treatments-xml/data/9C/CD/5B/9CCD5BDE97C15F7E9439578AB6C80B85.xml

<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="9CCD5BDE97C15F7E9439578AB6C80B85" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) jordanensis Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="A95FB1A6-43F7-4FFC-A202-05073CC72F90" docUuidSource="ZooBank" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wrase, David W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Machard, Patrice</mods:namePart>
<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:date>2022</mods:date>
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18. 
<taxonomicName LSID="http://zoobank.org/A95FB1A6-43F7-4FFC-A202-05073CC72F90" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) jordanensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4C</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7K</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9F</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11E</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12L</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15E</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
Jordan, 
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
Governorate, 
<normalizedToken originalValue="Al-Betrā’">Al-Betra'</normalizedToken>
District, Little Petra, archaeological area, 30.3667, 35.4333.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'JORDAN (
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
) / Little Petra, ca. 1000 m / archaeological area / 
<geoCoordinate degrees="30" direction="north" minutes="22" orientation="latitude" precision="925" value="30.366667">30°22'N</geoCoordinate>
/ 
<geoCoordinate degrees="35" direction="east" minutes="26" orientation="longitude" precision="925" value="35.433334">35°26'E</geoCoordinate>
/ (narrow shaft beside / stony staircase) / 25.III.2016 Wrase &amp; Laser [12B] [w, p]' (cWR). Paratypes: 1♀, 'Jordanien 1024 m / Prov 
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
/ W. Ziegler 9.3.2015 [w, p] // N Wadi Musa / 30°24'.15&quot;N, 35°26'.53&quot;E [w, p]' (cZIEG); 1♂, 'Jordania / Prov. 
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
/ n Wadi Musa 1023m / W.Ziegler 25.3.2016 [w, p] // Jabal al Bayda (Project / 30°24'291&quot;N, 35°26'743&quot;E [w, p]' (NMNHS); 1♂ 'S-Jordan: Wadi Rum / Qatar Spring / 
<geoCoordinate degrees="29.51" direction="north" orientation="latitude" precision="555" value="29.51">29.51°N</geoCoordinate>
/ 
<geoCoordinate degrees="35.41" direction="east" orientation="longitude" precision="555" value="35.41">35.41°E</geoCoordinate>
/ 1000-1100 m asl [w, p] // dry wadi with 
<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees, 21.III.2017 / leg. Th. Assmann [w, p]&quot; (cAL); 1♂, 
<normalizedToken originalValue="ʻS-Jordan">ʻS-Jordan</normalizedToken>
: Little Petra, ca. 1000m / wadis with Mediterranean trees in vicinity to the archaeological site / 
<geoCoordinate degrees="30.34" direction="north" orientation="latitude" precision="555" value="30.34">30.34°N</geoCoordinate>
, 
<geoCoordinate degrees="035.46" direction="east" orientation="longitude" precision="555" value="35.46">035.46°E</geoCoordinate>
/ 25.III.2016 / leg. Th. Assmann [w, p]' (cAL).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 5 specimens. TGE: 4♂♂, 1♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, based on the name of the country in which this species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
Differs from all other species of the &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the following set of characters: (1) meso- and metatarsomeres dorsally convex and smooth (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D</figureCitation>
); (2) posterior angles of pronotum incompletely rounded, with tips protruding laterally (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
); and (3) ventral margin of apex of median lobe (lateral view) evidently convex thus apex appears bent up (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus jordanensis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
share a trait that can be a mark for their close relationships, the apical gonocoxite without nematiform setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.70-9.90 mm; BW: 3.00-3.45 mm), with rather elongate and moderately convex body (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly light reddish-brown, palpomeres and legs somewhat lighter than rest of body. Integument slightly to moderately shiny, head and pronotum shinier than elytra. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Surface of pronotum with evident microreticulation, sculpticells regular isodiametric. Elytra (intervals, scutellum, basal margin and lateral gutter) with distinct isodiametric sculpticells. Ventral surface largely with more or less impressed isodiametric or slight transverse sculpticells. Head impunctate, nearly smooth, with a few, fine and shallow wrinkles in and laterally of frontal furrows. Pronotum surface mostly smooth, only basal area between foveae longitudinally wrinkled and adjacent lateral areas with rare and superficial punctures that do not reach anterior half. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled medially, smooth laterally, other ventrites neither wrinkled nor punctate. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third or more as wide as pronotum wide (PW/HW= 1.35-1.42). Eyes long, moderately convex. Labrum subrectangular, shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows small, punctiform. Paraorbital sulci straight, backward surpassing posterior margin of eye, hardly reaching level of posterior supraorbital pore. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a tenth wider than long (PW/PL= 1.08-1.13), with widest point at medial third. Anterior transverse impression absent; posterior transverse impression barely distinct. Sides sinuate, convex medially and anteriorly, slightly concave posteriorly; anterior bead present laterally, lacking or present in medial 1/10; basal bead present laterally, reduced to absent in medial third or medial half. Metepisternum somewhat longer than wide, MA/MI= 0.88-0.91. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times or little more as long as elytra width (EL/EW= 1.56-1.60), two-and-a-half-times as long as pronotum (EL/PL= 2.41-2.60), and a bit less than a half as wide as pronotum (EW/PW= 1.39-1.46), with widest point at medial third. Parascutellar striole linear, as impressed as other striae, not joining stria 1; striae 1-8 and striole slightly to indistinctly punctate; base of stria 1 ending in parascutellar pore, striae 2-5 reaching or barely not reaching basal bead, 6 and 7 ending little before basal bead. Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 punctures on each elytron in three specimens examined, including holotype. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemora with 6 (holotype) or 5 (both paratypes) setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX subtriangular, with proximal margin symmetrical and pointed at apex (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7K</figureCitation>
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, shaft broad with proximal part slightly constricted, and short, straight apex (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
); median lobe in ventral view straight, 3.7 times longer than wide; apical lamella (dorsal view) rather short, nearly symmetrical, rounded at tip, with sides straight to slightly concave. Internal sac in lateral view (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
) with ventral sclerite slightly broadened and rounded distally; same in ventral view (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11E</figureCitation>
), with dorsal sclerite large, having small left-sided and large right-sided protuberance, and ventral sclerite straight and rather narrow. Right paramere as on Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12L</figureCitation>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15E</figureCitation>
). Apical gonocoxite with widely rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
In additions to characters mentioned in 
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. differs from geographically close 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferranei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferranei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferranei</emphasis>
</taxonomicName>
by pronotum less wide (PW/PL: 1.08-1.13, vs. PW/PL: 1.14-1.21), with anterior margin narrower than posterior margin (PA/PB: 0.78-0.87, vs. PA/PB: 0.88-0.92).
</paragraph>
<paragraph pageId="0" pageNumber="71">
For differences between 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
, from one side, and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
, from other side, see sections 
<normalizedToken originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
(under the first two species) and 
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
(under 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
In Little Petra 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanansis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanansis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanansis</emphasis>
</taxonomicName>
sp. nov. lives in wadis that have a canyon-like profile and some Mediterranean tree species. There it lives together with 
<taxonomicName class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus quadricollisquadricollis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="quadricollisquadricollis">
<emphasis italics="true" pageId="0" pageNumber="71">Laemostenus quadricollisquadricollis</emphasis>
</taxonomicName>
L. Redtenbacher, 1843, 
<taxonomicName authorityName="Piochard de la Brulerie" authorityYear="1876" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus crucifer" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="crucifer">
<emphasis italics="true" pageId="0" pageNumber="71">Trechus crucifer</emphasis>
</taxonomicName>
Piochard de la 
<normalizedToken originalValue="Brûlerie">Brulerie</normalizedToken>
, 1876, 
<taxonomicName authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" lsidName="Calathus cinctus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis italics="true" pageId="0" pageNumber="71">Calathus cinctus</emphasis>
</taxonomicName>
Motschulsky, 1850, 
<taxonomicName authorityName="Menetries" authorityYear="1832" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis andreae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="andreae">
<emphasis italics="true" pageId="0" pageNumber="71">Cymindis andreae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
, 1832, and 
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus impressus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="impressus">
<emphasis italics="true" pageId="0" pageNumber="71">Carabus impressus</emphasis>
</taxonomicName>
Klug, 1832. In Wadi Rum the single specimen was found in a dry wadi with 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" lsidName="Retama" order="Fabales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Retama</emphasis>
</taxonomicName>
shrubs and single 
<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees in the shade of the walls of the head of the canyon-like valley. It occurs together with ground beetles that are mainly typical for desert habitats (
<taxonomicName authorityName="Schatzmayr" authorityYear="1936" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis italics="true" pageId="0" pageNumber="71">Laemostenus aegyptiacus</emphasis>
</taxonomicName>
Schatzmayr, 1936, 
<taxonomicName authorityName="Peyerimhoff" authorityYear="1907" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" lsidName="Singilis filicornis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis italics="true" pageId="0" pageNumber="71">Singilis filicornis</emphasis>
</taxonomicName>
Peyerimhoff, 1907, 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis hierichontica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="hierichontica">
<emphasis italics="true" pageId="0" pageNumber="71">Cymindis hierichontica</emphasis>
</taxonomicName>
Reiche &amp; Saulcy, 1855, 
<taxonomicName baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" lsidName="Merizomena castanea" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="71">Merizomena castanea</emphasis>
</taxonomicName>
Klug, 1832, and 
<taxonomicName authorityName="Bedel" authorityYear="1907" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" lsidName="Amara maindroni" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="0" pageNumber="71">Amara maindroni</emphasis>
</taxonomicName>
Bedel, 1907). A photograph of the habitat of the latter is given by 
<bibRefCitation DOI="https://doi.org/10.4081/fe.2017.226" author="Casale, A" journalOrPublisher="Fragmenta Entomologica" pageId="0" pageNumber="71" pagination="13 - 24" refId="B14" refString="Casale, A, Assmann, T, 2017. The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini). Fragmenta Entomologica 49 (1): 13 - 24, DOI: https://doi.org/10.4081/fe.2017.226" title="The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini)." url="https://doi.org/10.4081/fe.2017.226" volume="49" year="2017">Casale and Assmann (2017</bibRefCitation>
: 22, fig. 35).
</paragraph>
<caption doi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 16.</emphasis>
Distribution of species from subgenus 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in the western Aegean Sea Region and Turkey: 
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus chatzakiae</emphasis>
</taxonomicName>
sp. nov. (red circle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
(blue circles), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. (brown circles), and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
</taxonomicName>
sp. nov. (yellow circle).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
According to the data available, the species is the representative of the group &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; in the southern part of Jordan, stretching from the vicinity of Little Petra to areas close to the Saudi Arabian border (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). As Wadi Rum is in the most southern part of Jordan, the species may also occur in Northwest Saudi Arabia.
</paragraph>
</subSubSection>
</treatment>
</document>