258 lines
46 KiB
XML
258 lines
46 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.994.56810" ID-GBIF-Dataset="8639369d-0b63-4725-aa0c-aed78224bbe5" ID-PMC="PMC7686225" ID-Pensoft-Pub="1313-2970-994-1" ID-Pensoft-UUID="11A23FB601685F7ABFCC123875393447" ID-PubMed="33273881" ID-ZooBank="DB1879C919044320BE674D9F863F6775" ModsDocID="1313-2970-994-1" checkinTime="1605693934925" checkinUser="pensoft" docAuthor="Ng, Peter K. L." docDate="2020" docId="9CE1056C862D5E29982255A0E22BFC13" docLanguage="en" docName="ZooKeys 994: 1-34" docOrigin="ZooKeys 994" docSource="http://dx.doi.org/10.3897/zookeys.994.56810" docTitle="Johora thaiana Leelawathanagoon, Lheknim & Ng 2005" docType="treatment" docVersion="4" id="11A23FB601685F7ABFCC123875393447" lastPageNumber="1" masterDocId="11A23FB601685F7ABFCC123875393447" masterDocTitle="Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus" masterLastPageNumber="34" masterPageNumber="1" pageNumber="1" updateTime="1668169916828" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ng, Peter K. L.</mods:namePart>
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<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
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<mods:nameIdentifier type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>994</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.994.56810</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.994.56810</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-994-1</mods:identifier>
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<treatment ID-GBIF-Taxon="169094343" LSID="urn:lsid:plazi:treatment:9CE1056C862D5E29982255A0E22BFC13" httpUri="http://treatment.plazi.org/id/9CE1056C862D5E29982255A0E22BFC13" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="9CE1056C-862D-5E29-9822-55A0E22BFC13" authority="Leelawathanagoon, Lheknim & Ng, 2005" authorityName="Leelawathanagoon, Lheknim & Ng" authorityYear="2005" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora thaiana" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thaiana">Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1E</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2E</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3E</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4E</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5E</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">, 10A-D</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12E</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13E</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Leelawathanagoon, Lheknim & Ng" authorityYear="2005" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora thaiana" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thaiana">Johora thaiana</taxonomicName>
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Leelawathanagoon, Lheknim & Ng, 2005: 60, figs 1, 2;
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<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
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: table.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: 1 male (22.2
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<normalizedToken originalValue="×">x</normalizedToken>
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18.6 mm) (ZRC 2006.0052), Huai Sam Sop, Ko Lok River Basin, Ban Ba La, 140 m above sea level,
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<geoCoordinate degrees="5.71583" direction="north" orientation="latitude" precision="1" value="5.71583">5.71583°N</geoCoordinate>
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,
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<geoCoordinate degrees="101.83917" direction="east" orientation="longitude" precision="1" value="101.83917">101.83917°E</geoCoordinate>
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, Wang District, Narathiwat Province, southern Thailand, coll. Lheknim V, 14 July 1999; 1 female (21.0
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<normalizedToken originalValue="×">x</normalizedToken>
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17.4 mm) (ZRC 2006.0053), Sirindthron Waterfall, Ban Ba La, 300 m above sea level,
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<geoCoordinate degrees="5.8" direction="north" orientation="latitude" precision="5555" value="5.8">5.8°N</geoCoordinate>
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,
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<geoCoordinate degrees="101.82083" direction="east" orientation="longitude" precision="1" value="101.82083">101.82083°E</geoCoordinate>
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, Wang District, southern Thailand, coll. Lheknim V, 15 July 1999.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult carapace width to length ratio 1.10-1.21 (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1E</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12E</figureCitation>
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); dorsal surface gently convex in frontal view, not inflated (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3E</figureCitation>
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); frontal margin almost straight (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
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); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig.
|
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3E</figureCitation>
|
||
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, not joining lateral margin (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
|
||
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
|
||
); anterolateral margin distinctly convex (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
|
||
); posterolateral margin medially concave, distinctly converging towards gently convex, entire posterior carapace margin (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
|
||
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3E</figureCitation>
|
||
); outer surfaces of third maxillipeds with dense, long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4E</figureCitation>
|
||
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12E</figureCitation>
|
||
); G1 subterminal segment gradually tapering from broad proximal part to slender distal part, without distinct shelf-like structure along gently concave outer margin; terminal segment almost straight, ca. half length of subterminal segment, surfaces with numerous short setae (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10A-C</figureCitation>
|
||
); G2 longer than G1, distal segment long, about two-thirds length of basal segment (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10D</figureCitation>
|
||
). Female pleon ovate; somites 3-6 progressively narrower; telson semicircular (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13E</figureCitation>
|
||
). Vulvae large, on anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semicircular (Fig.
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
||
<taxonomicName authorityName="Leelawathanagoon, Lheknim & Ng" authorityYear="2005" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora thaiana" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thaiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Johora thaiana</emphasis>
|
||
</taxonomicName>
|
||
Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
18.6 mm) (ZRC 2006.0052), Thailand
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E-I</emphasis>
|
||
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (34.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
30.0 mm) (ZRC 2020.0072), Pahang
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, E</emphasis>
|
||
left G1 (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, F</emphasis>
|
||
distal part of left G1 (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C, H</emphasis>
|
||
distal part of left G1 (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
distal part of left G1 (mesial view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D, I</emphasis>
|
||
left G2 (ventral view). Scale bas: 1.0 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, D, E, I</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, C, F-H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The G1 structure of
|
||
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles that of
|
||
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
in that the terminal segment is straight; but their subterminal segments differ. In
|
||
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
, the subterminal segment gradually tapers from the broad proximal part to a slender distal section, without a shelf-like structure along the outer margin (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10A</figureCitation>
|
||
); in
|
||
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
, the distal part of the subterminal segment becomes slender more abruptly, resulting in distinct shelf-like structure on the outer margin (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E</figureCitation>
|
||
). Biogeographically, the type localities of
|
||
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
are less than 20 km from the
|
||
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in Jeli, Kelantan; but their carapace features and G1s differ markedly (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10A-C</figureCitation>
|
||
versus Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Johora erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternum and pleon." figureDoi="10.3897/zookeys.994.56810.figure7" httpUri="https://binary.pensoft.net/fig/476318" pageId="0" pageNumber="1">7B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
||
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora erici" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="erici">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Johora erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (38.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
32.0 mm) (ZRC 2020.0360), Perak
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E-H</emphasis>
|
||
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male (41.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
33.8 mm) (ZRC 1995.0269), Kelantan
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, E</emphasis>
|
||
left G1 (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, F</emphasis>
|
||
distal part of left G1 (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C, G</emphasis>
|
||
distal part of left G1 (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D, H</emphasis>
|
||
left G2 (ventral view). Scale bars: 1.0 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, D, E, H</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, C, F, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Leelawathanagoon, Lheknim & Ng" authorityYear="2005" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora thaiana" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thaiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Johora thaiana</emphasis>
|
||
</taxonomicName>
|
||
is known from the forest tributaries in southern Thailand, near the Malaysian border at Kelantan (Fig.
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng & Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim & Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
|
||
). Its distribution is adjacent to that of
|
||
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. which lives in the highlands of northern Malaysia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="conservation">
|
||
<paragraph pageId="0" pageNumber="1">Conservation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The species has so far only been found in protected forests in southern Thailand and is not under any immediate threat;
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. (2009)</bibRefCitation>
|
||
list it as of least concern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |