244AF97F8CDFBD01679" box="[1018,1094,1864,1892]" pageId="11" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="D3E42A7CEC0A2244A834F8CDFB7F1679" box="[1113,1257,1864,1892]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="14.[119,186,1863,1884]" captionStartId-1="15.[119,186,1863,1884]" captionStartId-2="16.[515,582,1414,1435]" captionTargetBox-0="[295,1305,306,1804]" captionTargetBox-1="[250,1342,307,1800]" captionTargetBox-2="[513,1474,235,1328]" captionTargetId-0="figure-2@14.[265,1333,297,1809]" captionTargetId-1="figure-2@15.[248,1350,297,1809]" captionTargetId-2="figure-2@16.[503,1487,221,1360]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="Figure 5 Atractides alaarchaensis n. sp., male holotype: A – idiosoma, ventral view; B – I-L-5 and -6; C – palp, medial view; D – palp, lateral view; E – genital field. Scale bars = 100 μm." captionText-1="Figure 6 Atractides alaarchaensis n. sp., female paratype: A – coxal field; B – I-L-5 and -6; C – palp, lateral view; D – genital field. Scale bars = 100 μm." captionText-2="Figure 7 Atractides alaarchaensis n. sp., deutonymph: A – I-L-5 and -6; B – palp, medial view; C – genital field. Scale bars = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.4487503" figureDoi-1="http://doi.org/10.5281/zenodo.4487505" figureDoi-2="http://doi.org/10.5281/zenodo.4487507" httpUri-0="https://zenodo.org/record/4487503/files/figure.png" httpUri-1="https://zenodo.org/record/4487505/files/figure.png" httpUri-2="https://zenodo.org/record/4487507/files/figure.png" pageId="11" pageNumber="175">Figure 5-7</figureCitation>
)
</emphasis>
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</paragraph>
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Zoobank: 
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51EA2777-8191-4C77-876B-0 
<collectingRegion id="891BF81BEC0A2244AF34F807FC101688" box="[857,902,1922,1941]" country="Indonesia" name="Aceh" pageId="11" pageNumber="175">ACE</collectingRegion>
5B269136
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<paragraph id="4B6036F9EC0A2242AE7EF841FD50103D" blockId="11.[491,1451,1987,2045]" lastBlockId="13.[491,1509,229,1356]" lastPageId="13" lastPageNumber="177" pageId="11" pageNumber="175">
Type material — 
<materialsCitation id="FBB73CA4EC0A2244AEA3F846FA0316E0" ID-GBIF-Occurrence="3029771305" collectingDate="2013-07-09" collectionCode="RMNH" collectorName="Ala Archa NP &amp; Pesic" country="Kyrgyzstan" elevation="1899" latitude="42.603367" location="Kyrgyzstan" longLatPrecision="1" longitude="74.48061" pageId="11" pageNumber="175" specimenCount="1" typeStatus="holotype">
<typeStatus id="9464885BEC0A2244AEA3F846FCBA16C6" box="[718,812,1987,2011]" pageId="11" pageNumber="175" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="2DCEAE3CEC0A2244AF1AF846FC5A16C6" box="[887,972,1987,2011]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="175" type="Museum">RMNH</collectionCode>
), 
<collectingCountry id="33C87669EC0A2244AF8DF841FBC216C1" box="[992,1108,1988,2012]" name="Kyrgyzstan" pageId="11" pageNumber="175">Kyrgyzstan</collectingCountry>
, KR3 
<collectorName id="E62A532FEC0A2244A8FBF846FAB916C6" box="[1174,1327,1987,2011]" pageId="11" pageNumber="175">Ala Archa NP</collectorName>
, rheocrenic spring and first order stream, 
<geoCoordinate id="2EEB503EEC0A2244AF40F860FC3C16E0" box="[813,938,2021,2045]" degrees="42" direction="north" minutes="36.202" orientation="latitude" pageId="11" pageNumber="175" precision="1" value="42.603367">42°36.202’N</geoCoordinate>
, 
<geoCoordinate id="2EEB503EEC0A2244AFDEF860FBBB16E0" box="[947,1069,2021,2045]" degrees="074" direction="east" minutes="28.837" orientation="longitude" pageId="11" pageNumber="175" precision="1" value="74.48061">074°28.837’E</geoCoordinate>
, 
<elevation id="C0F2D1CAEC0A2244A858F860FB0D16E0" box="[1077,1179,2021,2045]" metricMagnitude="3" metricUnit="m" metricValue="1.899" pageId="11" pageNumber="175" unit="m" value="1899.0">
<quantity id="8C279B1CEC0A2244A858F860FBE516E0" box="[1077,1139,2021,2045]" metricMagnitude="3" metricUnit="m" metricValue="1.899" pageId="11" pageNumber="175" unit="m" value="1899.0">1899 m</quantity>
asl
</elevation>
., 
<date id="3F611039EC0A2244A8C4F860FA9F16E0" box="[1193,1289,2021,2045]" pageId="11" pageNumber="175" value="2013-07-09">
<collectingDate id="2F25E9D1EC0A2244A8C4F860FA9F16E0" box="[1193,1289,2021,2045]" pageId="11" pageNumber="175" value="2013-07-09">9.vii.2013</collectingDate>
</date>
, leg. 
<collectorName id="E62A532FEC0A2244A92FF860FAEC16E0" box="[1346,1402,2021,2045]" pageId="11" pageNumber="175">Pešić</collectorName>
&amp;
</materialsCitation>
Smit, dissected and slide mounted. 
<typeStatus id="9464885BEC0C2242AFEFFF63FC7F11E3" box="[898,1001,230,254]" pageId="13" pageNumber="177" type="paratype">Paratypes</typeStatus>
: 12/27/3, same data as 
<typeStatus id="9464885BEC0C2242A8BFFF60FABB11E0" box="[1234,1325,229,253]" pageId="13" pageNumber="177" type="holotype">holotype</typeStatus>
, 2/1/1, dissected and slide mounted.
</paragraph>
<caption id="1FA06671EC0D2243AC1AF8C2FCB7166E" ID-DOI="http://doi.org/10.5281/zenodo.4487501" ID-Zenodo-Dep="4487501" httpUri="https://zenodo.org/record/4487501/files/figure.png" pageId="12" pageNumber="176" startId="12.[119,186,1863,1884]" targetBox="[274,1327,301,1795]" targetPageId="12">
<paragraph id="4B6036F9EC0D2243AC1AF8C2FCB7166E" blockId="12.[119,1500,1862,1907]" pageId="12" pageNumber="176">
<emphasis id="79ABEAEBEC0D2243AC1AF8C2FF5B1641" bold="true" box="[119,205,1863,1884]" pageId="12" pageNumber="176">Figure 4</emphasis>
<taxonomicName id="8CDF4D7AEC0D2243ACB4F8C3FE111641" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[217,391,1862,1884]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="176" phylum="Arthropoda" rank="species" species="manasi" status="sp. nov.">
<emphasis id="79ABEAEBEC0D2243ACB4F8C3FE111641" box="[217,391,1862,1884]" italics="true" pageId="12" pageNumber="176">Atractides manasi</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC0D2243ADE0F8C2FE501641" bold="true" box="[397,454,1863,1884]" pageId="12" pageNumber="176">
<taxonomicNameLabel id="62985790EC0D2243ADE0F8C2FE501641" box="[397,454,1863,1884]" pageId="12" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
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, female holotype: A – idiosoma, dorsal view; B – idiosoma, ventral view; C – genital field; D – I-L-5 and -6; E – palp, medial view; F – palp, lateral view. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="4B6036F9EC0C2242AE7EFEAFFAD810B7" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">Diagnosis — Coxae without sexual dimorphism (in both sexes Cx-I+II close to Cx-III+IV at their touching edges, but not forming a coxal shield, Vgl-3 not fused to Cx-IV); P-2 ventral margin convex, without distal projection, P-3 with one seta each on medial and lateral surface extending beyond ventral margin of segment; in male S-1-2 separation 19-21 μm.</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFE31FC411238" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">
Description — Adults: 
<emphasis id="79ABEAEBEC0C2242AF62FE31FC5410D1" box="[783,962,436,460]" italics="true" pageId="13" pageNumber="177">General features</emphasis>
— Integument striated, muscle insertions unsclerotized. Excretory pore smooth; Vgl-1 not fused to Vgl-2. Palp with weak sexual dimorphism, P-2 ventral margin convex, without projection, P-3 with one seta each on medial and lateral surface, P-4 sword seta between ventral setae. I-L-5 with S-1 and -2 separated, S-1 longish and blunt, S-2 pointed and thicker than S-1, I-L-6 curved, basally thickened, from the centre to the claw furrow with parallel dorsal and ventral margins (
<figureCitation id="D3E42A7CEC0C2242A810FDE4FB781364" box="[1149,1262,609,633]" captionStart="Figure 5" captionStartId="14.[119,186,1863,1884]" captionTargetBox="[295,1305,306,1804]" captionTargetId="figure-2@14.[265,1333,297,1809]" captionTargetPageId="14" captionText="Figure 5 Atractides alaarchaensis n. sp., male holotype: A – idiosoma, ventral view; B – I-L-5 and -6; C – palp, medial view; D – palp, lateral view; E – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487503" httpUri="https://zenodo.org/record/4487503/files/figure.png" pageId="13" pageNumber="177">Figure 5A</figureCitation>
). Male: Genital plate anterior margin equally convex, with a narrow border of secondary sclerotization, posterior margin indented, acetabula in an obtuse triangle (
<figureCitation id="D3E42A7CEC0C2242A860FD20FBEC13A0" box="[1037,1146,677,701]" captionStart="Figure 5" captionStartId="14.[119,186,1863,1884]" captionTargetBox="[295,1305,306,1804]" captionTargetId="figure-2@14.[265,1333,297,1809]" captionTargetPageId="14" captionText="Figure 5 Atractides alaarchaensis n. sp., male holotype: A – idiosoma, ventral view; B – I-L-5 and -6; C – palp, medial view; D – palp, lateral view; E – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487503" httpUri="https://zenodo.org/record/4487503/files/figure.png" pageId="13" pageNumber="177">Figure 5E</figureCitation>
); P-2 ventrodistally convex, P-4 thickened, ventral margin slightly projecting near insertion of proximoventral seta (
<figureCitation id="D3E42A7CEC0C2242A91DFD4DFE61121F" captionStart="Figure 5" captionStartId="14.[119,186,1863,1884]" captionTargetBox="[295,1305,306,1804]" captionTargetId="figure-2@14.[265,1333,297,1809]" captionTargetPageId="14" captionText="Figure 5 Atractides alaarchaensis n. sp., male holotype: A – idiosoma, ventral view; B – I-L-5 and -6; C – palp, medial view; D – palp, lateral view; E – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487503" httpUri="https://zenodo.org/record/4487503/files/figure.png" pageId="13" pageNumber="177">Figures 5</figureCitation>
C-D). Female. Acetabula in a weakly curved line (
<figureCitation id="D3E42A7CEC0C2242A872FD6FFB04121F" box="[1055,1170,746,770]" captionStart="Figure 6" captionStartId="15.[119,186,1863,1884]" captionTargetBox="[250,1342,307,1800]" captionTargetId="figure-2@15.[248,1350,297,1809]" captionTargetPageId="15" captionText="Figure 6 Atractides alaarchaensis n. sp., female paratype: A – coxal field; B – I-L-5 and -6; C – palp, lateral view; D – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487505" httpUri="https://zenodo.org/record/4487505/files/figure.png" pageId="13" pageNumber="177">Figure 6D</figureCitation>
); P-2 ventral margin slightly convex, P-4 slender than in male (
<figureCitation id="D3E42A7CEC0C2242AF37FC88FC5C1238" box="[858,970,781,805]" captionStart="Figure 6" captionStartId="15.[119,186,1863,1884]" captionTargetBox="[250,1342,307,1800]" captionTargetId="figure-2@15.[248,1350,297,1809]" captionTargetPageId="15" captionText="Figure 6 Atractides alaarchaensis n. sp., female paratype: A – coxal field; B – I-L-5 and -6; C – palp, lateral view; D – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487505" httpUri="https://zenodo.org/record/4487505/files/figure.png" pageId="13" pageNumber="177">Figure 6C</figureCitation>
).
</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFCAAFB1E1277" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">
Deutonymph: Provisional genital field with two pairs of acetabula (
<figureCitation id="D3E42A7CEC0C2242A88FFCAAFADD125A" box="[1250,1355,815,839]" captionStart="Figure 7" captionStartId="16.[515,582,1414,1435]" captionTargetBox="[513,1474,235,1328]" captionTargetId="figure-2@16.[503,1487,221,1360]" captionTargetPageId="16" captionText="Figure 7 Atractides alaarchaensis n. sp., deutonymph: A – I-L-5 and -6; B – palp, medial view; C – genital field. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487507" httpUri="https://zenodo.org/record/4487507/files/figure.png" pageId="13" pageNumber="177">Figure 7C</figureCitation>
); P-2 ventral margin convex, P-3 without seta on medial and lateral surface.
</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFCF0FDFD150B" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">
Measurements — Male (
<typeStatus id="9464885BEC0C2242AF73FCF1FCEA1291" box="[798,892,884,908]" pageId="13" pageNumber="177" type="holotype">holotype</typeStatus>
; in parentheses 
<typeStatus id="9464885BEC0C2242A847FCF0FB181290" box="[1066,1166,885,909]" pageId="13" pageNumber="177" type="paratype">paratypes</typeStatus>
, n = 2) — Idiosoma L 628 (631-656), W 416 (425-475); maximum diameter Dgl 26 (27-28); coxal field: L 328 (339-359), Cx-III W 347 (344-375), mL 113 (113-116), lL 204 (216); genital field L/W 117 (120-128)/123 (127-134), maximum diameter Ac 1-3: 38 (41-45), 39 (38-46), 40 (39-46); ejaculatory complex L 111 (125).
</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFBA4FB7815BD" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">Gnathosoma — palp total L 307 (304-317); dL/H, dL/H ratio: P-1, 33/28, 1.17 (31-34/28- 30, 1.1-1.2); P-2, 66/47, 1.4 (66/48, 1.36-1.37); P-3, 77/41, 1.89 (75-77/41, 1.85-1.89); P-4, 97/27, 3.6 (101-106/25, 4.0-4.2); P-5, 34/12, 2.9 (31-34/13, 2.5-2.7); L ratio P-2/P-4 0.68 (0.63-0.65). Gnathosomal base vL (106-111); chelicera total L (212-213).</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFB2EFD011451" blockId="13.[491,1509,229,1356]" pageId="13" pageNumber="177">Legs — I-L-5 dL 184 (186-192), vL 123 (119-126), dL/vL ratio 1.5 (1.53-1.56), maximum H 47 (47-52), dL/maximum H 3.9 (3.7-4.0), S-1 L 86 (89-91), L/W ratio 9.2 (9.0-9.5), S-2 L 72 (75-78), L/W ratio 6.2 (5.7-6.2), distance S-1-2 19 (20-21), L ratio S-1/2 1.19 (1.17-1.19); I-L-6 L 129 (128-130), central H 22 (20-21), L/central H ratio 5.9 (6.1-6.4); L ratio I-L-5/6 1.43 (1.45-1.48).</paragraph>
<paragraph id="4B6036F9EC0C2242AD86FAFCFA9514CB" blockId="13.[491,1478,1401,1701]" pageId="13" pageNumber="177">
Female (
<typeStatus id="9464885BEC0C2242AE27FAFCFD30148C" box="[586,678,1401,1425]" pageId="13" pageNumber="177" type="paratype">paratype</typeStatus>
, n = 1) — Idiosoma L 781, W 578; maximum diameter Dgl 27; coxal field: L 403; Cx-III W 456; Cx-I+II mL 141, lL 267; genital field L/W 182/180, genital plates L 130-136, pregenital sclerite W 81, maximum diameter Ac 1-3: 45, 42, 44.
</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EFA64FA0D1706" blockId="13.[491,1478,1401,1701]" pageId="13" pageNumber="177">Gnathosoma — palp total L 391; dL/H, dL/H ratio: P-1, 37/34, 1.07; P-2, 84/56, 1.5; P-3, 105/46, 2.28; P-4, 125/30, 4.2; P-5, 42/16, 2.7; L ratio P-2/P-4 0.68. Gnathosomal base vL</paragraph>
<paragraph id="4B6036F9EC0C2242AD86F9A0FD6C1720" blockId="13.[491,1478,1401,1701]" box="[491,762,1573,1597]" pageId="13" pageNumber="177">128; chelicera total L 266.</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EF9CCFBFF17B8" blockId="13.[491,1478,1401,1701]" pageId="13" pageNumber="177">Legs — I-L-5 dL 231, vL 148, dL/vL ratio 1.57, maximum H 59, dL/maximum H 3.9, S-1 L 113, L/W ratio 10.6, S-2 L 89, L/W ratio 6.8, distance S-1-2 31, L ratio S-1/2 1.26; I-L-6 L 148, central H 23, L/central H ratio 6.3; L ratio I-L-5/6, 1.56.</paragraph>
<paragraph id="4B6036F9EC0C2242AD86F957FDDA1611" blockId="13.[491,1482,1746,1976]" pageId="13" pageNumber="177">Deutonymph (n = 1) — Idiosoma L 409, W 298; provisionally genital field L/W 63/80, ratio 0.79.</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EF892FC29164C" blockId="13.[491,1482,1746,1976]" pageId="13" pageNumber="177">Palp: total L 219; dL/H, dL/H ratio: P-1, 23/22, 1.1; P-2, 45/34, 1.35; P-3, 56/28, 2.0; P-4, 69/19, 3.7; P-5, 26/9, 2.75; L ratio P-2/P-4 0.66.</paragraph>
<paragraph id="4B6036F9EC0C2242AE7EF8D9FBAE16A5" blockId="13.[491,1482,1746,1976]" pageId="13" pageNumber="177">Legs — I-L-5 dL 144, vL 97, dL/vL ratio 1.49, maximum H 41, dL/maximum H 3.55, S-1 L 69, L/W ratio 7.3, S-2 L 63, L/W ratio 5.7, distance S-1-2 13, L ratio S-1/2 1.1; I-L-6 L 100, central H 19, L/central H ratio 5.3; L ratio I-L-5/6, 1.44.</paragraph>
<paragraph id="4B6036F9EC0C225FAD86F860FAAF1612" blockId="13.[491,1473,2021,2045]" lastBlockId="16.[491,1511,1576,1807]" lastPageId="16" lastPageNumber="180" pageId="13" pageNumber="177">
Discussion — Due to a similar shape of palp and I-L-5 and -6, the new species is closely related to 
<taxonomicName id="8CDF4D7AEC11225FAE38F9ADFCDC175D" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[597,842,1576,1600]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="180" phylum="Arthropoda" rank="species" species="sonkulensis" status="sp. nov.">
<emphasis id="79ABEAEBEC11225FAE38F9ADFCDC175D" box="[597,842,1576,1600]" italics="true" pageId="16" pageNumber="180">Atractides sonkulensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC11225FAF27F9ADFC1E175D" bold="true" box="[842,904,1576,1600]" pageId="16" pageNumber="180">
<taxonomicNameLabel id="62985790EC11225FAF27F9ADFC1E175D" box="[842,904,1576,1600]" pageId="16" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(see below) and differs from all other species in having one seta each on medial and lateral surface of P-3. 
<taxonomicName id="8CDF4D7AEC11225FA863F9CEFA95177E" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[1038,1283,1611,1635]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="180" phylum="Arthropoda" rank="species" species="sonkulensis" status="sp. nov.">
<emphasis id="79ABEAEBEC11225FA863F9CEFA95177E" box="[1038,1283,1611,1635]" italics="true" pageId="16" pageNumber="180">Atractides sonkulensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC11225FA96EF9CEFAD4177E" bold="true" box="[1283,1346,1611,1635]" pageId="16" pageNumber="180">
<taxonomicNameLabel id="62985790EC11225FA96EF9CEFAD4177E" box="[1283,1346,1611,1635]" pageId="16" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs in the male in generally smaller dimensions of idiosoma, I-L-5/6 and palps (e.g., idiosoma L &lt;500 μm, -L-5 dL 145, palp total L &lt;300 μm), a coxal field merged to an unique plate including Vgl-3, a nearly straight ventral margin of P-2 and a shorter S-1/2 interspace (9-10 μm). In the female 
<taxonomicName id="8CDF4D7AEC11225FAE56F950FD4D17F0" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[571,731,1749,1773]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="180" phylum="Arthropoda" rank="species" species="sonkulensis" status="sp. nov.">
<emphasis id="79ABEAEBEC11225FAE56F950FD4D17F0" box="[571,731,1749,1773]" italics="true" pageId="16" pageNumber="180">A. sonkulensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC11225FAEB6F950FC8D17F0" bold="true" box="[731,795,1749,1773]" pageId="16" pageNumber="180">
<taxonomicNameLabel id="62985790EC11225FAEB6F950FC8D17F0" box="[731,795,1749,1773]" pageId="16" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs in a smaller dimensions of medial suture line of Cx-I+II, genital plates and acetabula, and a relatively longer I-L-6 (L ratio I-L-5/6 1.4).
</paragraph>
<caption id="1FA06671EC0F2241AC1AF8C2FDBB166E" ID-DOI="http://doi.org/10.5281/zenodo.4487503" ID-Zenodo-Dep="4487503" httpUri="https://zenodo.org/record/4487503/files/figure.png" pageId="14" pageNumber="178" startId="14.[119,186,1863,1884]" targetBox="[295,1305,306,1804]" targetPageId="14">
<paragraph id="4B6036F9EC0F2241AC1AF8C2FDBB166E" blockId="14.[119,1491,1862,1907]" pageId="14" pageNumber="178">
<emphasis id="79ABEAEBEC0F2241AC1AF8C2FF5D1641" bold="true" box="[119,203,1863,1884]" pageId="14" pageNumber="178">Figure 5</emphasis>
<taxonomicName id="8CDF4D7AEC0F2241ACBBF8C3FE541641" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[214,450,1862,1884]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="178" phylum="Arthropoda" rank="species" species="alaarchaensis" status="sp. nov.">
<emphasis id="79ABEAEBEC0F2241ACBBF8C3FE541641" box="[214,450,1862,1884]" italics="true" pageId="14" pageNumber="178">Atractides alaarchaensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC0F2241ADA5F8C2FD961641" bold="true" box="[456,512,1863,1884]" pageId="14" pageNumber="178">
<taxonomicNameLabel id="62985790EC0F2241ADA5F8C2FD961641" box="[456,512,1863,1884]" pageId="14" pageNumber="178" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype: A – idiosoma, ventral view; B – I-L-5 and -6; C – palp, medial view; D – palp, lateral view; E – genital field. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="1FA06671EC0E2240AC1AF8C2FE9D166E" ID-DOI="http://doi.org/10.5281/zenodo.4487505" ID-Zenodo-Dep="4487505" httpUri="https://zenodo.org/record/4487505/files/figure.png" pageId="15" pageNumber="179" startId="15.[119,186,1863,1884]" targetBox="[250,1342,307,1800]" targetPageId="15">
<paragraph id="4B6036F9EC0E2240AC1AF8C2FE9D166E" blockId="15.[119,1490,1862,1907]" pageId="15" pageNumber="179">
<emphasis id="79ABEAEBEC0E2240AC1AF8C2FF5B1641" bold="true" box="[119,205,1863,1884]" pageId="15" pageNumber="179">Figure 6</emphasis>
<taxonomicName id="8CDF4D7AEC0E2240ACB7F8C3FE5E1641" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[218,456,1862,1884]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="179" phylum="Arthropoda" rank="species" species="alaarchaensis" status="sp. nov.">
<emphasis id="79ABEAEBEC0E2240ACB7F8C3FE5E1641" box="[218,456,1862,1884]" italics="true" pageId="15" pageNumber="179">Atractides alaarchaensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC0E2240ADA3F8C2FD9E1641" bold="true" box="[462,520,1863,1884]" pageId="15" pageNumber="179">
<taxonomicNameLabel id="62985790EC0E2240ADA3F8C2FD9E1641" box="[462,520,1863,1884]" pageId="15" pageNumber="179" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female paratype: A – coxal field; B – I-L-5 and -6; C – palp, lateral view; D – genital field. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="1FA06671EC11225FAE6EFA03FCE914AF" ID-DOI="http://doi.org/10.5281/zenodo.4487507" ID-Zenodo-Dep="4487507" httpUri="https://zenodo.org/record/4487507/files/figure.png" pageId="16" pageNumber="180" startId="16.[515,582,1414,1435]" targetBox="[513,1474,235,1328]" targetPageId="16">
<paragraph id="4B6036F9EC11225FAE6EFA03FCE914AF" blockId="16.[515,1458,1413,1458]" pageId="16" pageNumber="180">
<emphasis id="79ABEAEBEC11225FAE6EFA03FDCF1486" bold="true" box="[515,601,1414,1435]" pageId="16" pageNumber="180">Figure 7</emphasis>
<taxonomicName id="8CDF4D7AEC11225FAE08FA00FCC41486" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[613,850,1413,1435]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="180" phylum="Arthropoda" rank="species" species="alaarchaensis" status="sp. nov.">
<emphasis id="79ABEAEBEC11225FAE08FA00FCC41486" box="[613,850,1413,1435]" italics="true" pageId="16" pageNumber="180">Atractides alaarchaensis</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC11225FAF35FA03FC061486" bold="true" box="[856,912,1414,1435]" pageId="16" pageNumber="180">
<taxonomicNameLabel id="62985790EC11225FAF35FA03FC061486" box="[856,912,1414,1435]" pageId="16" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, deutonymph: A – I-L-5 and -6; B – palp, medial view; C – genital field. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="4B6036F9EC11225FAD86F8B9FAD61649" blockId="16.[491,1344,1852,1876]" box="[491,1344,1852,1876]" pageId="16" pageNumber="180">Etymology — Named after the National Park where the new species was found.</paragraph>
<paragraph id="4B6036F9EC11225FAD86F804FAB21684" blockId="16.[491,1316,1921,1946]" box="[491,1316,1921,1946]" pageId="16" pageNumber="180">
Distribution — 
<collectingCountry id="33C87669EC11225FAEF8F807FC9C1687" box="[661,778,1922,1946]" name="Kyrgyzstan" pageId="16" pageNumber="180">Kyrgyzstan</collectingCountry>
; known only from the 
<emphasis id="79ABEAEBEC11225FA86FF804FB051684" box="[1026,1171,1921,1945]" italics="true" pageId="16" pageNumber="180">locus typicus</emphasis>
(
<figureCitation id="D3E42A7CEC11225FA8F7F804FA8E1684" box="[1178,1304,1921,1945]" captionStart="Figure 10" captionStartId="20.[119,186,1853,1874]" captionTargetBox="[175,1421,220,1800]" captionTargetId="figure-2@20.[175,1421,220,1800]" captionTargetPageId="20" captionText="Figure 10 Photographs of sampling sites: A – KR3, Ala Archa NP, spring outlet (locus typicusof Atractides alaarchaensis n. sp.); B – KR32, stream near Son Kul lake (locus typicusof Atractides sonkulensisn. sp.); C – KR10, spring in Grigorievka canyon (locus typicusof Atractides grigorievkan. sp.); D – KR8, Chon-Kemin NP, upper part of stream near Ashu resort (locus typicusof Torrenticola kyrgyzican. sp.); E – KR15, river on the road to May Saz Pass (locus typicusof Wandesia albertii n. sp.); F – KR22, stream in Karakal region (locus typicusof Atractides manasi n. sp.); G – KR23, rheohelocrenic spring in Karakal region (sampling site of Atractides grigorievkan. sp.); H – KR31, river near Son Kol Lake (sampling site of Atractides sonkulensisn. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4487513" httpUri="https://zenodo.org/record/4487513/files/figure.png" pageId="16" pageNumber="180">Figure 10A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>