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<document id="F46415AA813F4F12BA2BBD39C15AFDF1" ID-DOI="10.21248/contrib.entomol.65.2.297-325" ID-ISSN="0005-805X" ID-Zenodo-Dep="4753971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620838389347" checkinUser="carolina" docAuthor="Irmler, Ulrich" docDate="2015" docId="050D87D8FFB35F033FCEE116FEC8287F" docLanguage="en" docName="ContribEntomol.65.2.297-325.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 65 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1885" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Glyptoma denieri" docType="treatment" docVersion="3" lastPageNumber="314" masterDocId="F934FFA0FFA35F123F2DE445FFAE2C1E" masterDocTitle="The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="325" masterPageNumber="297" pageNumber="313" updateTime="1698916894542" updateUser="plazi">
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<mods:title id="DDF3438E67F887430EE63F2095619CDD">The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<treatment id="050D87D8FFB35F033FCEE116FEC8287F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F" httpUri="http://treatment.plazi.org/id/050D87D8FFB35F033FCEE116FEC8287F" lastPageId="17" lastPageNumber="314" pageId="16" pageNumber="313">
<subSubSection id="C5BE6545FFB35F023FCEE116FDF12970" box="[227,607,1363,1390]" pageId="16" pageNumber="313" type="nomenclature">
<paragraph id="8D1B36CEFFB35F023FCEE116FDF12970" blockId="16.[227,607,1363,1423]" box="[227,607,1363,1390]" pageId="16" pageNumber="313">
<heading id="D65381A2FFB35F023FCEE116FDF12970" box="[227,607,1363,1390]" centered="true" fontSize="12" level="2" pageId="16" pageNumber="313" reason="2">
<taxonomicName id="4AA44D4DFFB35F023FCEE116FDF12970" ID-CoL="3GHR9" authority="(BERNHAUER, 1939)" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[227,607,1363,1390]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023FCEE116FE3F2970" box="[227,401,1363,1390]" italics="true" pageId="16" pageNumber="313">Glyptoma denieri</emphasis>
(
<bibRefCitation id="E9354B3FFFB35F023EB2E116FDF72970" author="BERNHAUER, M." box="[415,601,1363,1390]" pageId="16" pageNumber="313" pagination="231 - 249" refId="ref22901" refString="BERNHAUER, M. 1939: Zur Staphylinidenfauna Argentiniens und Brasiliens (Col.). (40. Beitrag zur sudamerikanischen Fauna). - Revista de Entomologia 10: 231 - 249." type="journal article" year="1939">BERNHAUER, 1939</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023FCEE13CFE242991" box="[227,394,1401,1423]" pageId="16" pageNumber="313" type="description">
<paragraph id="8D1B36CEFFB35F023FCEE13CFE242991" blockId="16.[227,607,1363,1423]" box="[227,394,1401,1423]" pageId="16" pageNumber="313">
(
<figureCitation id="159F2A4BFFB35F023FCAE13CFEF32991" box="[231,349,1401,1423]" captionStart="Figs 1317" captionStartId="15.[151,191,2027,2049]" captionTargetBox="[197,1366,506,2003]" captionTargetId="figure-143@15.[151,1398,482,2026]" captionTargetPageId="15" captionText="Figs 1317: Aedeagus (A), antenna (B) and microsculpture of head (C) ofGlyptoma nitidum (13), G. obscuricolle (14), G. nitens (15), G. cognatum (16), G. denieri (17); scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4753985" httpUri="https://zenodo.org/record/4753985/files/figure.png" pageId="16" pageNumber="313">Figs 17A B, C</figureCitation>
,
<figureCitation id="159F2A4BFFB35F023E49E13CFE2B2991" box="[356,389,1401,1423]" captionStart="Figs 22" captionStartId="27.[151,191,1934,1956]" captionTargetBox="[164,1386,260,1919]" captionTargetId="figure-41@27.[151,1398,253,1933]" captionTargetPageId="27" captionText="Figs 22: Fore body of Glyptoma nitidum (A), G. obscuricollis (B), G. nitens (C), G. cognatum (D), G. sanctaecatharinae (E), G. klimschi (F); scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753999" httpUri="https://zenodo.org/record/4753999/files/figure.png" pageId="16" pageNumber="313">22E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023F90E1EEFE8C2A1E" pageId="16" pageNumber="313" type="reference_group">
<paragraph id="8D1B36CEFFB35F023F90E1EEFDEC29DE" blockId="16.[189,794,1451,1536]" box="[189,578,1451,1472]" pageId="16" pageNumber="313">
<treatmentCitationGroup id="ADB411E0FFB35F023F90E1EEFDEC29DE" box="[189,578,1451,1472]" pageId="16" pageNumber="313">
<treatmentCitation id="0C0510DFFFB35F023F90E1EEFDEC29DE" author="BERNHAUER, M." box="[189,578,1451,1472]" page="232" pageId="16" pageNumber="313" year="1939">
<taxonomicName id="4AA44D4DFFB35F023F90E1EEFDEC29DE" ID-CoL="PX68" authority="BERNHAUER, 1939: 232" authorityName="BERNHAUER" authorityYear="1939" box="[189,578,1451,1472]" class="Insecta" family="Staphylinidae" genus="Calocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023F90E1EEFECF29DE" box="[189,353,1451,1472]" italics="true" pageId="16" pageNumber="313">Calocerus denieri</emphasis>
<bibRefCitation id="E9354B3FFFB35F023E4BE1EEFDEC29DE" author="BERNHAUER, M." box="[358,578,1451,1472]" pageId="16" pageNumber="313" pagination="231 - 249" refId="ref22901" refString="BERNHAUER, M. 1939: Zur Staphylinidenfauna Argentiniens und Brasiliens (Col.). (40. Beitrag zur sudamerikanischen Fauna). - Revista de Entomologia 10: 231 - 249." type="journal article" year="1939">BERNHAUER, 1939: 232</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8D1B36CEFFB35F023F90E18EFE8C2A1E" blockId="16.[189,794,1451,1536]" pageId="16" pageNumber="313">
<treatmentCitationGroup id="ADB411E0FFB35F023F90E18EFE8C2A1E" pageId="16" pageNumber="313">
<treatmentCitation id="0C0510DFFFB35F023F90E18EFD7129FE" author="SCHEERPELTZ, O." box="[189,735,1483,1504]" page="13" pageId="16" pageNumber="313" year="1970">
<taxonomicName id="4AA44D4DFFB35F023F90E18EFD7129FE" ID-CoL="PX6F" authority="SCHEERPELTZ, 1970: 13" authorityName="SCHEERPELTZ" authorityPageNumber="13" authorityYear="1970" box="[189,735,1483,1504]" class="Insecta" family="Staphylinidae" genus="Calocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="BFD0EADCFFB35F023F90E18EFE7B29FE" box="[189,469,1483,1504]" italics="true" pageId="16" pageNumber="313">Calocerus sanctaecatharinae</emphasis>
<bibRefCitation id="E9354B3FFFB35F023ECAE18EFD7129FE" author="SCHEERPELTZ, O." box="[487,735,1483,1504]" pageId="16" pageNumber="313" pagination="130 - 142" refId="ref23245" refString="SCHEERPELTZ, O. 1970: Studien uber die Arten der GattungCalocerusFauvel (Col. Staphylinidae, Subfam. Piestinae, Tribus Thorscophorini). (19. Beitrag zur Kenntnis der neotropischen Staphyliniden). - Mitteilungen der Munchener Entomologischen Gesellschaft 59: 130 - 142." type="journal article" year="1970">SCHEERPELTZ, 1970: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="BFD0EADCFFB35F023DDCE18EFE8C2A1E" bold="true" pageId="16" pageNumber="313">
<taxonomicNameLabel id="A4E357A7FFB35F023DDCE18EFE8C2A1E" pageId="16" pageNumber="313">new synonymy</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023F90E26CFCAC2A7F" pageId="16" pageNumber="313" type="materials_examined">
<paragraph id="8D1B36CEFFB35F023F90E26CFCAC2A7F" blockId="16.[189,794,1577,1698]" pageId="16" pageNumber="313">
<emphasis id="BFD0EADCFFB35F023F90E26CFE6D2A5F" bold="true" box="[189,451,1577,1601]" pageId="16" pageNumber="313">Type material examined</emphasis>
:
<taxonomicName id="4AA44D4DFFB35F023EE2E26CFD2A2A5F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[463,644,1577,1601]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023EE2E26CFD2A2A5F" box="[463,644,1577,1601]" italics="true" pageId="16" pageNumber="313">Glyptoma denieri</emphasis>
</taxonomicName>
:
<materialsCitation id="3DCC3C93FFB35F023DA2E26CFD502A7F" collectingDate="1933-06-28" collectionCode="FMNH" collectorName="Denier" country="Argentina" location="Salta" pageId="16" pageNumber="313" specimenCount="1" specimenCount-male="1" stateProvince="Salta" typeStatus="holotype">
<specimenCount id="9BA2FD47FFB35F023DA2E26CFD792A5F" box="[655,727,1577,1601]" pageId="16" pageNumber="313" type="male">1 male</specimenCount>
,
<typeStatus id="521F886CFFB35F023DCDE26CFF402A7C" pageId="16" pageNumber="313" type="holotype">holotype</typeStatus>
:
<collectingCountry id="F5B3765EFFB35F023FDAE20CFECB2A7F" box="[247,357,1609,1633]" name="Argentina" pageId="16" pageNumber="313">Argentina</collectingCountry>
:
<collectingRegion id="4F60F82CFFB35F023E42E20CFE0A2A7F" box="[367,420,1609,1633]" country="Argentina" name="Salta" pageId="16" pageNumber="313">Salta</collectingRegion>
,
<date id="F91A100EFFB35F023E83E20CFDBB2A7F" box="[430,533,1609,1633]" pageId="16" pageNumber="313" value="1933-06-28">
<collectingDate id="E95EE9E6FFB35F023E83E20CFDBB2A7F" box="[430,533,1609,1633]" pageId="16" pageNumber="313" value="1933-06-28">28.6.1933</collectingDate>
</date>
, leg.
<collectorName id="20515318FFB35F023D67E20CFD3A2A7F" box="[586,660,1609,1633]" pageId="16" pageNumber="313">Denier</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023D8FE20CFD582A7F" box="[674,758,1609,1633]" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="313" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023F90E22CFCB42ABF" pageId="16" pageNumber="313" type="reference_group">
<paragraph id="8D1B36CEFFB35F023F90E22CFCB42ABF" blockId="16.[189,794,1577,1698]" pageId="16" pageNumber="313">
<treatmentCitationGroup id="ADB411E0FFB35F023F90E22CFCB42ABF" pageId="16" pageNumber="313">
<treatmentCitation id="0C0510DFFFB35F023F90E22CFD632A9F" author="SCHEERPELTZ, O." box="[189,717,1641,1665]" page="2" pageId="16" pageNumber="313" year="1970">
<taxonomicName id="4AA44D4DFFB35F023F90E22CFD632A9F" ID-CoL="PX6F" authority="SCHEERPELTZ, 1970: 2" authorityName="SCHEERPELTZ" authorityPageNumber="2" authorityYear="1970" box="[189,717,1641,1665]" class="Insecta" family="Staphylinidae" genus="Calocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="BFD0EADCFFB35F023F90E22CFE382A9F" box="[189,406,1641,1665]" italics="true" pageId="16" pageNumber="313">C. sanctaecatharinae</emphasis>
<materialsCitation id="3DCC3C93FFB35F023E99E22CFD632A9F" box="[436,717,1641,1665]" collectionCode="NHMW" collectorName="Klimsch" country="Brazil" location="Brazil" pageId="16" pageNumber="313" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
<bibRefCitation id="E9354B3FFFB35F023E99E22CFD632A9F" author="SCHEERPELTZ, O." box="[436,717,1641,1665]" pageId="16" pageNumber="313" pagination="130 - 142" refId="ref23245" refString="SCHEERPELTZ, O. 1970: Studien uber die Arten der GattungCalocerusFauvel (Col. Staphylinidae, Subfam. Piestinae, Tribus Thorscophorini). (19. Beitrag zur Kenntnis der neotropischen Staphyliniden). - Mitteilungen der Munchener Entomologischen Gesellschaft 59: 130 - 142." type="journal article" year="1970">
SCHEERPELTZ, 1970:
<specimenCount id="9BA2FD47FFB35F023DEDE22CFD632A9F" box="[704,717,1641,1665]" pageId="16" pageNumber="313" type="female">2</specimenCount>
</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
female
<typeStatus id="521F886CFFB35F023F90E2CFFEB22ABC" box="[189,284,1674,1698]" pageId="16" pageNumber="313" type="syntype">syntypes</typeStatus>
:
<collectingCountry id="F5B3765EFFB35F023E09E2CCFECB2ABF" box="[292,357,1673,1697]" name="Brazil" pageId="16" pageNumber="313">Brazil</collectingCountry>
: Santa Catharina, leg.
<collectorName id="20515318FFB35F023D67E2CCFD0D2ABF" box="[586,675,1673,1697]" pageId="16" pageNumber="313">Klimsch</collectorName>
(NHMW).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023F96E28CFC052D9F" pageId="16" pageNumber="313" type="materials_examined">
<paragraph id="8D1B36CEFFB35F023F96E28CFC052D9F" blockId="16.[187,795,1737,2081]" lastBlockId="16.[831,1437,233,385]" pageId="16" pageNumber="313">
<emphasis id="BFD0EADCFFB35F023F96E28CFE452AFF" bold="true" box="[187,491,1737,1761]" pageId="16" pageNumber="313">Further material examined</emphasis>
:
<materialsCitation id="3DCC3C93FFB35F023ED0E28CFD622B5F" collectionCode="AMNH" collectorName="J. Carpenter &amp; D. Agosti" country="Argentina" county="Aguas Blancas-Yaculica" elevation="520" latitude="-22.706333" location="Yungas forest" longLatPrecision="12" longitude="-64.392" municipality="Argentina-Bolivian" pageId="16" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Salta">
<collectingCountry id="F5B3765EFFB35F023ED0E28CFDC52AFF" box="[509,619,1737,1761]" name="Argentina" pageId="16" pageNumber="313">Argentina</collectingCountry>
:
<specimenCount id="9BA2FD47FFB35F023D54E28CFD772AFF" box="[633,729,1737,1761]" pageId="16" pageNumber="313" type="female">1 female</specimenCount>
,
<collectingRegion id="4F60F82CFFB35F023DCAE28CFF582B1F" country="Argentina" name="Salta" pageId="16" pageNumber="313">Salta Prov.</collectingRegion>
,
<collectingCounty id="647A4E42FFB35F023FD2E2ACFE522B1F" box="[255,508,1769,1793]" pageId="16" pageNumber="313">Aguas Blancas-Yaculica</collectingCounty>
, (
<collectingMunicipality id="6D7FACB4FFB35F023D20E2ACFD732B1F" box="[525,733,1769,1793]" pageId="16" pageNumber="313">Argentina-Bolivian</collectingMunicipality>
frontier) (
<geoCoordinate id="E8905009FFB35F023E2DE34CFED52B3F" box="[256,379,1801,1825]" degrees="64" direction="west" minutes="23.52" orientation="longitude" pageId="16" pageNumber="313" precision="9" value="-64.392">64°23.52'W</geoCoordinate>
,
<geoCoordinate id="E8905009FFB35F023EA0E34CFE502B3F" box="[397,510,1801,1825]" degrees="22" direction="south" minutes="42.38" orientation="latitude" pageId="16" pageNumber="313" precision="9" value="-22.706333">22°42.38'S</geoCoordinate>
),
<location id="887B6015FFB35F023D35E34FFD012B3F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023D35E34FFD012B3F" box="[536,687,1801,1826]" country="Argentina" county="Aguas Blancas-Yaculica" latitude="-22.706333" longLatPrecision="12" longitude="-64.392" municipality="Argentina-Bolivian" name="Yungas forest" pageId="16" pageNumber="313" stateProvince="Salta">Yungas forest</location>
leaflitter,
<quantity id="4A5C9B2BFFB35F023F90E36CFF512B5F" box="[189,255,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="16" pageNumber="313" unit="m" value="520.0">
<elevation id="0689D1FDFFB35F023F90E36CFF512B5F" box="[189,255,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="16" pageNumber="313" unit="m" value="520.0">520 m</elevation>
</quantity>
elev., leg.
<collectorName id="20515318FFB35F023E48E36FFE4D2B5F" box="[357,483,1833,1857]" pageId="16" pageNumber="313">J. Carpenter</collectorName>
&amp;
<collectorName id="20515318FFB35F023D2DE36FFDCC2B5F" box="[512,610,1833,1857]" pageId="16" pageNumber="313">D. Agosti</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023D43E36CFD682B5F" box="[622,710,1833,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="313">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023DF8E36CFE132B9F" collectionCode="NHMW" collectorName="A. Hetschko" country="Brazil" location="Santa" pageId="16" pageNumber="313" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<collectingCountry id="F5B3765EFFB35F023DF8E36CFCB82B5F" box="[725,790,1833,1857]" name="Brazil" pageId="16" pageNumber="313">Brazil</collectingCountry>
:
<specimenCount id="9BA2FD47FFB35F023F96E30CFEBE2B7F" box="[187,272,1865,1889]" pageId="16" pageNumber="313" type="male">2 males</specimenCount>
,
<specimenCount id="9BA2FD47FFB35F023E36E30CFE2C2B7F" box="[283,386,1865,1889]" pageId="16" pageNumber="313" type="female">4 females</specimenCount>
,
<location id="887B6015FFB35F023EA3E30CFE662B7F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023EA3E30CFE662B7F" box="[398,456,1865,1889]" country="Brazil" name="Santa" pageId="16" pageNumber="313">Santa</location>
<taxonomicName id="4AA44D4DFFB35F023EFDE30CFD472B7F" authority=", Blumeau, 1889" authorityName=", Blumenau" authorityYear="1889" box="[464,745,1865,1889]" class="Polyplacophora" family="Mopaliidae" genus="Catharina" kingdom="Animalia" order="Chitonida" pageId="16" pageNumber="313" phylum="Mollusca" rank="genus">Catharina, Blumeau, 1889</taxonomicName>
, leg.
<collectorName id="20515318FFB35F023F92E32CFEEF2B9F" box="[191,321,1897,1921]" pageId="16" pageNumber="313">A. Hetschko</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023E7BE32CFE1B2B9F" box="[342,437,1897,1921]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="313" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023EE2E32CFDFD2BBF" collectionCode="BMNH" collectorName="Cameron" country="Brazil" location="Sao Paulo" pageId="16" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<specimenCount id="9BA2FD47FFB35F023EE2E32CFD9C2B9F" box="[463,562,1897,1921]" pageId="16" pageNumber="313" type="female">1 female</specimenCount>
,
<collectingRegion id="4F60F82CFFB35F023D69E32CFD1B2B9F" box="[580,693,1897,1921]" country="Brazil" name="Sao Paulo" pageId="16" pageNumber="313">São Paulo</collectingRegion>
, without further data, coll.
<collectorName id="20515318FFB35F023EAEE3CCFE492BBF" box="[387,487,1929,1953]" pageId="16" pageNumber="313">Cameron</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023EDAE3CCFDE22BBF" box="[503,588,1929,1953]" pageId="16" pageNumber="313">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023D4DE3CCFDA92BFF" collectingDate="1960-05-22" collectionCode="UIC" collectorName="H. Schubart &amp; J. Becker" country="Brazil" county="Itatiaia" location="da Serra" municipality="Fazenda da Serra" pageId="16" pageNumber="313" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="9BA2FD47FFB35F023D4DE3CCFD052BBF" box="[608,683,1929,1953]" pageId="16" pageNumber="313" type="male">1 male</specimenCount>
,
<specimenCount id="9BA2FD47FFB35F023D95E3CCFCB82BBF" box="[696,790,1929,1953]" pageId="16" pageNumber="313" type="female">1 female</specimenCount>
,
<collectingRegion id="4F60F82CFFB35F023F90E3ECFEF62BDF" box="[189,344,1961,1985]" country="Brazil" name="Rio de Janeiro" pageId="16" pageNumber="313">Rio de Janeiro</collectingRegion>
,
<collectingCounty id="647A4E42FFB35F023E49E3ECFE1E2BDF" box="[356,432,1961,1985]" pageId="16" pageNumber="313">Itatiaia</collectingCounty>
,
<collectingMunicipality id="6D7FACB4FFB35F023E96E3ECFDD82BDF" box="[443,630,1961,1985]" pageId="16" pageNumber="313">
Fazenda
<location id="887B6015FFB35F023D37E3ECFDD82BDF" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023D37E3ECFDD82BDF" box="[538,630,1961,1985]" country="Brazil" county="Itatiaia" municipality="Fazenda da Serra" name="da Serra" pageId="16" pageNumber="313" stateProvince="Rio de Janeiro">da Serra</location>
</collectingMunicipality>
,
<date id="F91A100EFFB35F023DAFE3ECFD472BDF" box="[642,745,1961,1985]" pageId="16" pageNumber="313" value="1960-05-22">
<collectingDate id="E95EE9E6FFB35F023DAFE3ECFD472BDF" box="[642,745,1961,1985]" pageId="16" pageNumber="313" value="1960-05-22">22.5.1960</collectingDate>
</date>
, leg.
<collectorName id="20515318FFB35F023F92E38FFE932BFF" box="[191,317,1993,2017]" pageId="16" pageNumber="313">H. Schubart</collectorName>
&amp;
<collectorName id="20515318FFB35F023E72E38FFE132BFF" box="[351,445,1993,2017]" pageId="16" pageNumber="313">J. Becker</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023EE1E38CFE512BFF" box="[460,511,1993,2017]" pageId="16" pageNumber="313">UIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023D3EE38CFD42241F" collectingDate="1960-07-17" collectionCode="UIC" collectorName="H. Schubart" country="Brazil" location="Floresta de Tijuca" pageId="16" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro">
<specimenCount id="9BA2FD47FFB35F023D3EE38CFDDE2BFF" box="[531,624,1993,2017]" pageId="16" pageNumber="313" type="female">1 female</specimenCount>
,
<collectingRegion id="4F60F82CFFB35F023D56E38CFCB82BFF" box="[635,790,1993,2017]" country="Brazil" name="Rio de Janeiro" pageId="16" pageNumber="313">Rio de Janeiro</collectingRegion>
,
<location id="887B6015FFB35F023F93E3ACFED3241F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023F93E3ACFED3241F" box="[190,381,2025,2049]" country="Brazil" name="Floresta de Tijuca" pageId="16" pageNumber="313" stateProvince="Rio de Janeiro">Floresta de Tijuca</location>
,
<date id="F91A100EFFB35F023EAAE3ACFE40241F" box="[391,494,2025,2049]" pageId="16" pageNumber="313" value="1960-07-17">
<collectingDate id="E95EE9E6FFB35F023EAAE3ACFE40241F" box="[391,494,2025,2049]" pageId="16" pageNumber="313" value="1960-07-17">17.7.1960</collectingDate>
</date>
, leg.
<collectorName id="20515318FFB35F023D0EE3AFFD0D241F" box="[547,675,2025,2049]" pageId="16" pageNumber="313">H. Schubart</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023D9CE3ACFD4A241F" box="[689,740,2025,2049]" pageId="16" pageNumber="313">UIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023DDBE3ACFCBB243F" collectionCode="BMNH" collectorName="Cameron" country="Brazil" location="Rio de Janeiro" pageId="16" pageNumber="313" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingRegion id="4F60F82CFFB35F023DDBE3ACFE87243F" country="Brazil" name="Rio de Janeiro" pageId="16" pageNumber="313">Rio de Janeiro</collectingRegion>
, without further data, coll.
<collectorName id="20515318FFB35F023D65EC4CFD02243F" box="[584,684,2057,2081]" pageId="16" pageNumber="313">Cameron</collectorName>
(
<collectionCode id="EBB5AE0BFFB35F023D94EC4CFCA0243F" box="[697,782,2057,2081]" pageId="16" pageNumber="313">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023C12E4ACFAF82D1F" box="[831,1366,233,257]" collectionCode="SDEI" collectorName="Kraatz" country="Brazil" location="Brasilia" pageId="16" pageNumber="313" specimenCount="1">
<location id="887B6015FFB35F023C12E4ACFC3F2D1F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023C12E4ACFC3F2D1F" box="[831,913,233,257]" country="Brazil" name="Brasilia" pageId="16" pageNumber="313">Brasilia</location>
, coll.
<collectorName id="20515318FFB35F023CFCE4AFFBB72D1F" box="[977,1049,234,257]" pageId="16" pageNumber="313">Kraatz</collectorName>
, without further data (
<collectionCode id="EBB5AE0BFFB35F023A3FE4ACFAFE2D1F" box="[1298,1360,233,257]" pageId="16" pageNumber="313">SDEI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFB35F023A4EE4AFFC092D9F" accessionNumber="F00105" collectingDate="2000-04-12" collectionCode="KNHM" collectorName="Z. Falin" country="Paraguay" county="Cazaapa" elevation="90" latitude="-26.319166" location="Sosa" longLatPrecision="12" longitude="-55.7425" municipality="Hermosa" pageId="16" pageNumber="313" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F5B3765EFFB35F023A4EE4AFFCDB2D3C" name="Paraguay" pageId="16" pageNumber="313">Paraguay</collectingCountry>
:
<specimenCount id="9BA2FD47FFB35F023CA9E54CFC482D3F" box="[900,998,265,289]" pageId="16" pageNumber="313" type="female">1 female</specimenCount>
,
<collectingCounty id="647A4E42FFB35F023CD8E54CFBFF2D3F" box="[1013,1105,265,289]" pageId="16" pageNumber="313">Cazaapa</collectingCounty>
,
<collectingMunicipality id="6D7FACB4FFB35F023B4CE54FFB6B2D3F" box="[1121,1221,266,289]" pageId="16" pageNumber="313">Hermosa</collectingMunicipality>
, prop.
<location id="887B6015FFB35F023A34E54CFAE72D3F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023A34E54CFAE72D3F" box="[1305,1353,265,289]" country="Paraguay" county="Cazaapa" latitude="-26.319166" longLatPrecision="12" longitude="-55.7425" municipality="Hermosa" name="Sosa" pageId="16" pageNumber="313">Sosa</location>
family,
<location id="887B6015FFB35F023C12E56CFBA72D5F" LSID="urn:lsid:plazi:treatment:050D87D8FFB35F033FCEE116FEC8287F:887B6015FFB35F023C12E56CFBA72D5F" box="[831,1033,297,321]" country="Paraguay" county="Cazaapa" latitude="-26.319166" longLatPrecision="12" longitude="-55.7425" municipality="Hermosa" name="San Rafael Reserve" pageId="16" pageNumber="313">San Rafael Reserve</location>
(
<geoCoordinate id="E8905009FFB35F023B3AE56CFB3C2D5F" box="[1047,1170,297,321]" degrees="55" direction="west" minutes="44.55" orientation="longitude" pageId="16" pageNumber="313" precision="9" value="-55.7425">55°44.55'W</geoCoordinate>
,
<geoCoordinate id="E8905009FFB35F023BB1E56CFAA32D5F" box="[1180,1293,297,321]" degrees="26" direction="south" minutes="19.15" orientation="latitude" pageId="16" pageNumber="313" precision="9" value="-26.319166">26°19.15'S</geoCoordinate>
), under bark,
<quantity id="4A5C9B2BFFB35F023C6DE50CFCDB2D7F" box="[832,885,329,353]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="313" unit="m" value="90.0">
<elevation id="0689D1FDFFB35F023C6DE50CFCDB2D7F" box="[832,885,329,353]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="313" unit="m" value="90.0">90 m</elevation>
</quantity>
elevation,
<date id="F91A100EFFB35F023CD8E50CFBF32D7F" box="[1013,1117,329,353]" pageId="16" pageNumber="313" value="2000-04-12">
<collectingDate id="E95EE9E6FFB35F023CD8E50CFBF32D7F" box="[1013,1117,329,353]" pageId="16" pageNumber="313" value="2000-04-12">4.12.2000</collectingDate>
</date>
, leg.
<collectorName id="20515318FFB35F023BB2E50FFB562D7F" box="[1183,1272,329,353]" pageId="16" pageNumber="313">Z. Falin</collectorName>
#PAR1F00105 (
<collectionCode id="EBB5AE0BFFB35F023C6AE52CFC312D9F" box="[839,927,361,385]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="16" pageNumber="313">KNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F023C12E5ECFA33287F" pageId="16" pageNumber="313" type="diagnosis">
<paragraph id="8D1B36CEFFB35F023C12E5ECFA33287F" blockId="16.[831,1438,425,1121]" pageId="16" pageNumber="313">
<emphasis id="BFD0EADCFFB35F023C12E5ECFC022DDF" bold="true" box="[831,940,425,449]" pageId="16" pageNumber="313">Diagnosis</emphasis>
: The
<typeStatus id="521F886CFFB35F023CC6E5ECFBE72DDF" box="[1003,1097,425,449]" pageId="16" pageNumber="313" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4AA44D4DFFB35F023B40E5ECFB7B2DDF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1133,1237,425,449]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023B40E5ECFB7B2DDF" box="[1133,1237,425,449]" italics="true" pageId="16" pageNumber="313">G. denieri</emphasis>
</taxonomicName>
was the only male specimen that could be studied from
<collectingCountry id="F5B3765EFFB35F023BD8E58CFACD2DFF" box="[1269,1379,457,481]" name="Argentina" pageId="16" pageNumber="313">Argentina</collectingCountry>
. The pattern of punctation slightly differs from the Brazilian and Paraguayan specimens in punctation of the area between suture line and discal carina. In contrast to the Brazilian and the Paraguayan specimens, the type specimen from
<collectingCountry id="F5B3765EFFB35F023CECE62CFB822E9F" box="[961,1068,617,641]" name="Argentina" pageId="16" pageNumber="313">Argentina</collectingCountry>
has one more puncture in the overlapping zone between the irregular punctation of the anterior half and the row of punctures of the posterior half. However, the pattern and number of punctures of the anterior punctation varies considerably between the specimens. It seems not to be a specific character. Thus,
<taxonomicName id="4AA44D4DFFB35F023C6CE76CFBB42F5F" authorityName="SCHEERPELTZ" authorityYear="1970" box="[833,1050,809,833]" class="Insecta" family="Staphylinidae" genus="Calocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="BFD0EADCFFB35F023C6CE76CFBB42F5F" box="[833,1050,809,833]" italics="true" pageId="16" pageNumber="313">G. sanctaecatharinae</emphasis>
</taxonomicName>
is synonymised, in particular, as no differences were found in the aedeagus between the Brazilian and the Paraguayan specimens and the Argentinean type. In addition to the differentiating characters described under
<taxonomicName id="4AA44D4DFFB35F023CDBE7ECFBD32FDF" authorityName="SHARP" authorityYear="1887" box="[1014,1149,937,961]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFB35F023CDBE7ECFBD32FDF" box="[1014,1149,937,961]" italics="true" pageId="16" pageNumber="313">G. cognatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4AA44D4DFFB35F023B9EE7ECFA862FDF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1203,1320,937,961]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="BFD0EADCFFB35F023B9EE7ECFA862FDF" box="[1203,1320,937,961]" italics="true" pageId="16" pageNumber="313">G. klimschi</emphasis>
</taxonomicName>
,
<taxonomicName id="4AA44D4DFFB35F023A18E7ECFA322FDF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1333,1436,937,961]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023A18E7ECFA322FDF" box="[1333,1436,937,961]" italics="true" pageId="16" pageNumber="313">G. denieri</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="4AA44D4DFFB35F023CC8E78CFBD02FFF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[997,1150,969,993]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="obscuricolle">
<emphasis id="BFD0EADCFFB35F023CC8E78CFBD02FFF" box="[997,1150,969,993]" italics="true" pageId="16" pageNumber="313">G. obscuricolle</emphasis>
</taxonomicName>
in the denser microsculpture of the fore-body. However,
<taxonomicName id="4AA44D4DFFB35F023B8AE7ACFABD281F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1191,1299,1001,1025]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFB35F023B8AE7ACFABD281F" box="[1191,1299,1001,1025]" italics="true" pageId="16" pageNumber="313">G. denieri</emphasis>
</taxonomicName>
is distinctly smaller than
<taxonomicName id="4AA44D4DFFB35F023CF8E04CFBDC283F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[981,1138,1033,1057]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="obscuricolle">
<emphasis id="BFD0EADCFFB35F023CF8E04CFBDC283F" box="[981,1138,1033,1057]" italics="true" pageId="16" pageNumber="313">G. obscuricolle</emphasis>
</taxonomicName>
and the microsculpture is weaker. In contrast to
<taxonomicName id="4AA44D4DFFB35F023B1BE06CFB7E285F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1078,1232,1065,1089]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="313" phylum="Arthropoda" rank="species" species="obscuricolle">
<emphasis id="BFD0EADCFFB35F023B1BE06CFB7E285F" box="[1078,1232,1065,1089]" italics="true" pageId="16" pageNumber="313">G. obscuricolle</emphasis>
</taxonomicName>
, the apical lobe of the aedeagus is longer and thinner and has fewer sensillae.
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFB35F033C13E0CCFEC8287F" lastPageId="17" lastPageNumber="314" pageId="16" pageNumber="313" type="description">
<paragraph id="8D1B36CEFFB35F023C13E0CCFC2528FF" blockId="16.[830,1438,1161,2081]" pageId="16" pageNumber="313">
<emphasis id="BFD0EADCFFB35F023C13E0CCFC6E28BF" bold="true" box="[830,960,1161,1185]" pageId="16" pageNumber="313">Description</emphasis>
: Length:
<quantity id="4A5C9B2BFFB35F023B09E0CCFBD928BF" box="[1060,1143,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="313" unit="mm" value="2.4">2.4 mm</quantity>
. Colouration: Brown; elytra slightly lighter reddish brown; legs still lighter; yellowish brown;
</paragraph>
<paragraph id="8D1B36CEFFB35F023C12E0ACFC522ADF" blockId="16.[830,1438,1161,2081]" pageId="16" pageNumber="313">
Head:
<quantity id="4A5C9B2BFFB35F023CA8E0ACFC4A291F" box="[901,996,1257,1281]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="16" pageNumber="313" unit="mm" value="0.36">0.36 mm</quantity>
long,
<quantity id="4A5C9B2BFFB35F023B0BE0ACFB2B291F" box="[1062,1157,1257,1281]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="313" unit="mm" value="0.48">0.48 mm</quantity>
wide; eyes slightly prominent; twice as long as temples; temples narrowed to neck in short convex curve; cheek-like; in front of eyes, sides parallel up to base of antennae; parallel part approximately as long as length of eyes; then abruptly narrowed to anterior edge of clypeus in sinuate curve; anterior edge of clypeus 2.5 times as wide as distance between base of antennae; setiferous punctation deep and moderately dense; on vertex denser than on clypeus; on clypeus, interstices between punctures twice as wide as diameter of punctures; on vertex, interstices between punctures approximately as wide as diameter of punctures; setae at least as long as interstices between punctures; microsculpture weak; transversely or longitudinally striate; surface moderately shiny.
</paragraph>
<paragraph id="8D1B36CEFFB35F023C12E28CFADD2B5F" blockId="16.[830,1438,1161,2081]" pageId="16" pageNumber="313">Antennae as long as head and pronotum combined; all antennomeres nearly equally wide; slightly wider than long; second to tenth antennomere with long yellow setae; setae at least as long as width of antennomeres.</paragraph>
<paragraph id="8D1B36CEFFB35F033C12E30FFF7B2D9F" blockId="16.[830,1438,1161,2081]" lastBlockId="17.[150,758,233,1121]" lastPageId="17" lastPageNumber="314" pageId="16" pageNumber="313">
Pronotum:
<quantity id="4A5C9B2BFFB35F023C94E30CFBB62B7F" box="[953,1048,1865,1889]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="16" pageNumber="313" unit="mm" value="0.37">0.37 mm</quantity>
long,
<quantity id="4A5C9B2BFFB35F023B77E30CFB172B7F" box="[1114,1209,1865,1889]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="313" unit="mm" value="0.43">0.43 mm</quantity>
wide; widest in anterior third; in anterior third slightly narrowed to nearly rectangular angles; anterior edge slightly prominent; in posterior two third, sides narrowed to posterior angles in concave curve; posterior angles obtusely rounded; setiferous punctation deep and moderately dense; size of punctures varying; interstices between punctures at least as wide as diameter of punctures; lateral punctation denser than discal punctation; partly coriaceous; posterior angles with elongate deep grooves; moderately deep microsculpture longitudinally striate; surface slightly shiny.
</paragraph>
<paragraph id="8D1B36CEFFB25F033FBAE5CCFD6A2F5F" blockId="17.[150,758,233,1121]" pageId="17" pageNumber="314">
Elytra:
<quantity id="4A5C9B2BFFB25F033FC1E5CCFEFD2DBF" box="[236,339,393,417]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="314" unit="mm" value="0.62">0.62 mm</quantity>
long,
<quantity id="4A5C9B2BFFB25F033E8BE5CCFDA32DBF" box="[422,525,393,417]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="314" unit="mm" value="0.55">0.55 mm</quantity>
wide; sides slightly curved, but nearly parallel; shoulders rounded; anterior edge slightly emarginate; discal carina distinct; lateral carina absent; suture line formed by dense small punctures; continued to posterior margin; between suture line and discal carina with irregular setiferous punctures in anterior half and longitudinal row of 5 to 6 setiferous punctures in posterior half close to discal carina; at outer side of discal carina, dense row of elongate setiferous punctures; laterad with additional rows of elongate setiferous punctures; partly irregular in posterior half and with additional punctures between rows; microsculpture weak; netlike on surface of punctures; densely elongate on surface between punctures; surface slightly shiny.
</paragraph>
<paragraph id="8D1B36CEFFB25F033FBAE70CFDBA2FDF" blockId="17.[150,758,233,1121]" pageId="17" pageNumber="314">Abdomen on segments III to VI with two transverse irregular rows of large setiferous punctures; in posterior middle of segments III to VI with impunctate area; with weak microsculpture; surface shiny.</paragraph>
<paragraph id="8D1B36CEFFB25F033FBAE78CFEC8287F" blockId="17.[150,758,233,1121]" pageId="17" pageNumber="314">Aedeagus angulate between basal and apical lobe; angle obtuse; apical lobe narrow ending in acute apex; apex shortly curved; pointing ventrad; irregular row of sensillae at base of apical lobe; paramere reaching end of basal third of apical lobe.</paragraph>
</subSubSection>
</treatment>
</document>