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<document id="0975EB58CC6F7BEE91DA2CDF003AEA0D" ID-DOI="10.11646/phytotaxa.345.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13705161" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725584629088" checkinUser="felipe" docAuthor="Guerrero, J. M., A. A. Vouilloud, Sala, S. E., Kociolek, J. P. &amp; Vijver, B. Van De" docDate="2018" docId="BC39245889494974FF719232FC166EC4" docLanguage="en" docName="phytotaxa.345.2.3.pdf" docOrigin="Phytotaxa 345 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.345.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Orthoseira frenguellii Guerrero, Kociolek et Vouilloud 2018, spec. nov." docType="treatment" docVersion="2" lastPageNumber="126" masterDocId="40005C20894E4973FFF99037FFA16B60" masterDocTitle="New species and a new genus of the Orthoseirales from Patagonia, Argentina, with comments on systematic affinities within the Order" masterLastPageNumber="132" masterPageNumber="119" pageNumber="126" updateTime="1725596485087" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="055AF698AB96A117971E90981C4A6BAF">New species and a new genus of the Orthoseirales from Patagonia, Argentina, with comments on systematic affinities within the Order</mods:title>
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<mods:namePart id="BD4DB2DEAAA12907313B5E687E5B7BD3">Guerrero, J. M.</mods:namePart>
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<mods:namePart id="BBFC17B737BCED0FBCACBAAEB5EAD342">A. A. Vouilloud</mods:namePart>
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<mods:namePart id="ADABADFE0897366C2E259D08D803B143">Sala, S. E.</mods:namePart>
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<mods:namePart id="912E4C22E27CB830F99A27AFABBF8D69">Kociolek, J. P.</mods:namePart>
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<mods:title id="188173490D9F294E13C916A8435DAB9E">Phytotaxa</mods:title>
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<treatment id="BC39245889494974FF719232FC166EC4" ID-DOI="http://doi.org/10.5281/zenodo.13704994" ID-Zenodo-Dep="13704994" LSID="urn:lsid:plazi:treatment:BC39245889494974FF719232FC166EC4" httpUri="http://treatment.plazi.org/id/BC39245889494974FF719232FC166EC4" lastPageNumber="126" pageId="7" pageNumber="126">
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<taxonomicName id="F390EECD89494974FF719232FCAA6940" authority="Guerrero, Kociolek et Vouilloud" authorityName="Guerrero, Kociolek et Vouilloud" authorityYear="2018" box="[136,779,517,544]" class="Bacillariophyceae" family="Orthoseiraceae" genus="Orthoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Orthoseirales" pageId="7" pageNumber="126" phylum="Ochrophyta" rank="species" species="frenguellii" status="sp. nov.">
<emphasis id="06E4495C89494974FF719232FE246940" bold="true" box="[136,389,517,544]" italics="true" pageId="7" pageNumber="126">Orthoseira frenguellii</emphasis>
Guerrero, Kociolek et Vouilloud
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(
<figureCitation id="ACAB89CB89494974FC6A9230FBB26940" box="[915,1043,519,544]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 5471</figureCitation>
)
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<subSubSection id="7C8AC6C589494974FF71921CFE7868E8" pageId="7" pageNumber="126" type="description">
<paragraph id="342F954E89494974FF71921CFE7868E8" blockId="7.[136,1452,555,1444]" pageId="7" pageNumber="126">
<emphasis id="06E4495C89494974FF71921CFEB36924" bold="true" box="[136,274,555,580]" pageId="7" pageNumber="126">Description</emphasis>
:—Valves rounded, showing an irregular, marginal ring of small spines (
<figureCitation id="ACAB89CB89494974FB93921CFAF56924" box="[1130,1364,555,580]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 59, 60, 63, 65</figureCitation>
). Valve dimensions (n=26): diameter 23.550.5 μm. Striae on the valve face of varying length, branching near the mantle,
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10 μm, composed of dash-like areolae,
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10 μm (
<figureCitation id="ACAB89CB89494974FC6F9244FC4C69EC" box="[918,1005,627,652]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Fig. 60</figureCitation>
). Central area hyaline with 2, rarely 3 carinoportulae (
<figureCitation id="ACAB89CB89494974FEC592A0FE6269D0" box="[316,451,663,688]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 5462</figureCitation>
). Small spine-like structures scattered on the valve face among the striae and onto the central area, clearly visible in LM (
<figureCitation id="ACAB89CB89494974FDDA928CFD7969B4" box="[547,728,699,724]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 55, 58, 61</figureCitation>
). Mantle striae parallel to the pervalvar axis,
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10 μm, composed of distinct areolae decreasing in size to the mantle edge,
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10 μm; interstriae on the valve mantle thickened (
<figureCitation id="ACAB89CB89494974FEFC9334FE29687C" box="[261,392,771,796]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 67, 71</figureCitation>
). Internally, areolar foramina occluded by small siliceous ingrowths extending into the lumen (
<figureCitation id="ACAB89CB89494974FF699310FF476820" box="[144,230,807,832]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Fig. 70</figureCitation>
). Carinoportula openings nearly round, partially occluded by a lobed silica structure (
<figureCitation id="ACAB89CB89494974FB499310FAA26820" box="[1200,1283,807,832]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Fig. 69</figureCitation>
, arrow). Small slits irregularly packed between the carinoportulae (
<figureCitation id="ACAB89CB89494974FD30937CFCBA6804" box="[713,795,843,868]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Fig. 69</figureCitation>
, arrowhead). Girdle with numerous copulae, each one with 12 rows of poroids (
<figureCitation id="ACAB89CB89494974FE819358FE6A68E8" box="[376,459,879,904]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Fig. 71</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7C8AC6C589494974FF4493A4FB1668B0" pageId="7" pageNumber="126" type="materials_examined">
<paragraph id="342F954E89494974FF4493A4FB1668B0" blockId="7.[136,1452,555,1444]" pageId="7" pageNumber="126">
<emphasis id="06E4495C89494974FF4493A4FF5968CC" bold="true" box="[189,248,915,940]" pageId="7" pageNumber="126">Type</emphasis>
:—
<collectingCountry id="4C87D5DE89494974FEE193A4FE6368CC" box="[280,450,915,940]" name="Argentina" pageId="7" pageNumber="126">ARGENTINA</collectingCountry>
, Small creek near Copahue Volcano,
<collectingRegion id="F6545BAC89494974FC7493A4FC5468CC" box="[909,1013,915,940]" country="Argentina" name="Neuquen" pageId="7" pageNumber="126">Neuquén</collectingRegion>
, Collector J. Frenguelli,
<date id="402EB38E89494974FADC93A4FA0868CC" box="[1317,1449,915,940]" pageId="7" pageNumber="126" value="1932-05-02">02/05/1932</date>
. (
<typeStatus id="EB2B2BEC89494974FF699380FF5568B0" box="[144,244,951,976]" pageId="7" pageNumber="126" type="holotype">holotype</typeStatus>
Frenguelli Collection, Series 375, Slide 2, finder J 29, here illustrated as
<figureCitation id="ACAB89CB89494974FBD09380FB0D68B0" box="[1065,1196,951,976]" captionStart="FIGURES 5471" captionStartId="9.[136,243,1634,1655]" captionTargetBox="[191,1406,191,1606]" captionTargetId="figure-14@9.[177,1411,187,1611]" captionTargetPageId="9" captionText="FIGURES 5471. Orthoseira frenguellii spec. nov. Specimens of type population.5464. LM. 6571. SEM. 5462. Valve views showing population variability. 5455. Holotype. 6364.Valves in girdle view. 6567. External valve views. 65. Whole frustule in oblique view. 66. Detail of valve center showing external openings of carinoportulae (arrow). 67. Detail of valve in girdle view; note the thickened interstriae. 6870. Internal valve views. 68. Whole valve. 69. Detail of valve center; note the internal opening of the carinoportula (arrow) and packed slits (arrowhead). 70. Detail of areolae showing the siliceous ingrowths on the foramina.71. Valve in girdle view; note copulae with rows of poroids. Scale bars = 10 μm (Figs 5465, 68, 71), 5 μm (Fig. 67), 2 μm (Fig. 66); 1 μm (Figs 6970). Bar length is the same in all LM figures." figureDoi="http://doi.org/10.5281/zenodo.13705173" httpUri="https://zenodo.org/record/13705173/files/figure.png" pageId="7" pageNumber="126">Figs 5455</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7C8AC6C589494974FF4493ECFD2E6F78" pageId="7" pageNumber="126" type="etymology">
<paragraph id="342F954E89494974FF4493ECFD2E6F78" blockId="7.[136,1452,555,1444]" pageId="7" pageNumber="126">
<emphasis id="06E4495C89494974FF4493ECFE9F6894" bold="true" box="[189,318,987,1012]" pageId="7" pageNumber="126">Etymology</emphasis>
:—named in honor of Dr. Joaquín Frenguelli for his many contributions to science in
<collectingCountry id="4C87D5DE89494974FACC93ECFA066894" box="[1333,1447,987,1012]" name="Argentina" pageId="7" pageNumber="126">Argentina</collectingCountry>
, especially with regard to his work on diatoms.
</paragraph>
</subSubSection>
<subSubSection id="7C8AC6C589494974FF449414FC166EC4" pageId="7" pageNumber="126" type="discussion">
<paragraph id="342F954E89494974FF449414FD136E58" blockId="7.[136,1452,555,1444]" pageId="7" pageNumber="126">
<emphasis id="06E4495C89494974FF449414FE8B6F5C" bold="true" box="[189,298,1059,1084]" pageId="7" pageNumber="126">Remarks</emphasis>
:—in a few specimens of the studied population, stria and areola densities on the valve face are beyond the range given in the description (see Table 1). This species differs in LM from
<taxonomicName id="F390EECD89494974FBC19470FB416F00" authorityName="Guerrero &amp; A.A.Vouilloud &amp; Sala &amp; Kociolek &amp; Vijver" authorityYear="2018" box="[1080,1248,1095,1120]" class="Bacillariophyceae" family="Orthoseiraceae" genus="Orthoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Orthoseirales" pageId="7" pageNumber="126" phylum="Ochrophyta" rank="species" species="ferrarioana">
<emphasis id="06E4495C89494974FBC19470FB416F00" box="[1080,1248,1095,1120]" italics="true" pageId="7" pageNumber="126">O. ferrarioana</emphasis>
</taxonomicName>
in the number of carinoportulae; stria and areola densities, both on the valve face and on the mantle. In SEM further differences are evident: presence of small ingrowths in the internal areolae openings; morphology of the internal occlusion of the carinoportulae and number and arrangement of the central slits. Neither this species nor
<taxonomicName id="F390EECD89494974FB959484FAB36FAC" authorityName="Guerrero &amp; A.A.Vouilloud &amp; Sala &amp; Kociolek &amp; Vijver" authorityYear="2018" box="[1132,1298,1203,1228]" class="Bacillariophyceae" family="Orthoseiraceae" genus="Orthoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Orthoseirales" pageId="7" pageNumber="126" phylum="Ochrophyta" rank="species" species="ferrarioana">
<emphasis id="06E4495C89494974FB959484FAB36FAC" box="[1132,1298,1203,1228]" italics="true" pageId="7" pageNumber="126">O. ferrarioana</emphasis>
</taxonomicName>
have internal valves or caverns. The structure of the slits placed between the carinoportulae is similar to that described in other species of the genus
<taxonomicName id="F390EECD89494974FE8D94CCFE4E6E74" box="[372,495,1275,1300]" class="Bacillariophyceae" family="Orthoseiraceae" genus="Orthoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Orthoseirales" pageId="7" pageNumber="126" phylum="Ochrophyta" rank="genus">
<emphasis id="06E4495C89494974FE8D94CCFE4E6E74" box="[372,495,1275,1300]" italics="true" pageId="7" pageNumber="126">Orthoseira</emphasis>
</taxonomicName>
(
<bibRefCitation id="5001E8BF89494974FE0794CCFD0F6E74" author="Crawford, R. M." box="[510,686,1275,1300]" pageId="7" pageNumber="126" pagination="174 - 192" refId="ref7612" refString="Crawford, R. M. (1981) The diatom genus Aulacoseira Thwaites: its structure and taxonomy. Phycologia 20: 174 - 192. https: // doi. org / 10.2216 / i 0031 - 8884 - 20 - 2 - 174.1" type="journal article" year="1981">Crawford 1981</bibRefCitation>
,
<bibRefCitation id="5001E8BF89494974FD4294CCFC5D6E74" author="Spaulding, S. A. &amp; Kociolek, J. P." box="[699,1020,1275,1300]" pageId="7" pageNumber="126" pagination="133 - 147" refId="ref8769" refString="Spaulding, S. A. &amp; Kociolek, J. P. (1998) The diatom genus Orthoseira: ultrastructure and morphological variation in two species from Madagascar with comments on nomenclature in the genus. Diatom Research 13: 133 - 147. https: // doi. org / 10.1080 / 0269249 X. 1998.9705439" type="journal article" year="1998">Spaulding &amp; Kociolek 1998</bibRefCitation>
,
<bibRefCitation id="5001E8BF89494974FBF194CCFAD96E74" author="Van de Vijver, B. &amp; Kopalova, K." box="[1032,1400,1275,1300]" pageId="7" pageNumber="126" pagination="105 - 118" refId="ref9029" refString="Van de Vijver, B. &amp; Kopalova, K. (2008) Orthoseira gremmenii, a new aerophilic diatom from, Gough Island (southern Atlantic Ocean). Cryptogamie, Algologie 29: 105 - 118. https: // doi. org / 10.7872 / crya. v 35. iss 3.2014.245" type="journal article" year="2008">Van de Vijver &amp; Kopalová 2008</bibRefCitation>
) but they clearly differ in their arrangement in groups.
</paragraph>
<paragraph id="342F954E89494974FF449574FC166EC4" blockId="7.[136,1452,555,1444]" pageId="7" pageNumber="126">
This species differs from other
<taxonomicName id="F390EECD89494974FDE59574FD4E6E3C" box="[540,751,1347,1372]" pageId="7" pageNumber="126">
<emphasis id="06E4495C89494974FDE59574FD366E3C" box="[540,663,1347,1372]" italics="true" pageId="7" pageNumber="126">Orthoseira</emphasis>
species
</taxonomicName>
in having a particular combination of characters such as valve face striae branching near the mantle; internal areola foramina with small siliceous ingrowths, carinoportula openings partially occluded by a lobed silica structure and slits irregularly packed.
</paragraph>
</subSubSection>
</treatment>
</document>