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<document id="D98721DBDEAB992F4CE43A29EA41A490" ID-DOI="10.21248/contrib.entomol.65.2.297-325" ID-ISSN="0005-805X" ID-Zenodo-Dep="4753971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620838389347" checkinUser="carolina" docAuthor="Irmler, Ulrich" docDate="2015" docId="050D87D8FFAD5F023C48E776FE22291F" docLanguage="en" docName="ContribEntomol.65.2.297-325.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 65 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1885" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Glyptoma cognatum SHARP 1887" docType="treatment" docVersion="3" lastPageNumber="313" masterDocId="F934FFA0FFA35F123F2DE445FFAE2C1E" masterDocTitle="The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="325" masterPageNumber="297" pageNumber="311" updateTime="1698916894542" updateUser="plazi">
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<mods:title id="079CFB3C3CA7A97D0575A9980B7B7F5F">The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:namePart id="D1A15A89060B5C9F08FB01D6712FD847">Irmler, Ulrich</mods:namePart>
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<mods:title id="3F3133D0C28BC479EF3AAEA14779F224">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<treatment id="050D87D8FFAD5F023C48E776FE22291F" LSID="urn:lsid:plazi:treatment:050D87D8FFAD5F023C48E776FE22291F" httpUri="http://treatment.plazi.org/id/050D87D8FFAD5F023C48E776FE22291F" lastPageId="16" lastPageNumber="313" pageId="14" pageNumber="311">
<subSubSection id="C5BE6545FFAD5F1C3C48E776FB122F50" box="[869,1212,819,846]" pageId="14" pageNumber="311" type="nomenclature">
<paragraph id="8D1B36CEFFAD5F1C3C48E776FB122F50" blockId="14.[869,1212,819,879]" box="[869,1212,819,846]" pageId="14" pageNumber="311">
<heading id="D65381A2FFAD5F1C3C48E776FB122F50" box="[869,1212,819,846]" centered="true" fontSize="12" level="2" pageId="14" pageNumber="311" reason="2">
<taxonomicName id="4AA44D4DFFAD5F1C3C48E776FB122F50" ID-CoL="3GHR5" authority="SHARP, 1887" authorityName="SHARP" authorityYear="1887" box="[869,1212,819,846]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3C48E776FB9B2F50" box="[869,1077,819,846]" italics="true" pageId="14" pageNumber="311">Glyptoma cognatum</emphasis>
<bibRefCitation id="E9354B3FFFAD5F1C3B16E776FB122F50" author="SHARP, D." box="[1083,1212,819,846]" pageId="14" pageNumber="311" pagination="1 - 747" refId="ref23354" refString="SHARP, D. 1887: Biologia Centrali Americana: Insecta Coleoptera: 1 - 747." type="book chapter" year="1887">SHARP, 1887</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFAD5F1C3C48E71CFC532F71" box="[869,1021,857,879]" pageId="14" pageNumber="311" type="description">
<paragraph id="8D1B36CEFFAD5F1C3C48E71CFC532F71" blockId="14.[869,1212,819,879]" box="[869,1021,857,879]" pageId="14" pageNumber="311">
(
<figureCitation id="159F2A4BFFAD5F1C3C47E71CFC602F71" box="[874,974,857,879]" captionStart="Figs 1317" captionStartId="15.[151,191,2027,2049]" captionTargetBox="[197,1366,506,2003]" captionTargetId="figure-143@15.[151,1398,482,2026]" captionTargetPageId="15" captionText="Figs 1317: Aedeagus (A), antenna (B) and microsculpture of head (C) ofGlyptoma nitidum (13), G. obscuricolle (14), G. nitens (15), G. cognatum (16), G. denieri (17); scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4753985" httpUri="https://zenodo.org/record/4753985/files/figure.png" pageId="14" pageNumber="311">Figs 16A, B</figureCitation>
,
<figureCitation id="159F2A4BFFAD5F1C3CF8E71CFC572F71" box="[981,1017,857,879]" captionStart="Figs 22" captionStartId="27.[151,191,1934,1956]" captionTargetBox="[164,1386,260,1919]" captionTargetId="figure-41@27.[151,1398,253,1933]" captionTargetPageId="27" captionText="Figs 22: Fore body of Glyptoma nitidum (A), G. obscuricollis (B), G. nitens (C), G. cognatum (D), G. sanctaecatharinae (E), G. klimschi (F); scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753999" httpUri="https://zenodo.org/record/4753999/files/figure.png" pageId="14" pageNumber="311">22D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFAD5F1C3C12E7CEFB092FBE" box="[831,1191,907,929]" pageId="14" pageNumber="311" type="reference_group">
<paragraph id="8D1B36CEFFAD5F1C3C12E7CEFB092FBE" blockId="14.[831,1191,907,929]" box="[831,1191,907,929]" pageId="14" pageNumber="311">
<treatmentCitationGroup id="ADB411E0FFAD5F1C3C12E7CEFB092FBE" box="[831,1191,907,929]" pageId="14" pageNumber="311">
<treatmentCitation id="0C0510DFFFAD5F1C3C12E7CEFB092FBE" author="SHARP, D." box="[831,1191,907,929]" page="724" pageId="14" pageNumber="311" year="1887">
<taxonomicName id="4AA44D4DFFAD5F1C3C12E7CEFB092FBE" ID-CoL="3GHR5" authority="SHARP, 1887: 724" authorityName="SHARP" authorityPageNumber="724" authorityYear="1887" box="[831,1191,907,929]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3C12E7CEFC512FBF" box="[831,1023,907,929]" italics="true" pageId="14" pageNumber="311">Glyptoma cognatum</emphasis>
<bibRefCitation id="E9354B3FFFAD5F1C3B29E7CEFB092FBE" author="SHARP, D." box="[1028,1191,907,928]" pageId="14" pageNumber="311" pagination="1 - 747" refId="ref23354" refString="SHARP, D. 1887: Biologia Centrali Americana: Insecta Coleoptera: 1 - 747." type="book chapter" year="1887">SHARP, 1887: 724</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFAD5F1C3C12E78CFB2F291F" pageId="14" pageNumber="311" type="materials_examined">
<paragraph id="8D1B36CEFFAD5F1C3C12E78CFAE6281F" blockId="14.[831,1437,969,1026]" pageId="14" pageNumber="311">
<emphasis id="BFD0EADCFFAD5F1C3C12E78CFBFF2FFF" bold="true" box="[831,1105,969,993]" pageId="14" pageNumber="311">Type material examined</emphasis>
:
<materialsCitation id="3DCC3C93FFAD5F1C3B4FE78CFAEA281F" collectionCode="BMNH" collectorName="El Reposo &amp; C. Champion" country="Guatemala" location="Guatemala" pageId="14" pageNumber="311" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="syntype">
<collectingCountry id="F5B3765EFFAD5F1C3B4FE78CFB762FFF" box="[1122,1240,969,993]" name="Guatemala" pageId="14" pageNumber="311">Guatemala</collectingCountry>
:
<specimenCount id="9BA2FD47FFAD5F1C3BCAE78CFA9A2FFF" box="[1255,1332,969,993]" pageId="14" pageNumber="311" type="male">1 male</specimenCount>
,
<specimenCount id="9BA2FD47FFAD5F1C3A6EE78CFA332FFF" box="[1347,1437,969,993]" pageId="14" pageNumber="311" type="female">1 female</specimenCount>
<typeStatus id="521F886CFFAD5F1C3C12E7AFFC30281C" box="[831,926,1002,1026]" pageId="14" pageNumber="311" type="syntype">syntypes</typeStatus>
:
<collectorName id="20515318FFAD5F1C3C85E7ACFBBB281F" box="[936,1045,1001,1025]" pageId="14" pageNumber="311">El Reposo</collectorName>
, leg.
<collectorName id="20515318FFAD5F1C3B64E7ACFB76281F" box="[1097,1240,1001,1025]" pageId="14" pageNumber="311">C. Champion</collectorName>
(
<collectionCode id="EBB5AE0BFFAD5F1C3BCBE7ACFA92281F" box="[1254,1340,1001,1025]" pageId="14" pageNumber="311">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8D1B36CEFFAD5F1C3C12E06CFB2F291F" blockId="14.[831,1437,1065,1281]" pageId="14" pageNumber="311">
<emphasis id="BFD0EADCFFAD5F1C3C12E06CFBD6285F" bold="true" box="[831,1144,1065,1089]" pageId="14" pageNumber="311">Further material examined</emphasis>
:
<materialsCitation id="3DCC3C93FFAD5F1C3BA3E06FFA36289F" collectingDate="1968-05-24" collectionCode="AMNH" collectorName="R. Hutton" country="Panama" county="Albrook Forest" location="Canal Zone" municipality="Albrook Forest Site" pageId="14" pageNumber="311" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F5B3765EFFAD5F1C3BA3E06FFB48285F" box="[1166,1254,1066,1089]" name="Panama" pageId="14" pageNumber="311">Panama</collectingCountry>
:
<specimenCount id="9BA2FD47FFAD5F1C3BD4E06CFAE5285F" box="[1273,1355,1065,1089]" pageId="14" pageNumber="311" type="male">1 male</specimenCount>
,
<location id="887B6015FFAD5F1C3A73E06CFCD7287F" LSID="urn:lsid:plazi:treatment:050D87D8FFAD5F023C48E776FE22291F:887B6015FFAD5F1C3A73E06CFCD7287F" country="Panama" county="Albrook Forest" municipality="Albrook Forest Site" name="Canal Zone" pageId="14" pageNumber="311">Canal Zone</location>
,
<collectingMunicipality id="6D7FACB4FFAD5F1C3CA3E00CFBDD287F" box="[910,1139,1097,1121]" pageId="14" pageNumber="311">
<collectingCounty id="647A4E42FFAD5F1C3CA3E00CFB94287F" box="[910,1082,1097,1121]" pageId="14" pageNumber="311">Albrook Forest</collectingCounty>
Site
</collectingMunicipality>
, collected by black light (ground), 23.
<date id="F91A100EFFAD5F1C3CC8E02CFBFF289F" box="[997,1105,1129,1153]" pageId="14" pageNumber="311" value="1968-05-24">
<collectingDate id="E95EE9E6FFAD5F1C3CC8E02CFBFF289F" box="[997,1105,1129,1153]" pageId="14" pageNumber="311" value="1968-05-24">24.5.1968</collectingDate>
</date>
, leg.
<collectorName id="20515318FFAD5F1C3B8EE02FFAB2289F" box="[1187,1308,1130,1153]" pageId="14" pageNumber="311">R. Hutton</collectorName>
(
<collectionCode id="EBB5AE0BFFAD5F1C3A14E02CFA3F289F" box="[1337,1425,1129,1153]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="311">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFAD5F1C3C12E0CCFB4028DF" collectingDate="1968-05-16" collectionCode="AMNH" collectorName="R. Hutton" country="Panama" county="Albrook Forest" elevation="30" location="Canal Zone" municipality="Albrook Forest Site" pageId="14" pageNumber="311" specimenCount="1" specimenCount-female="1">
<specimenCount id="9BA2FD47FFAD5F1C3C12E0CCFC3028BF" box="[831,926,1161,1185]" pageId="14" pageNumber="311" type="female">1 female</specimenCount>
, same location,
<quantity id="4A5C9B2BFFAD5F1C3B62E0CCFB3B28BF" box="[1103,1173,1161,1185]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="14" pageNumber="311" unit="ft" value="100.0">
<elevation id="0689D1FDFFAD5F1C3B62E0CCFB3B28BF" box="[1103,1173,1161,1185]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="14" pageNumber="311" unit="ft" value="100.0">100 ft.</elevation>
</quantity>
elev., collected by black light,
<date id="F91A100EFFAD5F1C3C51E0ECFC4D28DF" box="[892,995,1193,1217]" pageId="14" pageNumber="311" value="1968-05-16">
<collectingDate id="E95EE9E6FFAD5F1C3C51E0ECFC4D28DF" box="[892,995,1193,1217]" pageId="14" pageNumber="311" value="1968-05-16">16.5.1968</collectingDate>
</date>
, leg.
<collectorName id="20515318FFAD5F1C3B35E0EFFB2C28DF" box="[1048,1154,1194,1217]" pageId="14" pageNumber="311">R. Hutton</collectorName>
(
<collectionCode id="EBB5AE0BFFAD5F1C3BA2E0ECFB4928DF" box="[1167,1255,1193,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="311">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DCC3C93FFAD5F1C3BD5E0ECFBD3291F" collectingDate="1999-03" collectingDateMax="1999-04-31" collectingDateMin="1999-03" collectionCode="AMNH, UIC" collectorName="A. Berkov" country="French Guiana" location="Saul" pageId="14" pageNumber="311" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry id="F5B3765EFFAD5F1C3BD5E0ECFA3728DF" box="[1272,1433,1193,1217]" name="French Guiana" pageId="14" pageNumber="311">French Guiana</collectingCountry>
:
<specimenCount id="9BA2FD47FFAD5F1C3C12E08CFC2628FF" box="[831,904,1225,1249]" pageId="14" pageNumber="311" type="male">1 male</specimenCount>
,
<specimenCount id="9BA2FD47FFAD5F1C3CB8E08CFC5028FF" box="[917,1022,1225,1249]" pageId="14" pageNumber="311" type="female">2 females</specimenCount>
,
<location id="887B6015FFAD5F1C3B26E08CFB9428FF" LSID="urn:lsid:plazi:treatment:050D87D8FFAD5F023C48E776FE22291F:887B6015FFAD5F1C3B26E08CFB9428FF" box="[1035,1082,1225,1249]" country="French Guiana" name="Saul" pageId="14" pageNumber="311">Saül</location>
, under bark,
<date id="F91A100EFFAD5F1C3BE2E08CFA3628FF" box="[1231,1432,1225,1249]" pageId="14" pageNumber="311" value="1999-03" valueMax="1999-04-31" valueMin="1999-03-00">
<collectingDate id="E95EE9E6FFAD5F1C3BE2E08CFA3628FF" box="[1231,1432,1225,1249]" pageId="14" pageNumber="311" value="1999-03" valueMax="1999-04-31" valueMin="1999-03-00">MarchApril 1999</collectingDate>
</date>
, leg.
<collectorName id="20515318FFAD5F1C3C47E0ACFC7B291F" box="[874,981,1257,1281]" pageId="14" pageNumber="311">A. Berkov</collectorName>
(
<collectionCode id="EBB5AE0BFFAD5F1C3CCFE0ACFB96291F" box="[994,1080,1257,1281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="311">AMNH</collectionCode>
,
<collectionCode id="EBB5AE0BFFAD5F1C3B6EE0ACFBDB291F" box="[1091,1141,1257,1281]" pageId="14" pageNumber="311">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFAD5F1C3C12E16CFB092ADF" pageId="14" pageNumber="311" type="diagnosis">
<paragraph id="8D1B36CEFFAD5F1C3C12E16CFB092ADF" blockId="14.[830,1437,1321,1729]" pageId="14" pageNumber="311">
<emphasis id="BFD0EADCFFAD5F1C3C12E16CFC02295F" bold="true" box="[831,940,1321,1345]" pageId="14" pageNumber="311">Diagnosis</emphasis>
: The three species
<taxonomicName id="4AA44D4DFFAD5F1C3B5FE16CFB58295F" authorityName="SHARP" authorityYear="1887" box="[1138,1270,1321,1345]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3B5FE16CFB58295F" box="[1138,1270,1321,1345]" italics="true" pageId="14" pageNumber="311">G. cognatum</emphasis>
</taxonomicName>
,
<taxonomicName id="4AA44D4DFFAD5F1C3A2EE16CFAC7295F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1283,1385,1321,1345]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3A2EE16CFAC7295F" box="[1283,1385,1321,1345]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4AA44D4DFFAD5F1C3C12E10CFC1D297F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[831,947,1353,1377]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="BFD0EADCFFAD5F1C3C12E10CFC1D297F" box="[831,947,1353,1377]" italics="true" pageId="14" pageNumber="311">G. klimschi</emphasis>
</taxonomicName>
are very similar in size and shape of the aedeagus.
<taxonomicName id="4AA44D4DFFAD5F1C3C42E12CFC5D299F" authorityName="SHARP" authorityYear="1887" box="[879,1011,1385,1409]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3C42E12CFC5D299F" box="[879,1011,1385,1409]" italics="true" pageId="14" pageNumber="311">G. cognatum</emphasis>
</taxonomicName>
can be differentiated from the two other species by the intermediate size from the slightly larger
<taxonomicName id="4AA44D4DFFAD5F1C3C12E1ECFC0929DF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[831,935,1449,1473]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3C12E1ECFC0929DF" box="[831,935,1449,1473]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
and the slightly smaller
<taxonomicName id="4AA44D4DFFAD5F1C3B93E1ECFA9929DF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1214,1335,1449,1473]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="BFD0EADCFFAD5F1C3B93E1ECFA9929DF" box="[1214,1335,1449,1473]" italics="true" pageId="14" pageNumber="311">G. klimschi</emphasis>
</taxonomicName>
. Regarding the microsculpture of the head and pronotum as well as both the thin apical lobe and few sensillae of the aedeagus, it resembles
<taxonomicName id="4AA44D4DFFAD5F1C3B00E24CFB3D2A3F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1069,1171,1545,1569]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3B00E24CFB3D2A3F" box="[1069,1171,1545,1569]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
. However, the microsculpture of
<taxonomicName id="4AA44D4DFFAD5F1C3CB3E26CFB8D2A5F" authorityName="SHARP" authorityYear="1887" box="[926,1059,1577,1601]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3CB3E26CFB8D2A5F" box="[926,1059,1577,1601]" italics="true" pageId="14" pageNumber="311">G. cognatum</emphasis>
</taxonomicName>
is weaker than in
<taxonomicName id="4AA44D4DFFAD5F1C3BCDE26CFAE92A5F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1248,1351,1577,1601]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3BCDE26CFAE92A5F" box="[1248,1351,1577,1601]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
and the number of punctures in the posterior half of the elytra is lower. In contrast to
<taxonomicName id="4AA44D4DFFAD5F1C3B0DE22CFB262A9F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1056,1160,1641,1665]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3B0DE22CFB262A9F" box="[1056,1160,1641,1665]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
, the apex of the aedeagus of
<taxonomicName id="4AA44D4DFFAD5F1C3C71E2CCFC4C2ABF" authorityName="SHARP" authorityYear="1887" box="[860,994,1673,1697]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAD5F1C3C71E2CCFC4C2ABF" box="[860,994,1673,1697]" italics="true" pageId="14" pageNumber="311">G. cognatum</emphasis>
</taxonomicName>
is curved pointing ventrad, whereas it is more or less straight in
<taxonomicName id="4AA44D4DFFAD5F1C3B14E2ECFB0F2ADF" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[1081,1185,1705,1729]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="311" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAD5F1C3B14E2ECFB0F2ADF" box="[1081,1185,1705,1729]" italics="true" pageId="14" pageNumber="311">G. denieri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C5BE6545FFAD5F023C12E2ACFE22291F" lastPageId="16" lastPageNumber="313" pageId="14" pageNumber="311" type="description">
<paragraph id="8D1B36CEFFAD5F1C3C12E2ACFACD2B3F" blockId="14.[831,1438,1769,2081]" pageId="14" pageNumber="311">
<emphasis id="BFD0EADCFFAD5F1C3C12E2ACFC6F2B1F" bold="true" box="[831,961,1769,1793]" pageId="14" pageNumber="311">Description</emphasis>
: Length:
<quantity id="4A5C9B2BFFAD5F1C3B18E2ACFB102B1F" box="[1077,1214,1769,1793]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="14" pageNumber="311" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.12.2 mm</quantity>
. Colouration: Dark brown; elytra lighter red; legs still lighter, yellowish.
</paragraph>
<paragraph id="8D1B36CEFFAD5F1D3C12E36CFF782DBF" blockId="14.[831,1438,1769,2081]" lastBlockId="15.[151,756,233,417]" lastPageId="15" lastPageNumber="312" pageId="14" pageNumber="311">
Head:
<quantity id="4A5C9B2BFFAD5F1C3CA8E36CFC4A2B5F" box="[901,996,1833,1857]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="311" unit="mm" value="0.27">0.27 mm</quantity>
long,
<quantity id="4A5C9B2BFFAD5F1C3B08E36CFB2A2B5F" box="[1061,1156,1833,1857]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="311" unit="mm" value="0.4">0.40 mm</quantity>
wide; eyes slightly prominent; nearly twice as long as temples; temples narrowed to neck in short strongly convex curve; sides in front of eyes parallel up to base of antennae; parallel part as long as length of eyes; at base of antennae abruptly narrowed to anterior edge of clypeus in nearly rectangular angle; anterior edge of clypeus slightly shorter than distance between base of antennae; setiferous punctures deep and moderately dense; interstices between punctures ranging between twice and three times as wide as diameter of punctures; close to neck, punctation denser; with weak irregular microsculpture; partly longitudinally striate; partly transversely striate; surface moderately shiny.
</paragraph>
<paragraph id="8D1B36CEFFAC5F1D3C37E4ACFAC42DDF" blockId="15.[794,1400,233,449]" pageId="15" pageNumber="312">Antennae as long as head and pronotum combined; all antennomeres nearly equally wide; second antennomere nearly twice as wide as long; third to fifth antennomere approximately 2.5 to 3 times as wide as long; tenth antennomere again twice as wide as long; second to tenth antennomere with long yellow setae; on average, setae approximately as long as half of width of antennomeres.</paragraph>
<caption id="D9DB6646FFAC5F1D3FBAE3AEFD222401" ID-DOI="http://doi.org/10.5281/zenodo.4753985" ID-Zenodo-Dep="4753985" httpUri="https://zenodo.org/record/4753985/files/figure.png" pageId="15" pageNumber="312" startId="15.[151,191,2027,2049]" targetBox="[197,1366,506,2003]" targetPageId="15">
<paragraph id="8D1B36CEFFAC5F1D3FBAE3AEFD222401" blockId="15.[151,1399,2027,2079]" pageId="15" pageNumber="312">
<emphasis id="BFD0EADCFFAC5F1D3FBAE3AEFEAE241F" bold="true" box="[151,256,2027,2049]" pageId="15" pageNumber="312">Figs 1317</emphasis>
: Aedeagus (A), antenna (B) and microsculpture of head (C) of
<taxonomicName id="4AA44D4DFFAC5F1D3C62E3A9FC55241F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[847,1019,2028,2049]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="312" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis id="BFD0EADCFFAC5F1D3C62E3A9FC55241F" box="[847,1019,2028,2049]" italics="true" pageId="15" pageNumber="312">Glyptoma nitidum</emphasis>
</taxonomicName>
(
<emphasis id="BFD0EADCFFAC5F1D3B24E3A9FB8F241F" bold="true" box="[1033,1057,2028,2049]" pageId="15" pageNumber="312">13</emphasis>
),
<taxonomicName id="4AA44D4DFFAC5F1D3B1EE3A9FB15241F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1075,1211,2028,2049]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="312" phylum="Arthropoda" rank="species" species="obscuricolle">
<emphasis id="BFD0EADCFFAC5F1D3B1EE3A9FB15241F" box="[1075,1211,2028,2049]" italics="true" pageId="15" pageNumber="312">G. obscuricolle</emphasis>
</taxonomicName>
(
<emphasis id="BFD0EADCFFAC5F1D3BE5E3A9FB4E241F" bold="true" box="[1224,1248,2028,2049]" pageId="15" pageNumber="312">14</emphasis>
),
<taxonomicName id="4AA44D4DFFAC5F1D3BDFE3A9FAEA241F" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1266,1348,2028,2049]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="312" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="BFD0EADCFFAC5F1D3BDFE3A9FAEA241F" box="[1266,1348,2028,2049]" italics="true" pageId="15" pageNumber="312">G. nitens</emphasis>
</taxonomicName>
(
<emphasis id="BFD0EADCFFAC5F1D3A7CE3A9FAC7241F" bold="true" box="[1361,1385,2028,2049]" pageId="15" pageNumber="312">15</emphasis>
),
<taxonomicName id="4AA44D4DFFAC5F1D3FBAEC4CFEBE2401" authorityName="SHARP" authorityYear="1887" box="[151,272,2057,2079]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="312" phylum="Arthropoda" rank="species" species="cognatum">
<emphasis id="BFD0EADCFFAC5F1D3FBAEC4CFEBE2401" box="[151,272,2057,2079]" italics="true" pageId="15" pageNumber="312">G. cognatum</emphasis>
</taxonomicName>
(
<emphasis id="BFD0EADCFFAC5F1D3E30EC4CFE9B2400" bold="true" box="[285,309,2057,2078]" pageId="15" pageNumber="312">16</emphasis>
),
<taxonomicName id="4AA44D4DFFAC5F1D3E65EC4CFE082400" baseAuthorityName="BERNHAUER" baseAuthorityYear="1939" box="[328,422,2057,2078]" class="Insecta" family="Staphylinidae" genus="Glyptoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="312" phylum="Arthropoda" rank="species" species="denieri">
<emphasis id="BFD0EADCFFAC5F1D3E65EC4CFE082400" box="[328,422,2057,2078]" italics="true" pageId="15" pageNumber="312">G. denieri</emphasis>
</taxonomicName>
(
<emphasis id="BFD0EADCFFAC5F1D3E9EEC4CFE652400" bold="true" box="[435,459,2057,2078]" pageId="15" pageNumber="312">17</emphasis>
); scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D1B36CEFFB35F023F90E4AFFD4D2FFF" blockId="16.[188,795,233,1281]" pageId="16" pageNumber="313">
Pronotum:
<quantity id="4A5C9B2BFFB35F023E15E4ACFE362D1F" box="[312,408,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="313" unit="mm" value="0.29">0.29 mm</quantity>
long
<quantity id="4A5C9B2BFFB35F023EF8E4ACFD9B2D1F" box="[469,565,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="313" unit="mm" value="0.35">0.35 mm</quantity>
wide; widest at anterior third; narrowed to anterior angles in short convex curve; narrowed to posterior angles in long, slightly concave curve; anterior and posterior angles obtuse; front edge slightly prominent; posterior edge as wide as neck; setiferous punctures large, but size irregular; partly coriaceous; on average, interstices between punctures mostly restricted to very narrow ridges; at posterior angles with irregular elongate impression; narrow midline impunctate; disc bordered by indistinct, irregularly sinuate carinae; insertion of setae partly centric, partly eccentric; microsculpture weak; longitudinally striate; surface shiny. Elytra:
<quantity id="4A5C9B2BFFB35F023E22E62CFEDA2E9F" box="[271,372,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="313" unit="mm" value="0.48">0.48 mm</quantity>
long,
<quantity id="4A5C9B2BFFB35F023EEDE62CFD8B2E9F" box="[448,549,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="313" unit="mm" value="0.45">0.45 mm</quantity>
wide; sides smoothly rounded; nearly parallel; shoulders distinct; shortly rounded to emarginate anterior edge; suture line formed by densely coriaceous small punctures; continued to posterior margin; discal carina distinct; lateral carina absent or only partly developed; discal carina margined at inner and outer edge by longitudinal row of elongate setiferous punctures; in anterior half on area between suture line and discal carina, several setiferous punctures; in posterior half impunctate; laterad to discal carina three longitudinal rows of setiferous punctures; extremely weak microsculpture; longitudinally striate; surface shiny.
</paragraph>
<paragraph id="8D1B36CEFFB35F023F90E7ACFE53287F" blockId="16.[188,795,233,1281]" pageId="16" pageNumber="313">Abdomen on segments III to VI with two transverse irregular rows of large setiferous punctures; posterior half of segments III to VI with impunctate area; without microsculpture; surface shiny.</paragraph>
<paragraph id="8D1B36CEFFB35F023F90E02CFE22291F" blockId="16.[188,795,233,1281]" pageId="16" pageNumber="313">Aedeagus angulate between basal and apical lobe; angle obtuse; apical lobe narrow ending in acute apex; apex shortly curved; pointing ventrad; very few small sensillae at base of apical lobe; paramere reaching end of basal third of apical lobe.</paragraph>
</subSubSection>
</treatment>
</document>