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<document id="10FD4B5BB33C5B9218BC8FC809ED87F4" ID-CLB-Dataset="6318" ID-DOI="10.11646/zootaxa.4970.2.1" ID-GBIF-Dataset="89d3c795-198b-4162-8903-9c6cf3f1e665" ID-ISSN="1175-5326" ID-Zenodo-Dep="4761544" ID-ZooBank="6DBA0C02-9E52-4FD0-8B48-8C7B405713A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620984897660" checkinUser="plazi" docAuthor="Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="052487CFFFD4E50346C6B50B8C62FF0E" docLanguage="en" docName="zootaxa.4970.2.1.pdf" docOrigin="Zootaxa 4970 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gangugia mula Bouzan &amp; Iniesta &amp; Brescovit 2021, n. sp." docType="treatment" docUuid="C71B69A8-40C3-4E39-8050-6ECA65AB930D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="234" masterDocId="F91DFFB7FFCAE5224651B2518D76FFF7" masterDocTitle="Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="231" updateTime="1698939836712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="442107C4E57DD83B2583B4B0D2B6277F">
<mods:title id="D7D01BAED54A884988AB8F2E075F466A">Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)</mods:title>
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<mods:name id="3A59D6C515BE35F519A2C64262A810F5" type="personal">
<mods:role id="51A2056BCCB5935971B785F8B3FF63FE">
<mods:roleTerm id="017291F0AA699B35E256633107DA197B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEA0D3FD21CCAC7B2255921BBC134107">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:affiliation id="7FDFD8A21801A6D6B6C9BA60CD170AB3">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil.</mods:affiliation>
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<mods:name id="38C385BBE3A89DCDF29075E6E41B80CC" type="personal">
<mods:role id="D2C951B32DC7184FE2F58145BBE7518F">
<mods:roleTerm id="E765A6EBE77BFAB92317A75DE6D655F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="955CF04110B129A95B057BA4DE1D371B">Iniesta, Luiz Felipe M.</mods:namePart>
<mods:nameIdentifier id="CB08BA7F7D9CAC3222AF090D87828FC7" type="ORCID">0000-0002-0529-4162</mods:nameIdentifier>
<mods:affiliation id="C6384CA921B3943A1C20B5B80C022C51">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. &amp; luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162</mods:affiliation>
<mods:nameIdentifier id="29C7977FA8523C2B5B9EBDBDEF2C96C6" type="email">luiz-moretti@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="4028629E4C5E79229FAED1BAD40FF1F6" type="personal">
<mods:role id="E17E2C13F181395C6E09980E42184203">
<mods:roleTerm id="FAFD45607AD6A68DE60F09CF093DA56D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4A6572C3C274C46FC42E6F99A3BADCC">Brescovit, Antonio Domingos</mods:namePart>
<mods:nameIdentifier id="89A99B242C13F2BCDDC8C758C4E8FBA0" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="E6EAC9DC2EAED6796E12A031D3C57CF7">antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="8A89F0946983A5D02C1DF0A9E94C32D9" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:typeOfResource id="8E632149130C35244FED296CD8C72DA0">text</mods:typeOfResource>
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<mods:title id="BE91F374E7D28497B96EA7A31C4A2108">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D2297F9CACA1DCD23406CAB21A2B8187">
<mods:date id="7CF8101D4CCE34B85AB57B813DF0FACB">2021</mods:date>
<mods:detail id="D828F3D47E76C2EC8AF076274FD46996" type="pubDate">
<mods:number id="14122AE65CB69F8C229C6FB8C766F018">2021-05-14</mods:number>
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<mods:number id="0AC4E86A6FC114353A67DE9CCE711619">4970</mods:number>
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<mods:identifier id="B5764623CF6D4AB20326953C4123ABB2" type="DOI">10.11646/zootaxa.4970.2.1</mods:identifier>
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<treatment id="052487CFFFD4E50346C6B50B8C62FF0E" ID-DOI="http://doi.org/10.5281/zenodo.4761613" ID-GBIF-Taxon="182486290" ID-Zenodo-Dep="4761613" LSID="urn:lsid:zoobank.org:act:C71B69A8-40C3-4E39-8050-6ECA65AB930D" httpUri="http://treatment.plazi.org/id/052487CFFFD4E50346C6B50B8C62FF0E" lastPageId="33" lastPageNumber="234" pageId="30" pageNumber="231">
<subSubSection id="C5976552FFD4E53C46C6B50B8CE9F883" box="[151,415,1882,1909]" pageId="30" pageNumber="231" type="nomenclature">
<paragraph id="8D3236D9FFD4E53C46C6B50B8CE9F883" blockId="30.[151,972,1882,1981]" box="[151,415,1882,1909]" pageId="30" pageNumber="231">
<heading id="D67A81B5FFD4E53C46C6B50B8CE9F883" bold="true" box="[151,415,1882,1909]" fontSize="11" level="1" pageId="30" pageNumber="231" reason="1">
<emphasis id="BFF9EACBFFD4E53C46C6B50B8CE9F883" bold="true" box="[151,415,1882,1909]" pageId="30" pageNumber="231">
<taxonomicName id="4A8D4D5AFFD4E53C46C6B50B8C23F882" ID-CoL="8H7SG" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[151,341,1882,1909]" class="Diplopoda" family="Chelodesmidae" genus="Gangugia" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="mula" status="sp. nov.">
<emphasis id="BFF9EACBFFD4E53C46C6B50B8C23F882" bold="true" box="[151,341,1882,1909]" italics="true" pageId="30" pageNumber="231">Gangugia mula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4CA57B0FFD4E53C470DB50B8CE9F883" box="[348,415,1882,1908]" pageId="30" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD4E53C46C6B52F8EBAF84A" pageId="30" pageNumber="231" type="description">
<paragraph id="8D3236D9FFD4E53C46C6B52F8CE1F86F" blockId="30.[151,972,1882,1981]" box="[151,407,1918,1944]" pageId="30" pageNumber="231">
<figureCitation id="15B62A5CFFD4E53C46C6B52F8C4DF86F" box="[151,315,1918,1944]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="31.[151,250,1939,1963]" captionStartId-1="32.[151,250,2002,2026]" captionTargetBox-0="[161,1426,417,1916]" captionTargetBox-1="[166,1421,849,1979]" captionTargetId-0="figure-120@31.[161,1426,417,1916]" captionTargetId-1="figure-319@32.[166,1421,849,1979]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." captionText-1="FIGURE 18. Distribution map of Gangugia Schubart, 1947 species." figureDoi-0="http://doi.org/10.5281/zenodo.4761609" figureDoi-1="http://doi.org/10.5281/zenodo.4761611" httpUri-0="https://zenodo.org/record/4761609/files/figure.png" httpUri-1="https://zenodo.org/record/4761611/files/figure.png" pageId="30" pageNumber="231">Figures 17–18</figureCitation>
,
<figureCitation id="15B62A5CFFD4E53C4717B52F8CE1F86F" box="[326,407,1918,1944]" captionStart="FIGURE 24" captionStartId="41.[151,250,1835,1859]" captionTargetBox="[303,1284,181,1811]" captionTargetId="figure-20@41.[303,1284,181,1811]" captionTargetPageId="41" captionText="FIGURE 24. Left vulva. Gangugia tapirapensis Schubart, 1947 (paralectotype, MZSP 1070): (A), ventral view; (B), lateral view. Gangugia mula n. sp. (paratype, IBSP 276): (C), ventral view; (D), lateral view. Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3569): (E), ventral view; (F), lateral view. Abiliodesmus cataractae (Hoffman, 1976) comb. nov. (paratype, MZSP 1066). (G), ventral view; (H), lateral view. Scale bars: 0,5 mm.Abbreviations: EV, external valve; IV, internal valve; Op, operculum." figureDoi="http://doi.org/10.5281/zenodo.4761631" httpUri="https://zenodo.org/record/4761631/files/figure.png" pageId="30" pageNumber="231">24C–D</figureCitation>
</paragraph>
<paragraph id="8D3236D9FFD4E53C46C6B5F38EBAF84A" blockId="30.[151,972,1882,1981]" box="[151,972,1954,1981]" pageId="30" pageNumber="231">
<uri id="F91C3ADBFFD4E53C46C6B5F38EBAF84A" box="[151,972,1954,1981]" pageId="30" pageNumber="231">
urn:lsid:zoobank.org:act:
<uuid id="F92B0C0CFFD4E53C47FDB5F38EBAF84A" box="[428,972,1954,1981]" pageId="30" pageNumber="231">C71B69A8-40C3-4E39-8050-6ECA65AB930D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD5E53D46C6B2C68FCCFF0E" pageId="31" pageNumber="232" type="materials_examined">
<paragraph id="8D3236D9FFD5E53D46C6B2C68FCCFF0E" blockId="31.[151,1437,150,393]" pageId="31" pageNumber="232">
<emphasis id="BFF9EACBFFD5E53D46C6B2C68C32FF47" bold="true" box="[151,324,150,177]" pageId="31" pageNumber="232">Type material.</emphasis>
<materialsCitation id="3DE53C84FFD5E53D4718B2C68FA1FF22" ID-GBIF-Occurrence="3125622354" collectionCode="IBSP" collectorName="Instituto Butantan Expedition" country="Brazil" county="Tucurui" latitude="-3.7002778" location="Male" longLatPrecision="21" longitude="-49.7" municipality="Tucurui" pageId="31" pageNumber="232" specimenCode="IBSP 277" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<location id="88526002FFD5E53D4718B2C68CF5FF46" LSID="urn:lsid:plazi:treatment:052487CFFFD4E50346C6B50B8C62FF0E:88526002FFD5E53D4718B2C68CF5FF46" box="[329,387,151,177]" country="Brazil" county="Tucurui" latitude="-3.7002778" longLatPrecision="21" longitude="-49.7" municipality="Tucurui" name="Male" pageId="31" pageNumber="232" stateProvince="Para">Male</location>
<typeStatus id="5236887BFFD5E53D47D9B2C68C9CFF46" box="[392,490,151,177]" pageId="31" pageNumber="232" type="holotype">holotype</typeStatus>
from UHE
<collectingMunicipality id="6D56ACA3FFD5E53D443BB2C68FB3FF46" box="[618,709,151,177]" pageId="31" pageNumber="232">Tucuruí</collectingMunicipality>
,
<collectingCounty id="64534E55FFD5E53D449FB2C68E50FF46" box="[718,806,151,177]" pageId="31" pageNumber="232">Tucuruí</collectingCounty>
(
<geoCoordinate id="E8B9501EFFD5E53D4565B2C68ECDFF46" box="[820,955,151,177]" degrees="03" direction="south" minutes="42" orientation="latitude" pageId="31" pageNumber="232" precision="15" seconds="01" value="-3.7002778">03°42’01”S</geoCoordinate>
,
<geoCoordinate id="E8B9501EFFD5E53D4597B2C6892FFF46" box="[966,1113,151,177]" degrees="49" direction="west" minutes="42" orientation="longitude" pageId="31" pageNumber="232" precision="15" seconds="00" value="-49.7">49°42’00”W</geoCoordinate>
),
<collectingRegion id="4F49F83BFFD5E53D423AB2C689D6FF46" box="[1131,1184,151,177]" country="Brazil" name="Para" pageId="31" pageNumber="232">Pará</collectingRegion>
,
<collectingCountry id="F59A7649FFD5E53D42F8B2C68986FF46" box="[1193,1264,151,177]" name="Brazil" pageId="31" pageNumber="232">Brazil</collectingCountry>
, 1984,
<collectorName id="2078530FFFD5E53D436EB2C68CF4FF22" pageId="31" pageNumber="232">Instituto Butantan Expedition</collectorName>
coll., deposited in
<specimenCode id="DD2B9EA2FFD5E53D4434B2EA8FA1FF22" box="[613,727,187,213]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="232">IBSP 277</specimenCode>
</materialsCitation>
.
<materialsCitation id="3DE53C84FFD5E53D44B4B2EA8CEEFF0E" ID-GBIF-Occurrence="3125622347" collectionCode="IBSP" collectorName="Instituto Butantan Expedition" country="Brazil" county="Tucurui" latitude="-3.7002778" location="Male" longLatPrecision="21" longitude="-49.7" municipality="Tucurui" pageId="31" pageNumber="232" specimenCode="IBSP 274" specimenCount="1" specimenCount-female="1" stateProvince="Para" typeStatus="paratype">
<typeStatus id="5236887BFFD5E53D44B4B2EA8E3DFF22" box="[741,843,187,213]" pageId="31" pageNumber="232" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9B8BFD50FFD5E53D450BB2EA8E85FF22" box="[858,1011,187,213]" pageId="31" pageNumber="232" type="female">three females</specimenCount>
with the same data of the
<typeStatus id="5236887BFFD5E53D4362B2EA88EEFF22" box="[1331,1432,187,213]" pageId="31" pageNumber="232" type="holotype">holotype</typeStatus>
, deposited in
<specimenCode id="DD2B9EA2FFD5E53D4777B28E8CE2FF0E" box="[294,404,223,249]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="232">IBSP 274</specimenCode>
,
</materialsCitation>
<materialsCitation id="3DE53C84FFD5E53D47CEB28E8F7BFF0E" ID-GBIF-Occurrence="3125622310" box="[415,525,223,249]" collectionCode="IBSP" collectorName="Instituto Butantan Expedition" country="Brazil" county="Tucurui" isTailEnumeration="true" latitude="-3.7002778" location="Male" longLatPrecision="21" longitude="-49.7" municipality="Tucurui" pageId="31" pageNumber="232" specimenCode="IBSP 276" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode id="DD2B9EA2FFD5E53D47CEB28E8F7BFF0E" box="[415,525,223,249]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="232">IBSP 276</specimenCode>
</materialsCitation>
,
<materialsCitation id="3DE53C84FFD5E53D4449B28E8FC0FF0E" ID-GBIF-Occurrence="3125622301" box="[536,694,223,249]" collectionCode="IBSP" country="Brazil" pageId="31" pageNumber="232" specimenCode="IBSP 278" specimenCount="1" stateProvince="Para" typeStatus="paratype">
and
<specimenCode id="DD2B9EA2FFD5E53D4419B28E8FC0FF0E" box="[584,694,223,249]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="232">IBSP 278</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD5E53D4696B3538C87FE7E" pageId="31" pageNumber="232" type="diagnosis">
<paragraph id="8D3236D9FFD5E53D4696B3538C87FE7E" blockId="31.[151,1437,150,393]" pageId="31" pageNumber="232">
<emphasis id="BFF9EACBFFD5E53D4696B3538C34FEEB" bold="true" box="[199,322,258,284]" pageId="31" pageNumber="232">Diagnosis.</emphasis>
Adult males differ from all other species of the genus by the combination of the following characters: legs from the fifth to the eighth segment with ventral projections in the coxae (
<figureCitation id="15B62A5CFFD5E53D4274B37689FBFEB6" box="[1061,1165,295,321]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); C–D) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); E–F) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="31" pageNumber="232">Fig. 31B</figureCitation>
); presence of triangular lateral expansion near the apex of the solenomere (
<figureCitation id="15B62A5CFFD5E53D44BCB31A8E27FE92" box="[749,849,331,357]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="31" pageNumber="232">Fig. 17F</figureCitation>
) and presence of a short secondary process in the prefemoral process (
<figureCitation id="15B62A5CFFD5E53D472DB33E8C93FE7E" box="[380,485,367,393]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="31" pageNumber="232">Fig. 17D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D9F26651FFD5E53D46C6B5C28E8AF804" ID-DOI="http://doi.org/10.5281/zenodo.4761609" ID-Zenodo-Dep="4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="31" pageNumber="232" startId="31.[151,250,1939,1963]" targetBox="[161,1426,417,1916]" targetPageId="31">
<paragraph id="8D3236D9FFD5E53D46C6B5C28E8AF804" blockId="31.[151,1437,1939,2035]" pageId="31" pageNumber="232">
<emphasis id="BFF9EACBFFD5E53D46C6B5C28C57F85C" bold="true" box="[151,289,1939,1963]" pageId="31" pageNumber="232">FIGURE 17.</emphasis>
<taxonomicName id="4A8D4D5AFFD5E53D4778B5C58CBFF85C" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[297,457,1939,1963]" class="Diplopoda" family="Chelodesmidae" genus="Gangugia" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="mula" status="sp. nov.">
<emphasis id="BFF9EACBFFD5E53D4778B5C58CBFF85C" box="[297,457,1939,1963]" italics="true" pageId="31" pageNumber="232">Gangugia mula</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFD5E53D4781B5C28F7DF85C" bold="true" box="[464,523,1939,1963]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="A4CA57B0FFD5E53D4781B5C28F7DF85C" box="[464,523,1939,1963]" pageId="31" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process.
</paragraph>
</caption>
<subSubSection id="C5976552FFEAE5024696B2C78CD5FCE2" pageId="32" pageNumber="233" type="description">
<paragraph id="8D3236D9FFEAE5024696B2C7885DFD96" blockId="32.[151,1437,150,825]" pageId="32" pageNumber="233">
<emphasis id="BFF9EACBFFEAE5024696B2C78C2FFF47" bold="true" box="[199,345,150,176]" pageId="32" pageNumber="233">Description.</emphasis>
Male (
<typeStatus id="5236887BFFEAE50247E0B2C68F68FF46" box="[433,542,151,177]" pageId="32" pageNumber="233" type="holotype">Holotype</typeStatus>
, IBSP 277). Coloration (long-preserved in 70% ethanol): Head light brown. Antennae ocher yellowish. Body brown, with the edge of the paranota lighter (
<figureCitation id="15B62A5CFFEAE5024258B2EA89E5FF22" box="[1033,1171,187,213]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17A–C</figureCitation>
). Legs ocher yellowish. Legs from the fifth to the eighth segment with ventral projections in the coxae being the posterior pair of the sixth segment extremely elongated. Telson ocher. Total length: 65.51. Total width: 11.50. Collum, length 3.44, width 10.82. Antennomere lengths (1&gt;7): 0.95; 2.38; 2.14; 1.88; 1.79; 1.70; 0.35. Podomeres lengths (1&gt;7): 1.19; 1.69; 2.99; 1.42; 1.86; 1.90; 0.45. Gonopod aperture, length 2.46, width 4.87. Telson, length 1.41. Gonopod: length 3.42, width 3.07. Gonocoxae: length 1.51, width 1.36. Gonocoxae globose, about 1/2 the size of the telopodite. Telopodite: length 3.33, width 1.62. Prefemoral region short (
<figureCitation id="15B62A5CFFEAE5024516B3C28ED9FE5A" box="[839,943,403,429]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17D</figureCitation>
). Prefemoral process (
<figureCitation id="15B62A5CFFEAE50242F8B3C2885BFE5A" box="[1193,1325,403,429]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17D–F</figureCitation>
) long and curve, presenting a dorso-basal lobe, starting from an angle of 225°, involving the solenomere in its initial portion and subsequently ascending parallel to the solenomere and partially covering it in ectal view (
<figureCitation id="15B62A5CFFEAE50242B3B38A883FFE02" box="[1250,1353,475,501]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17F</figureCitation>
); in its anterior portion presents an elongated, thin and acuminate secondary process (
<figureCitation id="15B62A5CFFEAE5024256B3AE8919FDEE" box="[1031,1135,511,537]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17D</figureCitation>
). Solenomere long, curved and with a lateral expansion (
<figureCitation id="15B62A5CFFEAE50247BDB0728F27FDCA" box="[492,593,547,573]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Fig. 17F</figureCitation>
), but not reaching the apex of the prefemoral process. Cingulum in basal position, covered by the prefemoral process. Apex of the solenomere abruptly descending (
<figureCitation id="15B62A5CFFEAE50242C3B0168869FD96" box="[1170,1311,583,609]" captionStart="FIGURE 17" captionStartId="31.[151,250,1939,1963]" captionTargetBox="[161,1426,417,1916]" captionTargetId="figure-120@31.[161,1426,417,1916]" captionTargetPageId="31" captionText="FIGURE 17. Gangugia mula n. sp. (holotype, IBSP 277), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761609" httpUri="https://zenodo.org/record/4761609/files/figure.png" pageId="32" pageNumber="233">Figs 17D–E</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFEAE5024696B03A8CD5FCE2" blockId="32.[151,1437,150,825]" pageId="32" pageNumber="233">
Female (IBSP 276). Body as in male, except for the legs; without modifications. Total length: 75.74. Total width: 11.46. Antennomere lengths (1&gt;7): 0.74; 2.07; 1.98; 1.70; 1.66; 1.75; 0.35. Podomeres lengths (1&gt;7): 1.10; 1.29;
<geoCoordinate id="E8B9501EFFEAE50246C6B0E28DAFFD3A" box="[151,217,691,717]" degrees="3.048" direction="north" orientation="latitude" pageId="32" pageNumber="233" precision="55" value="3.048">3.048</geoCoordinate>
;
<geoCoordinate id="E8B9501EFFEAE50246B6B0E28C5FFD3A" box="[231,297,691,717]" degrees="1.040" direction="east" orientation="longitude" pageId="32" pageNumber="233" precision="55" value="1.04">1.040</geoCoordinate>
; 1.28; 1.62; 0.32. Telson, length 1.35. Epigyne rounded with irregular edges. Vulvae (
<figureCitation id="15B62A5CFFEAE5024351B0E288E6FD3A" box="[1280,1424,691,717]" captionStart="FIGURE 24" captionStartId="41.[151,250,1835,1859]" captionTargetBox="[303,1284,181,1811]" captionTargetId="figure-20@41.[303,1284,181,1811]" captionTargetPageId="41" captionText="FIGURE 24. Left vulva. Gangugia tapirapensis Schubart, 1947 (paralectotype, MZSP 1070): (A), ventral view; (B), lateral view. Gangugia mula n. sp. (paratype, IBSP 276): (C), ventral view; (D), lateral view. Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3569): (E), ventral view; (F), lateral view. Abiliodesmus cataractae (Hoffman, 1976) comb. nov. (paratype, MZSP 1066). (G), ventral view; (H), lateral view. Scale bars: 0,5 mm.Abbreviations: EV, external valve; IV, internal valve; Op, operculum." figureDoi="http://doi.org/10.5281/zenodo.4761631" httpUri="https://zenodo.org/record/4761631/files/figure.png" pageId="32" pageNumber="233">Figs 24C–D</figureCitation>
): length 2.20, width 0.88. External valve: length 1.17, width 0.39. Internal valve: length 1.17, width 0.43. Operculum: length 0.32, width 0.62.
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFEAE5024696B14F8E2DFCCE" box="[199,859,798,825]" pageId="32" pageNumber="233" type="distribution">
<paragraph id="8D3236D9FFEAE5024696B14F8E2DFCCE" blockId="32.[151,1437,150,825]" box="[199,859,798,825]" pageId="32" pageNumber="233">
<emphasis id="BFF9EACBFFEAE5024696B14F8C17FCCF" bold="true" box="[199,353,798,824]" pageId="32" pageNumber="233">Distribution.</emphasis>
Known only from the
<typeStatus id="5236887BFFEAE5024431B14E8FE6FCCE" box="[608,656,799,825]" pageId="32" pageNumber="233">type</typeStatus>
locality (
<figureCitation id="15B62A5CFFEAE50244AAB14E8E38FCCE" box="[763,846,799,825]" captionStart="FIGURE 18" captionStartId="32.[151,250,2002,2026]" captionTargetBox="[166,1421,849,1979]" captionTargetId="figure-319@32.[166,1421,849,1979]" captionTargetPageId="32" captionText="FIGURE 18. Distribution map of Gangugia Schubart, 1947 species." figureDoi="http://doi.org/10.5281/zenodo.4761611" httpUri="https://zenodo.org/record/4761611/files/figure.png" pageId="32" pageNumber="233">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D9F26651FFEAE50246C6B5838E26F81D" ID-DOI="http://doi.org/10.5281/zenodo.4761611" ID-Zenodo-Dep="4761611" box="[151,848,2002,2026]" httpUri="https://zenodo.org/record/4761611/files/figure.png" pageId="32" pageNumber="233" startId="32.[151,250,2002,2026]" targetBox="[166,1421,849,1979]" targetPageId="32">
<paragraph id="8D3236D9FFEAE50246C6B5838E26F81D" blockId="32.[151,848,2002,2026]" box="[151,848,2002,2026]" pageId="32" pageNumber="233">
<emphasis id="BFF9EACBFFEAE50246C6B5838C56F81D" bold="true" box="[151,288,2002,2026]" pageId="32" pageNumber="233">FIGURE 18.</emphasis>
Distribution map of
<taxonomicName id="4A8D4D5AFFEAE50247A5B5828E26F81D" box="[500,848,2002,2026]" class="Diplopoda" family="Chelodesmidae" genus="Gangugia" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BFF9EACBFFEAE50247A5B5828F2FF81D" box="[500,601,2003,2026]" italics="true" pageId="32" pageNumber="233">Gangugia</emphasis>
<bibRefCitation id="E91C4B28FFEAE5024431B5838F8CF81D" author="Schubart, O." box="[608,762,2002,2026]" pageId="32" pageNumber="233" pagination="1 - 74" refId="ref33135" refString="Schubart, O. (1947) Os Diplopodos da viagem do naturalista Antenor Leitao de Carvalho aos rios Araguaia e Amazonas em 1939 e 1940. Boletim do Museu Nacional, 82, 1 - 74." type="journal article" year="1947">Schubart, 1947</bibRefCitation>
species.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C5976552FFEBE5034696B2C68C62FF0E" pageId="33" pageNumber="234" type="etymology">
<paragraph id="8D3236D9FFEBE5034696B2C68C62FF0E" blockId="33.[151,1436,151,249]" pageId="33" pageNumber="234">
<emphasis id="BFF9EACBFFEBE5034696B2C68C38FF46" bold="true" box="[199,334,151,177]" pageId="33" pageNumber="234">Etymology.</emphasis>
The species epithet is a reference of the Brazilian folkloric character “Mula-sem-cabeça”.According to the legend, of uncertain origin, it’s a mule of brown color with a torch of fire in the place of the head. Noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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