305 lines
32 KiB
XML
305 lines
32 KiB
XML
<document id="B47C08D9D874A8713718CD23FD2AC945" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas" docDate="2015" docId="03A25264CADDFF2DEE91FAEB4380FDDA" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Orthetrum umbratum Dijkstra & Meziere, sp. nov." docType="treatment" docVersion="21" lastPageNumber="644" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="641" updateTime="1695001031402" updateUser="admin">
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<mods:titleInfo id="D5E8842E90F32E982072758E26EF4053">
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<mods:title id="31A8EC5F5AEFC6F29E6E1BA5466D4C35">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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</mods:titleInfo>
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<mods:name id="94A1D8A57D87E1A0FD88E27316FC2825" type="personal">
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<mods:roleTerm id="266852A8B3F50B6E5CB4D0BEEE61A181">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5D01934F338D4A312F8905F3532BBFD6">Dijkstra, Klaas-Douwe B.</mods:namePart>
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</mods:name>
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<mods:name id="348F05B1536C047D77D5BF9BAD1A5D4C" type="personal">
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<mods:roleTerm id="0E5E671CE426395E938FCD19C3F6A11B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DE02D877A05890A48B768CA72C90AC94">Kipping, Jens</mods:namePart>
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</mods:name>
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<mods:name id="715193181987BA59EEAD4A262D42C52C" type="personal">
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<mods:role id="AC2E69E259DB77FC7FCBC73B0245C4D5">
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<mods:roleTerm id="2843E8AA9FC44A5492AD82E1453770E0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="06388DCA61145327F55D0397F50242B6">Mézière, Nicolas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0277E9C4DE0C21DF96687838DB2E1A9D">text</mods:typeOfResource>
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<mods:titleInfo id="E3888C67D636DB7BA3E587E33497902A">
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<mods:title id="DFB7CAB4A1F1DD00E8AE558F0ED9BF70">Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part id="ED3969EB8AF13E19D24C3FABFAAF8C81">
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<mods:date id="59E0494422BE3C23EE3E22B3D2EC3038">2015</mods:date>
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<mods:detail id="0431794C957CDAC324FED90D4C04A011" type="volume">
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<mods:number id="3EFB954E253EB8A4271DC72CEF37DC48">44</mods:number>
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</mods:detail>
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<mods:detail id="3095418CD3DDB28140CF1A286AB5F073" type="issue">
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<mods:number id="C6E10DD7DE50218BDD2135E0D6CDFA26">4</mods:number>
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<mods:start id="6DCCB09C2E1A8985EFA72EB4C1D63B84">447</mods:start>
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<mods:end id="2A7507E3051F4C0310D49E4031679270">678</mods:end>
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<mods:classification id="D30959880AEDFD238E80ABAFF91C0172">journal article</mods:classification>
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<mods:identifier id="F9937EE077356B989F1223D6F68CC2E6" type="DOI">10.5281/zenodo.35388</mods:identifier>
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</mods:mods>
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<treatment id="03A25264CADDFF2DEE91FAEB4380FDDA" ID-DOI="http://doi.org/10.5281/zenodo.5640298" ID-GBIF-Taxon="114923463" ID-Zenodo-Dep="5640298" LSID="urn:lsid:zoobank.org:act:03A25264-CADD-FF2D-EE91-FAEB4380FDDA" httpUri="http://treatment.plazi.org/id/03A25264CADDFF2DEE91FAEB4380FDDA" lastPageId="199" lastPageNumber="644" pageId="196" pageNumber="641">
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<subSubSection id="C311B0F9CADDFF2EEE91FAEB421BFAFF" pageId="196" pageNumber="641" type="nomenclature">
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<paragraph id="8BB4E372CADDFF2EEE91FAEB421BFAFF" pageId="196" pageNumber="641">
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||
<heading id="D0FC541ECADDFF2EEE91FAEB421BFAFF" bold="true" centered="true" fontSize="11" level="2" pageId="196" pageNumber="641" reason="4">
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||
<emphasis id="B97F3F60CADDFF2EEE91FAEB42FDFA90" bold="true" box="[179,1016,1285,1314]" pageId="196" pageNumber="641">
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||
<taxonomicName id="4C0B98F1CADDFF2EEE91FAEB43A1FA90" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CADD-FF2D-EE91-FAEB4380FDDA" authority="Dijkstra & Meziere" authorityName="Dijkstra & Meziere" box="[179,676,1285,1314]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="196" pageNumber="641" phylum="Arthropoda" rank="species" species="umbratum" status="sp. nov.">
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<emphasis id="B97F3F60CADDFF2EEE91FAEB40B9FA90" bold="true" box="[179,444,1286,1314]" italics="true" pageId="196" pageNumber="641">Orthetrum umbratum</emphasis>
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Dijkstra & Mézière
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</taxonomicName>
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<taxonomicNameLabel id="A24C821BCADDFF2EEC89FAEA420DFA90" box="[683,776,1287,1314]" pageId="196" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
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– Shadow Skimmer
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</emphasis>
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||
(
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<typeStatus id="54B05DD0CADDFF2EEFB1FADC40CBFAFF" box="[403,462,1329,1357]" pageId="196" pageNumber="641">Type</typeStatus>
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||
<figureCitation id="1330FFF7CADDFF2EEFF6FADC4341FAFE" box="[468,580,1329,1356]" captionStart="Photo 53" captionStartId="153.[159,225,1465,1490]" captionTargetBox="[159,1035,771,1423]" captionTargetId="figure@153.[159,1038,771,1425]" captionTargetPageId="153" captionText="Photo 53. Paragomphus dispar, male, near Moyabi, Gabon. When flying rapidly low over the water, males are hard to see except for their unique markings. Photo: JK (17 - ix- 2013)" pageId="196" pageNumber="641">Photo 53</figureCitation>
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,
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<figureCitation id="1330FFF7CADDFF2EEC6CFADC43B8FAFE" box="[590,701,1329,1356]" captionStart="Photo 68" captionStartId="198.[159,225,1498,1522]" captionTargetBox="[159,1035,877,1457]" captionTargetId="figure@198.[159,1038,874,1457]" captionTargetPageId="198" captionText="Photo 68. Orthetrum umbratum, male; Moukoumbi, Gabon. Photo: NM (10 - iv- - 2012)" pageId="196" pageNumber="641">Photo 68</figureCitation>
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,
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<figureCitation id="1330FFF7CADDFF2EECE5FADC4213FAFF" box="[711,790,1329,1357]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="196" pageNumber="641">Fig. 31</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C311B0F9CADDFF2EEEBDFA8D401AFAC8" box="[159,287,1376,1402]" pageId="196" pageNumber="641" type="description">
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||
<paragraph id="8BB4E372CADDFF2EEEBDFA8D401AFAC8" box="[159,287,1376,1402]" pageId="196" pageNumber="641">
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||
<heading id="D0FC541ECADDFF2EEEBDFA8D401AFAC8" bold="true" box="[159,287,1376,1402]" fontSize="11" level="3" pageId="196" pageNumber="641" reason="6">
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<emphasis id="B97F3F60CADDFF2EEEBDFA8D401AFAC8" bold="true" box="[159,287,1376,1402]" pageId="196" pageNumber="641">Taxonomy</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C311B0F9CADDFF2EEEBDFA6941F2F9BD" pageId="196" pageNumber="641" type="nomenclature">
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<paragraph id="8BB4E372CADDFF2EEEBDFA6941F2F9BD" pageId="196" pageNumber="641">
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Morphologically and ecologically similar to
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<taxonomicName id="4C0B98F1CADDFF2EEC9FFA684236FA2D" box="[701,819,1413,1439]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="196" pageNumber="641" phylum="Arthropoda" rank="species" species="saegeri">
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<emphasis id="B97F3F60CADDFF2EEC9FFA684236FA2D" box="[701,819,1413,1439]" italics="true" pageId="196" pageNumber="641">O. saegeri</emphasis>
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</taxonomicName>
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(see
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<taxonomicName id="4C0B98F1CADDFF2EED5AFA684509FA2D" box="[888,1036,1413,1439]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="196" pageNumber="641" phylum="Arthropoda" rank="species" species="agaricum" status="sp. nov.">
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<emphasis id="B97F3F60CADDFF2EED5AFA684509FA2D" box="[888,1036,1413,1439]" italics="true" pageId="196" pageNumber="641">O. agaricum</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A24C821BCADDFF2EEEBDFA4641FBFA76" box="[159,254,1450,1476]" pageId="196" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
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), but differs clearly in the shape of the hamule. May genetically be distinct from sympatric
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<taxonomicName id="4C0B98F1CADDFF2EEFC1FA224353FA58" box="[483,598,1487,1514]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="196" pageNumber="641" phylum="Arthropoda" rank="species" species="saegeri">
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<emphasis id="B97F3F60CADDFF2EEFC1FA224353FA58" box="[483,598,1487,1514]" italics="true" pageId="196" pageNumber="641">O. saegeri</emphasis>
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</taxonomicName>
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, but this is not entirely resolved (see below).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C311B0F9CADCFF2FEEBDFF1343D0FDDA" pageId="197" pageNumber="642" type="materials_examined">
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||
<paragraph id="8BB4E372CADCFF2FEEBDFF13406DFEAA" box="[159,360,254,280]" pageId="197" pageNumber="642">
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||
<heading id="D0FC541ECADCFF2FEEBDFF13406DFEAA" bold="true" box="[159,360,254,280]" fontSize="11" level="3" pageId="197" pageNumber="642" reason="6">
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||
<emphasis id="B97F3F60CADCFF2FEEBDFF13406DFEAA" bold="true" box="[159,360,254,280]" pageId="197" pageNumber="642">Material studied</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BB4E372CADCFF2FEEBDFEC9406CFE1F" pageId="197" pageNumber="642">
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||
<materialsCitation id="3B63E92FCADCFF2FEEBDFEC94060FE1F" ID-GBIF-Occurrence="3396361375" collectingDate="2012-03-10" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="532 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF370-15" latitude="-1.0836" location="Moanda- Lastoursville road about 4 km after Moukoumbi" longLatPrecision="7" longitude="12.8923" pageId="197" pageNumber="642" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-Ogooué Province" typeStatus="holotype">
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||
<typeStatus id="54B05DD0CADCFF2FEEBDFEC94015FE8F" box="[159,272,292,317]" pageId="197" pageNumber="642">
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||
<emphasis id="B97F3F60CADCFF2FEEBDFEC94015FE8F" bold="true" box="[159,272,292,317]" pageId="197" pageNumber="642">Holotype</emphasis>
|
||
</typeStatus>
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||
<emphasis id="B97F3F60CADCFF2FEF34FECF402FFE8E" bold="true" box="[278,298,290,316]" pageId="197" pageNumber="642">
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||
<specimenCount id="A7AE7317BDC5412271E2BF6816D390C8" type="male">♂</specimenCount>
|
||
</emphasis>
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||
<emphasis id="B97F3F60CADCFF2FEF08FEC94034FE8F" bold="true" box="[298,305,292,317]" pageId="197" pageNumber="642">.</emphasis>
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||
<specimenCode id="FEFDD036288AF0CAD43D33C0F2D8638E">RMNH.INS.506140</specimenCode>
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||
,
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||
<collectingCountry id="F31CA3E2CADCFF2FEC08FECF4379FE8F" box="[554,636,290,317]" name="Gabon" pageId="197" pageNumber="642" value="Gabon">Gabon</collectingCountry>
|
||
,
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||
<collectingRegion id="49CF2D90CADCFF2FECA7FECF4293FE8F" box="[645,918,290,317]" pageId="197" pageNumber="642">Haut-Ogooué Province</collectingRegion>
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||
,
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||
<location id="C98A4C2AB489C9E950674CB1ACD1D402" LSID="urn:lsid:plazi:treatment:03A25264CADDFF2DEE91FAEB4380FDDA:C98A4C2AB489C9E950674CB1ACD1D402" country="Gabon" latitude="-1.0836" longLatPrecision="7" longitude="12.8923" name="Moanda- Lastoursville road about 4 km after Moukoumbi" stateProvince="Haut-Ogooué Province">
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||
Moanda- Lastoursville road about
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||
<quantity id="4CF34E97CADCFF2FEFE7FEA54307FED0" box="[453,514,328,354]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="197" pageNumber="642" unit="km" value="4.0">4 km</quantity>
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after Moukoumbi
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||
</location>
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, roadside stream, swamp, forest and spring,
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||
<quantity id="4CF34E97CADCFF2FEFA3FE8340CAFE3A" box="[385,463,366,392]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="197" pageNumber="642" unit="m" value="532.0">
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||
<elevation id="2EB7E4D9FA62F404B82D90FA36981699" box="[385,463,366,392]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="197" pageNumber="642" unit="m" value="532.0">532 m</elevation>
|
||
</quantity>
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||
a.s.l. (
|
||
<geoCoordinate id="EE3F85B5CADCFF2FEC00FE80438CFE3A" box="[546,649,365,392]" direction="south" orientation="latitude" pageId="197" pageNumber="642" precision="5" value="-1.0836">1.0836 ° S</geoCoordinate>
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<geoCoordinate id="EE3F85B5CADCFF2FECB7FE80420AFE3A" box="[661,783,365,392]" direction="east" orientation="longitude" pageId="197" pageNumber="642" precision="5" value="12.8923">12.8923 ° E</geoCoordinate>
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||
),
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||
<date id="FFB5C5B2CADCFF2FED05FE8042AFFE3A" box="[807,938,365,392]" pageId="197" pageNumber="642" value="2012-03-10">
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||
<collectingDate id="527A4ADF006B9C82B9A7405BA82C6FD6" box="[807,938,365,392]" pageId="197" pageNumber="642" value="2012-03-10">10 -iii- 2012</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="0B1E4902923616AB174A0F6CB4225440">N. Mézière</collectorName>
|
||
,
|
||
<collectionCode id="ED1A7BB7CADCFF2FEF2BFE7E4060FE1F" box="[265,357,403,429]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="197" pageNumber="642">RMNH</collectionCode>
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
<paragraph id="8BB4E372CADCFF2FEEBDFE5443D0FDDA" pageId="197" pageNumber="642">
|
||
<emphasis id="B97F3F60CADCFF2FEEBDFE544087FE61" bold="true" box="[159,386,440,467]" pageId="197" pageNumber="642">Further material.</emphasis>
|
||
<materialsCitation id="3B63E92FCADCFF2FEFBAFE554339FE4A" ID-GBIF-Occurrence="3396361329" collectingDate="2012-03-10" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="532 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF379-15" latitude="-1.0836" location="Moanda- Lastoursville road about 4 km after Moukoumbi" longLatPrecision="7" longitude="12.8923" pageId="197" pageNumber="642" specimenCode="RMNH.INS.554425" specimenCount="5" specimenCount-male="5" stateProvince="Haut-Ogooué Province">
|
||
<collectingCountry id="F31CA3E2CADCFF2FEFBAFE554305FE60" box="[408,512,440,466]" name="Gabon" pageId="197" pageNumber="642" value="Gabon">GABON</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CF2D90CADCFF2FEC3DFE5A4243FE60" box="[543,838,439,466]" pageId="197" pageNumber="642">Haut-Ogooué Province</collectingRegion>
|
||
):
|
||
<specimenCount id="9D0D28FBCADCFF2FED4BFE54428EFE60" box="[873,907,440,466]" pageId="197" pageNumber="642" type="generic">5 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DAE645ACC52640DF4A50C7EA57A6C016">RMNH.INS.506180</specimenCode>
|
||
), as holotype,
|
||
<collectionCode id="ED1A7BB7CADCFF2FEFC2FE334339FE4A" box="[480,572,478,504]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="197" pageNumber="642">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B63E92FCADCFF2FEC68FE3343D4FDDA" ID-GBIF-Occurrence="3396361306" collectingDate="2013-05-08" collectionCode="RMNH" collectorName="N. Mézière, J. Renoult" country="Gabon" elevation="577 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF512-15" latitude="-1.7437" location="Bakoumba, Parc de la Lékédi, Module 3" longLatPrecision="7" longitude="12.9354" pageId="197" pageNumber="642" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-ogooue Province">
|
||
<specimenCount id="9D0D28FBCADCFF2FEC68FE334369FE45" box="[586,620,477,504]" pageId="197" pageNumber="642" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="CA7E43481723E400CC70D275D3161BB2">RMNH.INS.508810</specimenCode>
|
||
),
|
||
<location id="013904A3D458021185F0EEC3382B41A1" LSID="urn:lsid:plazi:treatment:03A25264CADDFF2DEE91FAEB4380FDDA:013904A3D458021185F0EEC3382B41A1" country="Gabon" latitude="-1.7437" longLatPrecision="7" longitude="12.9354" name="Bakoumba, Parc de la Lékédi, Module 3" stateProvince="Haut-ogooue Province">Bakoumba, Parc de la Lékédi, Module 3</location>
|
||
, sandy forest stream (
|
||
<quantity id="4CF34E97CADCFF2FECCAFDEF422BFDAF" box="[744,814,514,541]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="197" pageNumber="642" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
||
wide,
|
||
<quantity id="4CF34E97CADCFF2FED59FDEE42C6FDAF" box="[891,963,515,541]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="197" pageNumber="642" unit="cm" value="20.0">20 cm</quantity>
|
||
deep) with bordering seep, sources and ponds,
|
||
<quantity id="4CF34E97CADCFF2FECA0FDC443C9FDF0" box="[642,716,553,578]" metricMagnitude="2" metricUnit="m" metricValue="5.77" pageId="197" pageNumber="642" unit="m" value="577.0">
|
||
<elevation id="38323A33B108490A1D049F481ABB7B70" box="[642,716,553,578]" metricMagnitude="2" metricUnit="m" metricValue="5.77" pageId="197" pageNumber="642" unit="m" value="577.0">577 m</elevation>
|
||
</quantity>
|
||
a.s.l. (
|
||
<geoCoordinate id="EE3F85B5CADCFF2FED34FDC54278FDF0" box="[790,893,552,578]" direction="south" orientation="latitude" pageId="197" pageNumber="642" precision="5" value="-1.7437">1.7437 ° S</geoCoordinate>
|
||
<geoCoordinate id="EE3F85B5CADCFF2FEDA7FDC542FAFDF0" box="[901,1023,552,578]" direction="east" orientation="longitude" pageId="197" pageNumber="642" precision="5" value="12.9354">12.9354 ° E</geoCoordinate>
|
||
),
|
||
<date id="FFB5C5B2CADCFF2FEEBDFDA3401DFDDA" box="[159,280,590,616]" pageId="197" pageNumber="642" value="2013-05-08">
|
||
<collectingDate id="6534DD5CC47C1ED9238B0D50310B8A2D" box="[159,280,590,616]" pageId="197" pageNumber="642" value="2013-05-08">08-v- 2013</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="3A7B1EA451FDFBA260116A93F0BA0F98">N. Mézière</collectorName>
|
||
&
|
||
<collectorName id="584D8BDE0CE8072BAF3815317538ED11">J. Renoult</collectorName>
|
||
,
|
||
<collectionCode id="ED1A7BB7CADCFF2FEC57FDA343D4FDDA" box="[629,721,590,616]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="197" pageNumber="642">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311B0F9CADCFF2FEEBDFD754271FCFA" pageId="197" pageNumber="642" type="description">
|
||
<paragraph id="8BB4E372CADCFF2FEEBDFD75400CFD01" box="[159,265,664,691]" pageId="197" pageNumber="642">
|
||
<heading id="D0FC541ECADCFF2FEEBDFD75400CFD01" bold="true" box="[159,265,664,691]" fontSize="11" level="3" pageId="197" pageNumber="642" reason="6">
|
||
<emphasis id="B97F3F60CADCFF2FEEBDFD75400CFD01" bold="true" box="[159,265,664,691]" pageId="197" pageNumber="642">Genetics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CADCFF2FEEBDFD504271FCFA" pageId="197" pageNumber="642">
|
||
Five unique haplotypes (n = 7) are quite distinct from five of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEDB4FD534509FD6A" box="[918,1036,702,728]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADCFF2FEDB4FD534509FD6A" box="[918,1036,702,728]" italics="true" pageId="197" pageNumber="642">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
(n = 6), but aside from three samples of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEC5CFD0E4218FD4F" box="[638,797,738,765]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="umbratum">
|
||
<emphasis id="B97F3F60CADCFF2FEC5CFD0E4218FD4F" box="[638,797,738,765]" italics="true" pageId="197" pageNumber="642">O. umbratum</emphasis>
|
||
</taxonomicName>
|
||
this group includes one of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEED0FCE54063FC91" box="[242,358,776,803]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="lusinga" status="sp. nov.">
|
||
<emphasis id="B97F3F60CADCFF2FEED0FCE54063FC91" box="[242,358,776,803]" italics="true" pageId="197" pageNumber="642">O. lusinga</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C821BCADCFF2FEF52FCE440CEFC90" box="[368,459,777,802]" pageId="197" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, one of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEC09FCE54398FC91" box="[555,669,776,803]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADCFF2FEC09FCE54398FC91" box="[555,669,776,803]" italics="true" pageId="197" pageNumber="642">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
from central Congo-Kinshasa, and two unverified samples of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEC21FCC34371FCFA" box="[515,628,814,840]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADCFF2FEC21FCC34371FCFA" box="[515,628,814,840]" italics="true" pageId="197" pageNumber="642">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
from Gabon (
|
||
<figureCitation id="1330FFF7CADCFF2FED3BFCC04260FCFA" box="[793,869,813,840]" captionStart="Tree 8" captionStartId="188.[159,207,1438,1463]" captionTargetBox="[159,1034,577,1399]" captionTargetId="figure@188.[156,1038,571,1402]" captionTargetPageId="188" captionText="Tree 8. COI gene tree from Bayesian inference analysis for the saegeri - group of Orthetrum and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages)." pageId="197" pageNumber="642">Tree 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF74B3FACADCFF2FEEBDFA1B426AF9BD" box="[159,879,1526,1551]" pageId="197" pageNumber="642" targetBox="[158,1035,875,1485]" targetPageId="197">
|
||
<paragraph id="8BB4E372CADCFF2FEEBDFA1B426AF9BD" box="[159,879,1526,1551]" pageId="197" pageNumber="642">
|
||
<emphasis id="B97F3F60CADCFF2FEEBDFA1B4041F9BD" bold="true" box="[159,324,1526,1551]" pageId="197" pageNumber="642">
|
||
Type
|
||
<figureCitation id="1330FFF7CADCFF2FEEF8FA1B4045F9BD" box="[218,320,1526,1551]" captionStart="Photo 53" captionStartId="153.[159,225,1465,1490]" captionTargetBox="[159,1035,771,1423]" captionTargetId="figure@153.[159,1038,771,1425]" captionTargetPageId="153" captionText="Photo 53. Paragomphus dispar, male, near Moyabi, Gabon. When flying rapidly low over the water, males are hard to see except for their unique markings. Photo: JK (17 - ix- 2013)" pageId="197" pageNumber="642">Photo 53</figureCitation>
|
||
.
|
||
</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C0B98F1CADCFF2FEFEDFA1B43BEF9BD" box="[463,699,1526,1551]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="197" pageNumber="642" phylum="Arthropoda" rank="species" species="umbratum" status="sp. nov.">
|
||
<emphasis id="B97F3F60CADCFF2FEFEDFA1B43BEF9BD" box="[463,699,1526,1551]" italics="true" pageId="197" pageNumber="642">Orthetrum umbratum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C821BCADCFF2FECE3FA1B4212F9BD" box="[705,791,1526,1551]" pageId="197" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, RMNH.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C311B0F9CADFFF2CEEBDFF1341DEFCFA" pageId="198" pageNumber="643" type="diagnosis">
|
||
<paragraph id="8BB4E372CADFFF2CEEBDFF134317FEAA" box="[159,530,254,280]" pageId="198" pageNumber="643">
|
||
<heading id="D0FC541ECADFFF2CEEBDFF134317FEAA" bold="true" box="[159,530,254,280]" fontSize="11" level="3" pageId="198" pageNumber="643" reason="6">
|
||
<emphasis id="B97F3F60CADFFF2CEEBDFF134317FEAA" bold="true" box="[159,530,254,280]" pageId="198" pageNumber="643">Male morphological diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CADFFF2CEEBDFECF41DEFCFA" pageId="198" pageNumber="643">
|
||
Recalls the sympatric
|
||
<taxonomicName id="4C0B98F1CADFFF2CEF8BFECE4318FE8F" box="[425,541,291,317]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="198" pageNumber="643" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADFFF2CEF8BFECE4318FE8F" box="[425,541,291,317]" italics="true" pageId="198" pageNumber="643">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
by (a) moderate size, Hw 27.5 –31.0 mm (n = 7); (b) the absence of cell-doublings in the radial planate and thus a single row of cells there; (c) all subcostal Ax dark; (d) medium-sized dark brown Pt that are about 11 % of Hw length, 3.0–
|
||
<quantity id="4CF34E97CADFFF2CECE8FE7E4220FE1F" box="[714,805,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="198" pageNumber="643" unit="mm" value="3.4">3.4 mm</quantity>
|
||
; (e) Hw base faintly yellow, but never with a dark brown patch; and (f) abdomen slender, although shorter than Hw. When not pruinose also similar by (g) the dorsa of S 4–7 with pairs of pale subapical spots that lie closer to the dorsal than lateral carinae, while S 8–10 are all dark. However, (1) the hook of the hamule tapers and curves gradually toward a sharp tip that is turned back- and outward, appearing like a claw or talon beside a rather narrow and distinctly higher lobe, with a wide space between the hook and lobe, and thus the overall shape is somewhat intermediate between
|
||
<taxonomicName id="4C0B98F1CADFFF2CECF9FD53423AFD6A" box="[731,831,701,728]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="198" pageNumber="643" phylum="Arthropoda" rank="species" species="hintzi">
|
||
<emphasis id="B97F3F60CADFFF2CECF9FD53423AFD6A" box="[731,831,701,728]" italics="true" pageId="198" pageNumber="643">O. hintzi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B98F1CADFFF2CED5BFD53450EFD6A" box="[889,1035,701,728]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="198" pageNumber="643" phylum="Arthropoda" rank="species" species="machadoi">
|
||
<emphasis id="B97F3F60CADFFF2CED5BFD53450EFD6A" box="[889,1035,701,728]" italics="true" pageId="198" pageNumber="643">O. machadoi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1330FFF7CADFFF2CEE85FD0F41FCFD4F" box="[167,249,738,765]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="198" pageNumber="643">Fig. 31</figureCitation>
|
||
); and (2) the appendages are always wholly black and never (partly) pale in young specimens, although sympatric
|
||
<taxonomicName id="4C0B98F1CADFFF2CECEFFCE54244FC91" box="[717,833,776,803]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="198" pageNumber="643" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADFFF2CECEFFCE54244FC91" box="[717,833,776,803]" italics="true" pageId="198" pageNumber="643">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
can be similarly dark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF74B3FACADFFF2CEEBDFA3741E0F9A2" pageId="198" pageNumber="643" targetBox="[159,1035,877,1457]" targetPageId="198">
|
||
<paragraph id="8BB4E372CADFFF2CEEBDFA3741E0F9A2" pageId="198" pageNumber="643">
|
||
<emphasis id="B97F3F60CADFFF2CEEBDFA374009FA40" bold="true" box="[159,268,1498,1522]" pageId="198" pageNumber="643">Photo 68.</emphasis>
|
||
<taxonomicName id="4C0B98F1CADFFF2CEF37FA374301FA40" box="[277,516,1498,1522]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="198" pageNumber="643" phylum="Arthropoda" rank="species" species="umbratum">
|
||
<emphasis id="B97F3F60CADFFF2CEF37FA374301FA40" box="[277,516,1498,1522]" italics="true" pageId="198" pageNumber="643">Orthetrum umbratum</emphasis>
|
||
</taxonomicName>
|
||
, male; Moukoumbi, Gabon. Photo: NM (10-iv- -2012)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C311B0F9CADEFF2DEEBDFF1340EFFED0" pageId="199" pageNumber="644" type="etymology">
|
||
<paragraph id="8BB4E372CADEFF2DEEBDFF134021FEAA" box="[159,292,254,280]" pageId="199" pageNumber="644">
|
||
<heading id="D0FC541ECADEFF2DEEBDFF134021FEAA" bold="true" box="[159,292,254,280]" fontSize="11" level="3" pageId="199" pageNumber="644" reason="6">
|
||
<emphasis id="B97F3F60CADEFF2DEEBDFF134021FEAA" bold="true" box="[159,292,254,280]" pageId="199" pageNumber="644">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CADEFF2DEEBDFECF40EFFED0" pageId="199" pageNumber="644">Latin “shaded” refers to the species’ deep forest habitat, which is unusual for the genus (neuter adjective).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311B0F9CADEFF2DEEBDFE794380FDDA" pageId="199" pageNumber="644" type="distribution">
|
||
<paragraph id="8BB4E372CADEFF2DEEBDFE794081FE1F" box="[159,388,404,429]" pageId="199" pageNumber="644">
|
||
<heading id="D0FC541ECADEFF2DEEBDFE794081FE1F" bold="true" box="[159,388,404,429]" fontSize="11" level="3" pageId="199" pageNumber="644" reason="6">
|
||
<emphasis id="B97F3F60CADEFF2DEEBDFE794081FE1F" bold="true" box="[159,388,404,429]" pageId="199" pageNumber="644">Range and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CADEFF2DEEBDFE554380FDDA" pageId="199" pageNumber="644">
|
||
Replaces
|
||
<taxonomicName id="4C0B98F1CADEFF2DEF32FE554081FE61" box="[272,388,440,467]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADEFF2DEF32FE554081FE61" box="[272,388,440,467]" italics="true" pageId="199" pageNumber="644">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
in south-eastern Gabon at the darkest and least disturbed rainforest pools with very much detritus, recorded at relatively high- er elevation between 530 and
|
||
<quantity id="4CF34E97CADEFF2DEC23FDEE4349FDAF" box="[513,588,515,541]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="199" pageNumber="644" unit="m" value="580.0">580 m</quantity>
|
||
a.s.l (
|
||
<figureCitation id="1330FFF7CADEFF2DECB3FDEF43EBFDAF" box="[657,750,514,541]" captionStart="Map 11" captionStartId="192.[159,210,1496,1521]" captionTargetBox="[158,1035,505,1454]" captionTargetId="figure@192.[156,1038,505,1458]" captionTargetPageId="192" captionText="Map 11. Distribution of the saegeri - group of Orthetrum (O. agaricum sp. nov., O. kafwi sp. nov., O. lusinga, O. umbratum sp. nov. and O. saegeri)." pageId="199" pageNumber="644">Map 11</figureCitation>
|
||
). While
|
||
<taxonomicName id="4C0B98F1CADEFF2DED77FDEE42C2FDAF" box="[853,967,515,541]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="saegeri">
|
||
<emphasis id="B97F3F60CADEFF2DED77FDEE42C2FDAF" box="[853,967,515,541]" italics="true" pageId="199" pageNumber="644">O. saegeri</emphasis>
|
||
</taxonomicName>
|
||
often basks at the forest edge,
|
||
<taxonomicName id="4C0B98F1CADEFF2DEF96FDC5434AFDF0" box="[436,591,552,578]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="umbratum">
|
||
<emphasis id="B97F3F60CADEFF2DEF96FDC5434AFDF0" box="[436,591,552,578]" italics="true" pageId="199" pageNumber="644">O.umbratum</emphasis>
|
||
</taxonomicName>
|
||
only comes down to the ponds during bright sun in the two hours around noon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |