256 lines
34 KiB
XML
256 lines
34 KiB
XML
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<mods:title id="416C48980C9AD086AB2DCE7E2718043B">Akarbatrus gen. nov., an unusual Batrisitae (Coleoptera: Staphylinidae: Pselaphinae) from Sumatra</mods:title>
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<mods:namePart id="C16A153EA2CD59775BEB84F8DE9001F0">Löbl, Ivan</mods:namePart>
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<mods:title id="B31682441C8381E4166585E4BDC1215E">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="5D747BDA562DAE6E72C6ED032BF218A1">
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<mods:date id="E94690BCC18106F0AEFEB54F3F625F85">2009</mods:date>
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<mods:number id="44B63C7E6F03A5D7440A31340E332157">2009-12-15</mods:number>
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<mods:number id="0EAB1CB147D255207A3AE38173D0D7E4">49</mods:number>
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<mods:identifier id="6F08A38D1BA1E054B64286A3E59337A0" type="DOI">10.5281/zenodo.5323058</mods:identifier>
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<treatment id="BC03A7484D6FFFBDA5AE4493FCDCDD04" ID-DOI="http://doi.org/10.5281/zenodo.5343545" ID-GBIF-Taxon="185202499" ID-Zenodo-Dep="5343545" LSID="urn:lsid:plazi:treatment:BC03A7484D6FFFBDA5AE4493FCDCDD04" httpUri="http://treatment.plazi.org/id/BC03A7484D6FFFBDA5AE4493FCDCDD04" lastPageId="3" lastPageNumber="664" pageId="1" pageNumber="662">
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<subSubSection id="7CB045D54D6FFFBFA5AE4493FD76DD3B" box="[485,717,754,779]" pageId="1" pageNumber="662" type="nomenclature">
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<paragraph id="3415165E4D6FFFBFA5AE4493FD76DD3B" blockId="1.[485,717,754,779]" box="[485,717,754,779]" pageId="1" pageNumber="662">
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<heading id="6F5DA1324D6FFFBFA5AE4493FD76DD3B" bold="true" box="[485,717,754,779]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="662" reason="3">
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<emphasis id="06DECA4C4D6FFFBFA5AE4493FD76DD3B" bold="true" box="[485,717,754,779]" pageId="1" pageNumber="662">
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<taxonomicName id="F3AA6DDD4D6FFFBFA5AE4493FDDADD3B" ID-CoL="8XJF3" authority="Löbl, 2009" authorityName="Löbl" authorityYear="2009" box="[485,609,754,779]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="662" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="06DECA4C4D6FFFBFA5AE4493FDDADD3B" bold="true" box="[485,609,754,779]" italics="true" pageId="1" pageNumber="662">Akarbatrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1DED77374D6FFFBFA6224492FD76DD3B" box="[617,717,755,779]" pageId="1" pageNumber="662" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CB045D54D6FFFBFA6634570FD30DD14" box="[552,651,785,804]" pageId="1" pageNumber="662" type="description">
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<paragraph id="3415165E4D6FFFBFA6634570FD30DD14" blockId="1.[552,651,785,804]" box="[552,651,785,804]" pageId="1" pageNumber="662">
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(
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<figureCitation id="AC910ADB4D6FFFBFA6664570FD3FDD14" box="[557,644,785,804]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[152,191,1213,1232]" captionStartId-1="4.[151,190,1515,1534]" captionTargetBox-0="[151,1126,219,1184]" captionTargetBox-1="[169,1126,220,1398]" captionTargetId-0="figure-101@2.[151,1126,219,1184]" captionTargetId-1="figure-10@4.[151,1126,204,1420]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="Figs. 1–4. Akarbatrus jelineki gen. nov.et sp. nov., SEM micrographs. 1–2 – head (1 – dorsal view; 2 – lateral view); 3–4 – pronotum (3 – dorsal view; 4 – lateral view)." captionText-1="Figs. 5–10. Akarbatrus jelineki gen. nov. et sp. nov. 5 – antennomeres 1 to 4, scale bar = 0.1 mm; 6 – pronotum in ventral view, scale bar = 0.2 mm; 7 – mesotrochanter, scale bar = 0.1 mm; 8 – metatrochanter, scale bar = 0.1 mm; 9–10 – aedeagus (9 – ventral view, 10 – lateral view; scale bar = 0.1 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.5323060" figureDoi-1="http://doi.org/10.5281/zenodo.5323062" httpUri-0="https://zenodo.org/record/5323060/files/figure.png" httpUri-1="https://zenodo.org/record/5323062/files/figure.png" pageId="1" pageNumber="662">Figs 1–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="7CB045D54D6FFFBFA43A455AFE62DD7E" box="[113,473,826,846]" pageId="1" pageNumber="662" type="type_taxon">
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<paragraph id="3415165E4D6FFFBFA43A455AFE62DD7E" blockId="1.[113,473,826,846]" box="[113,473,826,846]" pageId="1" pageNumber="662">
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<emphasis id="06DECA4C4D6FFFBFA43A455AFF53DD7E" bold="true" box="[113,232,826,846]" pageId="1" pageNumber="662">
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<typeStatus id="EB11A8FC4D6FFFBFA43A455AFF25DD7E" box="[113,158,827,846]" pageId="1" pageNumber="662">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="F3AA6DDD4D6FFFBFA4A6455DFE2ADD7E" ID-CoL="5TV5M" authority="Löbl, 2009" authorityName="Löbl" authorityYear="2009" box="[237,401,828,846]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="662" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
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<emphasis id="06DECA4C4D6FFFBFA4A6455DFE2ADD7E" box="[237,401,828,846]" italics="true" pageId="1" pageNumber="662">Akarbatrus jelineki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1DED77374D6FFFBFA5DD455AFE62DD7E" box="[406,473,827,846]" pageId="1" pageNumber="662" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CB045D54D6FFFBFA43A4507FDB1DA6E" pageId="1" pageNumber="662" type="diagnosis">
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<paragraph id="3415165E4D6FFFBFA43A4507FDB1DA6E" blockId="1.[113,1089,870,1566]" pageId="1" pageNumber="662">
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<emphasis id="06DECA4C4D6FFFBFA43A4507FF5DDD4E" bold="true" box="[113,230,870,894]" pageId="1" pageNumber="662">Diagnosis.</emphasis>
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Head rounded triangular. Eyes situated in posterior half of head. Antenna long, club loosely 3-segmented, ultimate segment conspicuously long. Pronotum lacking tubercles, spines and antebasal foveae, with antebasal stria and pair of laterobasal foveae. Elytra lacking foveae and dorsal striae. Marginal elytral striae present. Abdomen moderately narrowed apically, with rounded apex. Abdominal tergite 1 slightly longer than tergites 2 or 3, with lateral margins rounded. Legs long. Male pronotum excavated.Aedeagus elongate, with loose, weakly attached associated sclerite and long paramere, median lobe moderately flattened, with dorsal membrane and apophyses.
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</paragraph>
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</subSubSection>
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<subSubSection id="7CB045D54D6FFFBFA43A4207FE75D82E" pageId="1" pageNumber="662" type="description">
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<paragraph id="3415165E4D6FFFBFA43A4207FE75D82E" blockId="1.[113,1089,870,1566]" pageId="1" pageNumber="662">
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<emphasis id="06DECA4C4D6FFFBFA43A4207FF45DA4E" bold="true" box="[113,254,1126,1150]" pageId="1" pageNumber="662">Description.</emphasis>
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Length
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<quantity id="F352BBBB4D6FFFBFA5154207FDBBDA4E" box="[350,512,1126,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.75" metricValueMin="1.35" pageId="1" pageNumber="662" unit="mm" value="1.55" valueMax="1.75" valueMin="1.35">1.35–1.75 mm</quantity>
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. Head rounded triangular. U-shaped vertexal sulcus present or absent. Vertexal foveae small. Eyes convex, not emarginate, situated in posterior half of head. Gular carina faint. Gular foveae in a common, fovea-like depression. Maxillary palpus lacking obvious sexual characters, with segment 3 about as long as wide and segment 4 narrowed toward base and apex, with a shallow impression. Antenna 11-segmented, long, with thick scape and loosely 3-segmented club, ultimate antennomere conspicuously elongate. Pronotum with pair of laterobasal foveae, lacking tubercles and spines, with mesal depression. Elytra lacking basal and laterobasal foveae, lacking discal striae, with marginal striae on elytral flanks, with or without lateroapical cleft. Prosternum with pair of simple lateral foveae. Lateral metasternal foveae touching mid-line, about as large as anterolateral metasternal foveae. Abdomen moderately narrowed and rounded apically. Tergite 1 lacking basal sulcus, with pair of basal foveae, angular laterobasally, rounded lateroapically, with only inner marginal carinae apparent. Following tergites lacking marginal carinae. Sternite 2 with two pairs of basal foveae.
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</paragraph>
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</subSubSection>
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<caption id="60D546D64D6CFFBCA4D342DCFDF6DADA" ID-DOI="http://doi.org/10.5281/zenodo.5323060" ID-Zenodo-Dep="5323060" httpUri="https://zenodo.org/record/5323060/files/figure.png" pageId="2" pageNumber="663" startId="2.[152,191,1213,1232]" targetBox="[151,1126,219,1184]" targetPageId="2">
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<paragraph id="3415165E4D6CFFBCA4D342DCFDF6DADA" blockId="2.[152,1128,1213,1259]" pageId="2" pageNumber="663">
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Figs. 1–4.
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<taxonomicName id="F3AA6DDD4D6CFFBCA4BB42DFFE28DAE0" authority="Löbl, 2009" authorityName="Löbl" authorityYear="2009" box="[240,403,1214,1232]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="species" species="jelineki" status="gen. nov.et sp. nov.">
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<emphasis id="06DECA4C4D6CFFBCA4BB42DFFE28DAE0" box="[240,403,1214,1232]" italics="true" pageId="2" pageNumber="663">Akarbatrus jelineki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1DED77374D6CFFBCA5DC42DCFD84DAE0" box="[407,575,1213,1232]" pageId="2" pageNumber="663" rank="species">gen. nov.et sp. nov.</taxonomicNameLabel>
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, SEM micrographs. 1–2 – head (1 – dorsal view; 2 – lateral view); 3–4 – pronotum (3 – dorsal view; 4 – lateral view).
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</paragraph>
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</caption>
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<subSubSection id="7CB045D54D6CFFBDA4DC4342FDD6DC44" lastPageId="3" lastPageNumber="664" pageId="2" pageNumber="663" type="diagnosis">
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<paragraph id="3415165E4D6CFFBCA4DC4342FE3BD82B" blockId="2.[151,1127,1315,1563]" pageId="2" pageNumber="663">
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<emphasis id="06DECA4C4D6CFFBCA4DC4342FE2CDB0B" bold="true" box="[151,407,1315,1339]" pageId="2" pageNumber="663">Differential diagnosis.</emphasis>
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<taxonomicName id="F3AA6DDD4D6CFFBCA5EA4345FDA1DB0B" authority="Löbl, 2009" authorityName="Löbl" authorityYear="2009" box="[417,538,1316,1339]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="06DECA4C4D6CFFBCA5EA4345FDA1DB0B" box="[417,538,1316,1339]" italics="true" pageId="2" pageNumber="663">Akarbatrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1DED77374D6CFFBCA6684342FD32DB0A" box="[547,649,1315,1339]" pageId="2" pageNumber="663" rank="genus">gen. nov.</taxonomicNameLabel>
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may be distinguished from most members of the supertribe
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<taxonomicName id="F3AA6DDD4D6CFFBCA51E4322FE00DB6B" authorityName="Reitter" authorityYear="1882" box="[341,443,1347,1371]" pageId="2" pageNumber="663" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
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by the elytra lacking foveae. The absence of elytral foveae is shared with several members of the temperate edaphic or cavernicolous ‘Amauropini’, and a few subtropical and tropical taxa, such as
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<taxonomicName id="F3AA6DDD4D6CFFBCA63D43E5FC7BDBAB" authority="Chandler, 2001" authorityName="Chandler" authorityYear="2001" box="[630,960,1411,1435]" class="Insecta" family="Staphylinidae" genus="Amblybatrisus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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<emphasis id="06DECA4C4D6CFFBCA63D43E5FCABDBAB" box="[630,784,1412,1435]" italics="true" pageId="2" pageNumber="663">Amblybatrisus</emphasis>
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<bibRefCitation id="503B6BAF4D6CFFBCA75343E2FC7BDBAB" author="CHANDLER D. S." box="[792,960,1411,1435]" pageId="2" pageNumber="663" pagination="1 - 560" refId="ref4201" refString="CHANDLER D. S. 2001: Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International 15: i - viii + 1 - 560." type="journal article" year="2001">Chandler, 2001</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="F3AA6DDD4D6CFFBCA78743E5FE91DB8B" authority="Leleup, 1975" authorityName="Leleup" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Batrisoconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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<emphasis id="06DECA4C4D6CFFBCA78743E5FBDDDBAB" box="[972,1126,1412,1435]" italics="true" pageId="2" pageNumber="663">Batrisoconnus</emphasis>
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||
<bibRefCitation id="503B6BAF4D6CFFBCA4DC43C2FE91DB8B" author="LELEUP N." box="[151,298,1443,1467]" pageId="2" pageNumber="663" pagination="210 - 247" refId="ref4289" refString="LELEUP N. 1975: Contribution a l'etude des coleopteres pselaphides de l'Afrique. 19. Genres et especes termitobies de la region ethiopienne. Bulletin et Annales de la Societe Royale Belge d'Entomologie 111: 210 - 247." type="journal article" year="1975">Leleup, 1975</bibRefCitation>
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||
</taxonomicName>
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,
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||
<taxonomicName id="F3AA6DDD4D6CFFBCA57C43C5FDEADB8B" authority="Raffray, 1894" authorityName="Raffray" authorityYear="1894" box="[311,593,1443,1467]" class="Insecta" family="Staphylinidae" genus="Batrisopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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||
<emphasis id="06DECA4C4D6CFFBCA57C43C5FE14DB8B" box="[311,431,1444,1467]" italics="true" pageId="2" pageNumber="663">Batrisopsis</emphasis>
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||
<bibRefCitation id="503B6BAF4D6CFFBCA5F243C2FDEADB8B" author="RAFFRAY A." box="[441,593,1443,1467]" pageId="2" pageNumber="663" pagination="197 - 282" refId="ref4593" refString="RAFFRAY A. 1894: Revision des Pselaphides des iles de Singapore et de Penang. Revue d'Entomologie 13: 197 - 282, pl. 1." type="journal article" year="1894">Raffray, 1894</bibRefCitation>
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||
</taxonomicName>
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,
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<taxonomicName id="F3AA6DDD4D6CFFBCA61443C5FC2EDB8B" authority="Leleup, 1976" authorityName="Leleup" authorityYear="1976" box="[607,917,1443,1467]" class="Insecta" family="Staphylinidae" genus="Jochmansiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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||
<emphasis id="06DECA4C4D6CFFBCA61443C5FD42DB8B" box="[607,761,1444,1467]" italics="true" pageId="2" pageNumber="663">Jochmansiella</emphasis>
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||
Leleup, 1976
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</taxonomicName>
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,
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<taxonomicName id="F3AA6DDD4D6CFFBCA7E843C5FF74DBEB" authority="Lobl, 1979" authorityName="Lobl" authorityYear="1979" class="Insecta" family="Staphylinidae" genus="Megabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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||
<emphasis id="06DECA4C4D6CFFBCA7E843C5FB9FDB8B" box="[931,1060,1444,1467]" italics="true" pageId="2" pageNumber="663">Megabatrus</emphasis>
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||
<bibRefCitation id="503B6BAF4D6CFFBCA06643C2FF74DBEB" author="LOBL I." pageId="2" pageNumber="663" pagination="189 - 198" refId="ref4493" refString="LOBL I. 1979: Deux Batrisini nouveaux de l'Asie tropicale (Coleoptera, Pselaphidae). Archives des Sciences (Geneve) 32: 189 - 198." type="journal article" year="1979">Löbl, 1979</bibRefCitation>
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||
</taxonomicName>
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||
,
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||
<taxonomicName id="F3AA6DDD4D6CFFBCA49043A5FD81DBEB" authority="Newton & Chandler, 1989" authorityName="Newton & Chandler" authorityYear="1989" box="[219,570,1475,1499]" class="Insecta" family="Staphylinidae" genus="Mnia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
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<emphasis id="06DECA4C4D6CFFBCA49043A5FEA8DBEB" box="[219,275,1476,1499]" italics="true" pageId="2" pageNumber="663">Mnia</emphasis>
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA55043A2FD81DBEB" author="NEWTON A. F. & CHANDLER D. S." box="[283,570,1475,1499]" pageId="2" pageNumber="663" pagination="1 - 92" refId="ref4556" refString="NEWTON A. F. & CHANDLER D. S. 1989: World catalogue of the genera of Pselaphidae (Coleoptera). Fieldiana: Zoology (New Series) 67: 1 - 92." type="journal article" year="1989">Newton & Chandler, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3AA6DDD4D6CFFBCA60E43A5FCE1DBEB" authority="Chandler, 2001" authorityName="Chandler" authorityYear="2001" box="[581,858,1475,1499]" class="Insecta" family="Staphylinidae" genus="Mossman" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6CFFBCA60E43A5FD10DBEB" box="[581,683,1476,1499]" italics="true" pageId="2" pageNumber="663">Mossman</emphasis>
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA6F843A2FCE1DBEB" author="CHANDLER D. S." box="[691,858,1475,1499]" pageId="2" pageNumber="663" pagination="1 - 560" refId="ref4201" refString="CHANDLER D. S. 2001: Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International 15: i - viii + 1 - 560." type="journal article" year="2001">Chandler, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3AA6DDD4D6CFFBCA72E43A5FBD9DBEB" authority="Raffray, 1904" authorityName="Raffray" authorityYear="1904" box="[869,1122,1475,1499]" class="Insecta" family="Staphylinidae" genus="Ophelius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6CFFBCA72E43A5FC7EDBEB" box="[869,965,1476,1499]" italics="true" pageId="2" pageNumber="663">Ophelius</emphasis>
|
||
Raffray, 1904
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="F3AA6DDD4D6CFFBCA48D4385FDB5DBCB" authority="Leleup, 1981" authorityName="Leleup" authorityYear="1981" box="[198,526,1507,1531]" class="Insecta" family="Staphylinidae" genus="Procheilophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="663" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6CFFBCA48D4385FEC3DBCB" box="[198,376,1508,1531]" italics="true" pageId="2" pageNumber="663">Procheilophorus</emphasis>
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA5CB4382FDB5DBCB" author="LELEUP N." box="[384,526,1507,1531]" pageId="2" pageNumber="663" pagination="3 - 76" refId="ref4435" refString="LELEUP N. 1981: Mission entomologique du Dr Ivan Lobl en Cote d'Ivoire. Coleoptera Pselaphidae III (Batrisinae). Revue Suisse de Zoologie 88: 3 - 76." type="journal article" year="1981">Leleup, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA6374382FCBDDBCB" author="LELEUP N." box="[636,774,1507,1531]" pageId="2" pageNumber="663" pagination="210 - 247" refId="ref4289" refString="LELEUP N. 1975: Contribution a l'etude des coleopteres pselaphides de l'Afrique. 19. Genres et especes termitobies de la region ethiopienne. Bulletin et Annales de la Societe Royale Belge d'Entomologie 111: 210 - 247." type="journal article" year="1975">LELEUP 1975</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA7594382FCECDBCB" author="LELEUP N." box="[786,855,1507,1531]" pageId="2" pageNumber="663" pagination="301 - 319" refId="ref4377" refString="LELEUP N. 1976 b: Contribution a l'etude des coleopteres pselaphides de Madagascar. 4. Deux genres nouveaux et sept especes inedites decouverts par le Professeur H. Franz (Faronini, Euplectini, Batrisini, Arhytodini, Tyrini). Bulletin et Annales de la Societe Royale Belge d'Entomologie 112: 301 - 319." type="journal article" year="1976">1976b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA7284382FC20DBCB" author="LELEUP N." box="[867,923,1507,1531]" pageId="2" pageNumber="663" pagination="3 - 76" refId="ref4435" refString="LELEUP N. 1981: Mission entomologique du Dr Ivan Lobl en Cote d'Ivoire. Coleoptera Pselaphidae III (Batrisinae). Revue Suisse de Zoologie 88: 3 - 76." type="journal article" year="1981">1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA7E34382FBA5DBCB" author="LOBL I." box="[936,1054,1507,1531]" pageId="2" pageNumber="663" pagination="151 - 206" refId="ref4467" refString="LOBL I. 1973: Revision der Gattung Mina Raffray (Coleoptera, Pselaphidae). Revue Suisse de Zoologie 108: 151 - 206." type="journal article" year="1973">LÖBL 1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA0614382FBD9DBCB" author="LOBL I." box="[1066,1122,1507,1531]" pageId="2" pageNumber="663" pagination="189 - 198" refId="ref4493" refString="LOBL I. 1979: Deux Batrisini nouveaux de l'Asie tropicale (Coleoptera, Pselaphidae). Archives des Sciences (Geneve) 32: 189 - 198." type="journal article" year="1979">1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA4DC4062FE8BD82B" author="RAFFRAY A." box="[151,304,1539,1563]" pageId="2" pageNumber="663" pagination="197 - 282" refId="ref4593" refString="RAFFRAY A. 1894: Revision des Pselaphides des iles de Singapore et de Penang. Revue d'Entomologie 13: 197 - 282, pl. 1." type="journal article" year="1894">RAFFRAY 1894</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6CFFBCA5714062FEC8D82B" author="RAFFRAY A." box="[314,371,1539,1563]" pageId="2" pageNumber="663" refId="ref4621" refString="RAFFRAY A. 1908: Coleoptera. Fam. Pselaphidae. In: WYTSMANN P. (ed.): Genera Insectorum. 64 th fascicule. P. Wytsmann, Bruxelles, 487 pp., 9 pls." type="book" year="1908">1908</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3415165E4D6DFFBDA4DB46BCFDD6DC44" blockId="3.[113,1089,220,820]" pageId="3" pageNumber="664">
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA4DB46BCFEBDDEC4" authorityName="Löbl" authorityYear="2009" box="[144,262,221,244]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA4DB46BCFEBDDEC4" box="[144,262,221,244]" italics="true" pageId="3" pageNumber="664">Akarbatrus</emphasis>
|
||
</taxonomicName>
|
||
is unique within the
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA59C46BDFD80DEC4" authorityName="Reitter" authorityYear="1882" box="[471,571,220,244]" pageId="3" pageNumber="664" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
|
||
in having both reduced elytral foveae and sexually modified male pronotum. Thus, the combination of these two features provides means for a reliable identification of the new genus.
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA669477CFD21DF04" authorityName="Löbl" authorityYear="2009" box="[546,666,285,308]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA669477CFD21DF04" box="[546,666,285,308]" italics="true" pageId="3" pageNumber="664">Akarbatrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA69B477CFC8DDF04" authorityName="Chandler" authorityYear="2001" box="[720,822,285,308]" class="Insecta" family="Staphylinidae" genus="Mossman" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA69B477CFC8DDF04" box="[720,822,285,308]" italics="true" pageId="3" pageNumber="664">Mossman</emphasis>
|
||
</taxonomicName>
|
||
are similar in their body shape and share long appendages, pronota without spines, presence of laterobasal pronotal foveae while other pronotal foveae are absent, abdominal tergites 1 lacking basal sulci and possessing only one pair of basal foveae and one pair of inner marginal carinae, and elongate and complex aedeagus with a long paramere.
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA61D47FCFD75DF84" authorityName="Löbl" authorityYear="2009" box="[598,718,413,436]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA61D47FCFD75DF84" box="[598,718,413,436]" italics="true" pageId="3" pageNumber="664">Akarbatrus</emphasis>
|
||
</taxonomicName>
|
||
may be readily distinguished from
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA43A47DCFF6DDFE4" authorityName="Chandler" authorityYear="2001" box="[113,214,445,468]" class="Insecta" family="Staphylinidae" genus="Mossman" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA43A47DCFF6DDFE4" box="[113,214,445,468]" italics="true" pageId="3" pageNumber="664">Mossman</emphasis>
|
||
</taxonomicName>
|
||
also by the apically moderately narrowed and rounded abdomen, while the abdomen is conical in
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA4BD47BCFEE7DFC4" authorityName="Chandler" authorityYear="2001" box="[246,348,477,500]" class="Insecta" family="Staphylinidae" genus="Mossman" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA4BD47BCFEE7DFC4" box="[246,348,477,500]" italics="true" pageId="3" pageNumber="664">Mossman</emphasis>
|
||
</taxonomicName>
|
||
. In addition,
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA5A247BCFDDADFC4" authorityName="Löbl" authorityYear="2009" box="[489,609,477,500]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA5A247BCFDDADFC4" box="[489,609,477,500]" italics="true" pageId="3" pageNumber="664">Akarbatrus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA6A047BCFCEADFC4" authorityName="Chandler" authorityYear="2001" box="[747,849,477,500]" class="Insecta" family="Staphylinidae" genus="Mossman" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA6A047BCFCEADFC4" box="[747,849,477,500]" italics="true" pageId="3" pageNumber="664">Mossman</emphasis>
|
||
</taxonomicName>
|
||
by the eyes situated in the basal half of the head and the pronotum lacking tubercles. The remaining genera listed above are not linked by any synapomorphy and each of them except members of the ‘Amauropini’ appears to be isolated within the supertribe
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA6D7445DFCB9DC64" authorityName="Reitter" authorityYear="1882" box="[668,770,572,596]" pageId="3" pageNumber="664" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA775445DFC51DC64" author="CHANDLER D. S." box="[830,1002,572,596]" pageId="3" pageNumber="664" pagination="1 - 560" refId="ref4201" refString="CHANDLER D. S. 2001: Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International 15: i - viii + 1 - 560." type="journal article" year="2001">CHANDLER 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA7BD445DFF0CDC43" author="LELEUP N." pageId="3" pageNumber="664" pagination="301 - 319" refId="ref4377" refString="LELEUP N. 1976 b: Contribution a l'etude des coleopteres pselaphides de Madagascar. 4. Deux genres nouveaux et sept especes inedites decouverts par le Professeur H. Franz (Faronini, Euplectini, Batrisini, Arhytodini, Tyrini). Bulletin et Annales de la Societe Royale Belge d'Entomologie 112: 301 - 319." type="journal article" year="1976">LELEUP 1976b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA489443DFE8DDC44" author="LOBL I." box="[194,310,604,628]" pageId="3" pageNumber="664" pagination="151 - 206" refId="ref4467" refString="LOBL I. 1973: Revision der Gattung Mina Raffray (Coleoptera, Pselaphidae). Revue Suisse de Zoologie 108: 151 - 206." type="journal article" year="1973">LÖBL 1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA50A443DFEC2DC44" author="LOBL I." box="[321,377,604,628]" pageId="3" pageNumber="664" pagination="189 - 198" refId="ref4493" refString="LOBL I. 1979: Deux Batrisini nouveaux de l'Asie tropicale (Coleoptera, Pselaphidae). Archives des Sciences (Geneve) 32: 189 - 198." type="journal article" year="1979">1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA5CF443DFDA6DC44" author="RAFFRAY A." box="[388,541,604,628]" pageId="3" pageNumber="664" pagination="197 - 282" refId="ref4593" refString="RAFFRAY A. 1894: Revision des Pselaphides des iles de Singapore et de Penang. Revue d'Entomologie 13: 197 - 282, pl. 1." type="journal article" year="1894">RAFFRAY 1894</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="503B6BAF4D6DFFBDA66C443DFDDBDC44" author="RAFFRAY A." box="[551,608,604,628]" pageId="3" pageNumber="664" refId="ref4621" refString="RAFFRAY A. 1908: Coleoptera. Fam. Pselaphidae. In: WYTSMANN P. (ed.): Genera Insectorum. 64 th fascicule. P. Wytsmann, Bruxelles, 487 pp., 9 pls." type="book" year="1908">1908</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CB045D54D6DFFBDA43A441DFE81DCE4" pageId="3" pageNumber="664" type="etymology">
|
||
<paragraph id="3415165E4D6DFFBDA43A441DFE81DCE4" blockId="3.[113,1089,220,820]" pageId="3" pageNumber="664">
|
||
<emphasis id="06DECA4C4D6DFFBDA43A441DFF49DCA4" bold="true" box="[113,242,636,660]" pageId="3" pageNumber="664">Etymology.</emphasis>
|
||
The name is derived from the Indonesian word Akar, meaning root, and combined with an arbitrary abbreviation of
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA64844FCFD5ADC84" authority="Aube, 1833" authorityName="Aube" authorityYear="1833" box="[515,737,668,692]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06DECA4C4D6DFFBDA64844FCFDE7DC84" box="[515,604,669,692]" italics="true" pageId="3" pageNumber="664">Batrisus</emphasis>
|
||
Aubé, 1833
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="EB11A8FC4D6DFFBDA75F44FDFCF9DC84" box="[788,834,668,692]" pageId="3" pageNumber="664">type</typeStatus>
|
||
genus of the
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA79F44FDFB85DC84" authorityName="Reitter" authorityYear="1882" box="[980,1086,668,692]" pageId="3" pageNumber="664" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
|
||
. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CB045D54D6DFFBDA43A44BAFE5BDCC4" box="[113,480,731,756]" pageId="3" pageNumber="664" type="distribution">
|
||
<paragraph id="3415165E4D6DFFBDA43A44BAFE5BDCC4" blockId="3.[113,1089,220,820]" box="[113,480,731,756]" pageId="3" pageNumber="664">
|
||
<emphasis id="06DECA4C4D6DFFBDA43A44BAFEBFDCC3" bold="true" box="[113,260,731,755]" pageId="3" pageNumber="664">Distribution.</emphasis>
|
||
<collectingCountry id="4CBD56CE4D6DFFBDA54144BDFECDDCC4" box="[266,374,732,756]" name="Indonesia" pageId="3" pageNumber="664">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="F66ED8BC4D6DFFBDA5CB44BDFE67DCC4" box="[384,476,732,756]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="664">Sumatra</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CB045D54D6DFFBDA43A449AFCDCDD04" pageId="3" pageNumber="664" type="discussion">
|
||
<paragraph id="3415165E4D6DFFBDA43A449AFCDCDD04" blockId="3.[113,1089,220,820]" pageId="3" pageNumber="664">
|
||
<emphasis id="06DECA4C4D6DFFBDA43A449AFF48DD23" bold="true" box="[113,243,763,787]" pageId="3" pageNumber="664">Comments.</emphasis>
|
||
All
|
||
<specimenCount id="22ACDDD74D6DFFBDA569449DFE09DD24" box="[290,434,764,788]" pageId="3" pageNumber="664" type="generic">12 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="F3AA6DDD4D6DFFBDA59C449CFDF4DD24" authority="Löbl, 2009" authorityName="Löbl" authorityYear="2009" box="[471,591,765,788]" class="Insecta" family="Staphylinidae" genus="Akarbatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="664" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="06DECA4C4D6DFFBDA59C449CFDF4DD24" box="[471,591,765,788]" italics="true" pageId="3" pageNumber="664">Akarbatrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1DED77374D6DFFBDA61D449DFD03DD23" box="[598,696,764,788]" pageId="3" pageNumber="664" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
collected are males. This is possibly due to differences in phenology (females may appear later than males).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |