309 lines
33 KiB
XML
309 lines
33 KiB
XML
<document id="26242CE086FDBCC777C0DDEDAE15D2C3" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas" docDate="2015" docId="03A25264CAC1FF36EF3AFF10406BFE1F" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Zygonyx annika Dijkstra, sp. nov." docType="treatment" docVersion="21" lastPageNumber="665" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="661" updateTime="1695001031402" updateUser="admin">
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<mods:titleInfo id="6F4FDBABA3A2D7AE188958C63490CF12">
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<mods:title id="BB19F5996C6449489DACD5960738DBEA">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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<mods:roleTerm id="8BC4B1CC04B80F8C1B5DF36662AACF06">Author</mods:roleTerm>
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<mods:namePart id="4525F6371A467178BBE5262E073E7614">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="207F1A6CD15FD28613CD2B62C598CC48">Kipping, Jens</mods:namePart>
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<mods:name id="740B238DCE34956A6C38096D65EE2449" type="personal">
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<mods:roleTerm id="728049B870647CAFCBCE032B594253C6">Author</mods:roleTerm>
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<mods:namePart id="7ACDD8F93EE877FFB9A5B210DAD8B7C7">Mézière, Nicolas</mods:namePart>
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<mods:titleInfo id="B1E60EEE87099D693E2FA1739CDD5047">
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<mods:title id="5FBACC04E874480A5686CDC8C1009542">Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part id="8890A5AE60FA76BAAB895CA23995A9BE">
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<mods:date id="6ED734184DA49F74AEC3E4440DB7545A">2015</mods:date>
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<mods:number id="9F5CD4CB7E1AEEB4D2DF3E26D6CF5AC4">44</mods:number>
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<mods:number id="FF0246EF82C446D5B0A31BB6F9D56EF2">4</mods:number>
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<mods:identifier id="A6DC0B038A35370676D0D9681C746DCC" type="DOI">10.5281/zenodo.35388</mods:identifier>
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<treatment id="03A25264CAC1FF36EF3AFF10406BFE1F" ID-DOI="http://doi.org/10.5281/zenodo.5640308" ID-GBIF-Taxon="114923470" ID-Zenodo-Dep="5640308" LSID="urn:lsid:zoobank.org:act:03A25264-CAC1-FF36-EF3A-FF10406BFE1F" httpUri="http://treatment.plazi.org/id/03A25264CAC1FF36EF3AFF10406BFE1F" lastPageId="220" lastPageNumber="665" pageId="216" pageNumber="661">
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<subSubSection id="C311B0F9CAC1FF32EF3AFF10421BFEF1" pageId="216" pageNumber="661" type="nomenclature">
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<paragraph id="8BB4E372CAC1FF32EF3AFF10421BFEF1" pageId="216" pageNumber="661">
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<heading id="D0FC541ECAC1FF32EF3AFF10421BFEF1" bold="true" centered="true" fontSize="11" level="2" pageId="216" pageNumber="661" reason="4">
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<emphasis id="B97F3F60CAC1FF32EF3AFF104297FEAA" bold="true" box="[280,914,251,280]" pageId="216" pageNumber="661">
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<taxonomicName id="4C0B98F1CAC1FF32EF3AFF10433CFEAA" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CAC1-FF36-EF3A-FF10406BFE1F" authority="Dijkstra" authorityName="Dijkstra" box="[280,569,253,280]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="annika" status="sp. nov.">
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<emphasis id="B97F3F60CAC1FF32EF3AFF1040D0FEAA" bold="true" box="[280,469,253,280]" italics="true" pageId="216" pageNumber="661">Zygonyx annika</emphasis>
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Dijkstra
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</taxonomicName>
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<taxonomicNameLabel id="A24C821BCAC1FF32EC62FF104398FEAA" box="[576,669,253,280]" pageId="216" pageNumber="661" rank="species">sp. nov.</taxonomicNameLabel>
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– Annika’s Cascader
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</emphasis>
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||
(
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<typeStatus id="54B05DD0CAC1FF32EFB1FECA40CBFEF1" box="[403,462,295,323]" pageId="216" pageNumber="661">Type</typeStatus>
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<figureCitation id="1330FFF7CAC1FF32EFF6FECA4341FEF0" box="[468,580,295,322]" captionStart="Photo 58" captionStartId="169.[159,225,1500,1525]" captionTargetBox="[158,1035,891,1460]" captionTargetId="figure@169.[156,1038,888,1462]" captionTargetPageId="169" captionText="Photo 58. Eleuthemis eogaster, male; Lumanie River near Uíge, Angola. Photo: KDBD (03 - x- 2013)" pageId="216" pageNumber="661">Photo 58</figureCitation>
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,
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<figureCitation id="1330FFF7CAC1FF32EC6CFECA43B8FEF0" box="[590,701,295,322]" captionStart="Photo 75" captionStartId="218.[159,225,1322,1347]" captionText="Photo 75. Rocky stretch of the Moa River at Tiwai Island, Sierra Leone. Type locality of Zygonyx annika and habitat of Pseudagrion pacale. Photo: KDBD (14 - iii- 2011)" pageId="216" pageNumber="661">Photo 75</figureCitation>
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,
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<figureCitation id="1330FFF7CAC1FF32ECE5FECA4212FEF1" box="[711,791,295,323]" captionStart="Figure 35" captionStartId="219.[744,812,1322,1347]" captionTargetBox="[158,706,638,1549]" captionTargetId="figure@219.[156,709,634,1552]" captionTargetPageId="219" captionText="Figure 35. Secondary genitalia (left) and male appendages (right) in lateral view of the flavicosta - complex of Zygonyx; illustrations of Z. flavicosta are based on a specimen from Sierra Leone." pageId="216" pageNumber="661">Fig. 35</figureCitation>
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)
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</heading>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C311B0F9CAC1FF32EEBDFEBB401AFEC2" box="[159,287,342,368]" pageId="216" pageNumber="661" type="description">
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||
<paragraph id="8BB4E372CAC1FF32EEBDFEBB401AFEC2" box="[159,287,342,368]" pageId="216" pageNumber="661">
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||
<heading id="D0FC541ECAC1FF32EEBDFEBB401AFEC2" bold="true" box="[159,287,342,368]" fontSize="11" level="3" pageId="216" pageNumber="661" reason="6">
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<emphasis id="B97F3F60CAC1FF32EEBDFEBB401AFEC2" bold="true" box="[159,287,342,368]" pageId="216" pageNumber="661">Taxonomy</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C311B0F9CAC1FF32EEBDFE9740C0FD57" pageId="216" pageNumber="661" type="discussion">
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<paragraph id="8BB4E372CAC1FF32EEBDFE9740C0FD57" pageId="216" pageNumber="661">
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Variation in markings and male genitalia suggests that a complex exists of species similar to
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<taxonomicName id="4C0B98F1CAC1FF32EF49FE4C43B7FE08" authority="Sjostedt, 1900" authorityName="Sjostedt" authorityYear="1900" box="[363,690,416,442]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
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<emphasis id="B97F3F60CAC1FF32EF49FE4C40F0FE08" box="[363,501,416,442]" italics="true" pageId="216" pageNumber="661">Z. flavicosta</emphasis>
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(Sjöstedt, 1900)
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</taxonomicName>
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, but thus far only
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<taxonomicName id="4C0B98F1CAC1FF32EDA2FE4C4070FE52" authority="Legrand, 1997" authorityName="Legrand" authorityYear="1997" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="geminuncus">
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<emphasis id="B97F3F60CAC1FF32EDA2FE4C41C1FE52" italics="true" pageId="216" pageNumber="661">Z. geminuncus</emphasis>
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Legrand, 1997
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</taxonomicName>
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has been separated. Morphology and COI data suggest that at least six species are involved, with two or even three occurring in close proximity. Variation among specimens nearest to the true
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<taxonomicName id="4C0B98F1CAC1FF32EDA0FDFC450EFD98" box="[898,1035,528,554]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
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<emphasis id="B97F3F60CAC1FF32EDA0FDFC450EFD98" box="[898,1035,528,554]" italics="true" pageId="216" pageNumber="661">Z. flavicosta</emphasis>
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</taxonomicName>
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is so great that we only name three species that can be separated from sympatric relatives by both genetics and morphology (
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<figureCitation id="1330FFF7CAC1FF32ECC0FDB74239FDC7" box="[738,828,602,629]" captionStart="Tree 10" captionStartId="217.[159,207,1437,1461]" captionTargetBox="[159,1032,380,1396]" captionTargetId="figure@217.[156,1038,378,1399]" captionTargetPageId="217" captionText="Tree 10. COI gene tree from Bayesian inference analysis for the flavicosta - complex of Zygonyx and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages)." pageId="216" pageNumber="661">Tree 10</figureCitation>
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). The present taxon is known only from a pair caught in copula, which overlaps geographically with morphologically and genetically distinct populations that were identified as
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<taxonomicName id="4C0B98F1CAC1FF32EF16FD2640BBFD57" box="[308,446,714,741]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
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<emphasis id="B97F3F60CAC1FF32EF16FD2640BBFD57" box="[308,446,714,741]" italics="true" pageId="216" pageNumber="661">Z. flavicosta</emphasis>
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</taxonomicName>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C311B0F9CAC1FF32EEBDFCF343C7FB8F" pageId="216" pageNumber="661" type="materials_examined">
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||
<paragraph id="8BB4E372CAC1FF32EEBDFCF3406DFC8A" box="[159,360,798,824]" pageId="216" pageNumber="661">
|
||
<heading id="D0FC541ECAC1FF32EEBDFCF3406DFC8A" bold="true" box="[159,360,798,824]" fontSize="11" level="3" pageId="216" pageNumber="661" reason="6">
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||
<emphasis id="B97F3F60CAC1FF32EEBDFCF3406DFC8A" bold="true" box="[159,360,798,824]" pageId="216" pageNumber="661">Material studied</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BB4E372CAC1FF32EEBDFCAE433EFC40" pageId="216" pageNumber="661">
|
||
<materialsCitation id="3B63E92FCAC1FF32EEBDFCAE40C6FC7F" ID-GBIF-Occurrence="3396361344" collectingDate="2011-03-14" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Sierra Leone" elevation="122 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF252-15" latitude="7.551" location="SW of Kenema, Moa River at Tiwai Island" longLatPrecision="55" longitude="-11.3549" pageId="216" pageNumber="661" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Province" typeStatus="holotype">
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||
<emphasis id="B97F3F60CAC1FF32EEBDFCAE4015FCEF" bold="true" box="[159,272,835,861]" pageId="216" pageNumber="661">
|
||
<typeStatus id="54B05DD0CAC1FF32EEBDFCAE4015FCEF" box="[159,272,835,861]" pageId="216" pageNumber="661">Holotype</typeStatus>
|
||
</emphasis>
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||
<emphasis id="B97F3F60CAC1FF32EF39FCAF402AFCEE" bold="true" box="[283,303,834,860]" pageId="216" pageNumber="661">
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||
<specimenCount id="AD2E67A70A751F9ACF934DBC99DA45AE" type="male">♂</specimenCount>
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||
</emphasis>
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||
<emphasis id="B97F3F60CAC1FF32EF0DFCAE4033FCEF" bold="true" box="[303,310,835,861]" pageId="216" pageNumber="661">.</emphasis>
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<specimenCode id="6E321A6E359314EC1D68E2FF08762516">RMNH.INS.503187</specimenCode>
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,
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<collectingCountry id="F31CA3E2CAC1FF32EC18FCAF43D6FCEF" box="[570,723,834,861]" name="Sierra Leone" pageId="216" pageNumber="661" value="Sierra Leone">Sierra Leone</collectingCountry>
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||
,
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<collectingRegion id="49CF2D90CAC1FF32ECC0FCAE42AAFCEF" box="[738,943,834,861]" pageId="216" pageNumber="661">Eastern Province</collectingRegion>
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||
,
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<quantity id="4CF34E97CAC1FF32ED9CFCAE4509FCEF" box="[958,1036,834,861]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="216" pageNumber="661" unit="km" value="40.0">
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<locationDeviation id="27AC539994967B7AC68212A170825C86" box="[958,1036,834,861]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="216" pageNumber="661" unit="km" value="40.0">40 km</locationDeviation>
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</quantity>
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<location id="8F3C344E8A91F753863710D39674D830" LSID="urn:lsid:plazi:treatment:03A25264CAC1FF36EF3AFF10406BFE1F:8F3C344E8A91F753863710D39674D830" country="Sierra Leone" latitude="7.551" longLatPrecision="55" longitude="-11.3549" name="SW of Kenema, Moa River at Tiwai Island" stateProvince="Eastern Province">SW of Kenema, Moa River at Tiwai Island</location>
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, forested island in large river with rocky rapids (
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||
<figureCitation id="1330FFF7CAC1FF32EF1DFC6040A8FC1A" box="[319,429,909,936]" captionStart="Photo 75" captionStartId="218.[159,225,1322,1347]" captionText="Photo 75. Rocky stretch of the Moa River at Tiwai Island, Sierra Leone. Type locality of Zygonyx annika and habitat of Pseudagrion pacale. Photo: KDBD (14 - iii- 2011)" pageId="216" pageNumber="661">Photo 75</figureCitation>
|
||
),
|
||
<quantity id="4CF34E97CAC1FF32EFE0FC60430EFC1A" box="[450,523,909,936]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="216" pageNumber="661" unit="m" value="122.0">
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<elevation id="28970EA5BACB023901BFD9ADED77AB7D" box="[450,523,909,936]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="216" pageNumber="661" unit="m" value="122.0">122 m</elevation>
|
||
</quantity>
|
||
a.s.l. (
|
||
<geoCoordinate id="EE3F85B5CAC1FF32EC76FC6043B1FC1A" box="[596,692,909,936]" direction="north" orientation="latitude" pageId="216" pageNumber="661" precision="55" value="7.551">7.551 ° N</geoCoordinate>
|
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<geoCoordinate id="EE3F85B5CAC1FF32EC99FC604244FC1A" box="[699,833,909,936]" direction="west" orientation="longitude" pageId="216" pageNumber="661" precision="5" value="-11.3549">11.3549 ° W</geoCoordinate>
|
||
),
|
||
<date id="FFB5C5B2CAC1FF32ED77FC6042DDFC1A" box="[853,984,909,936]" pageId="216" pageNumber="661" value="2011-03-14">
|
||
<collectingDate id="0EEC0F5FAED7DA4A48C89F1CFD478D96" box="[853,984,909,936]" pageId="216" pageNumber="661" value="2011-03-14">14 -iii- 2011</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="27DCFD496B75ADA4DE1CCF023D68BD59">K.-D.B. Dijkstra</collectorName>
|
||
,
|
||
<collectionCode id="ED1A7BB7CAC1FF32EF45FC5E40C6FC7F" box="[359,451,947,973]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="216" pageNumber="661">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
. The right Hw and leg are aberrant in shape, which is probably a developmental defect.
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CAC1FF32EEBDFC1343C7FB8F" pageId="216" pageNumber="661">
|
||
<emphasis id="B97F3F60CAC1FF32EEBDFC134078FBAA" bold="true" box="[159,381,1022,1048]" pageId="216" pageNumber="661">Further material.</emphasis>
|
||
<materialsCitation id="3B63E92FCAC1FF32EFACFC1043BBFB8F" ID-GBIF-Occurrence="3396361345" collectingDate="2011-03-14" collectionCode="RMNH" country="Sierra Leone" elevation="122 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF253-15" latitude="7.551" location="SW of Kenema, Moa River at Tiwai Island" longitude="-11.35549" pageId="216" pageNumber="661" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Province">
|
||
<collectingCountry id="F31CA3E2CAC1FF32EFACFC10435BFBAA" box="[398,606,1021,1048]" name="Sierra Leone" pageId="216" pageNumber="661" value="Sierra Leone">SIERRA LEONE</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CF2D90CAC1FF32EC5AFC104254FBAA" box="[632,849,1021,1048]" pageId="216" pageNumber="661">Eastern Province</collectingRegion>
|
||
):
|
||
<specimenCount id="4D11E35C515827EB7557268F62A911C4" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="395EBA8BE6AEC6D9D2BB87C85BFF3BD4">RMNH.INS.503188</specimenCode>
|
||
), in copula with holotype,
|
||
<collectionCode id="ED1A7BB7CAC1FF32EC40FBCE43BBFB8F" box="[610,702,1059,1085]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="216" pageNumber="661">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311B0F9CAC1FF32EEBDFB9B42D2FB68" pageId="216" pageNumber="661" type="description">
|
||
<paragraph id="8BB4E372CAC1FF32EEBDFB9B400CFB22" box="[159,265,1142,1168]" pageId="216" pageNumber="661">
|
||
<heading id="D0FC541ECAC1FF32EEBDFB9B400CFB22" bold="true" box="[159,265,1142,1168]" fontSize="11" level="3" pageId="216" pageNumber="661" reason="6">
|
||
<emphasis id="B97F3F60CAC1FF32EEBDFB9B400CFB22" bold="true" box="[159,265,1142,1168]" pageId="216" pageNumber="661">Genetics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CAC1FF32EEBDFB7642D2FB68" pageId="216" pageNumber="661">
|
||
One unique haplotype (n = 2) is distinct and nearer to
|
||
<taxonomicName id="4C0B98F1CAC1FF32ED33FB764278FB07" box="[785,893,1178,1205]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="dionyx" status="sp. nov.">
|
||
<emphasis id="B97F3F60CAC1FF32ED33FB764278FB07" box="[785,893,1178,1205]" italics="true" pageId="216" pageNumber="661">Z. dionyx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C821BCAC1FF32EDA1FB7642DFFB07" box="[899,986,1179,1205]" pageId="216" pageNumber="661" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C0B98F1CAC1FF32EEBDFB2C4043FB68" box="[159,326,1217,1242]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="geminuncus">
|
||
<emphasis id="B97F3F60CAC1FF32EEBDFB2C4043FB68" box="[159,326,1217,1242]" italics="true" pageId="216" pageNumber="661">Z. geminuncus</emphasis>
|
||
</taxonomicName>
|
||
than the morphologically similar
|
||
<taxonomicName id="4C0B98F1CAC1FF32ECF7FB2C425AFB68" box="[725,863,1216,1242]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC1FF32ECF7FB2C425AFB68" box="[725,863,1216,1242]" italics="true" pageId="216" pageNumber="661">Z. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1330FFF7CAC1FF32ED4CFB2D42CCFB68" box="[878,969,1216,1242]" captionStart="Tree 10" captionStartId="217.[159,207,1437,1461]" captionTargetBox="[159,1032,380,1396]" captionTargetId="figure@217.[156,1038,378,1399]" captionTargetPageId="217" captionText="Tree 10. COI gene tree from Bayesian inference analysis for the flavicosta - complex of Zygonyx and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages)." pageId="216" pageNumber="661">Tree 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311B0F9CAC1FF31EEBDFAFE4213FE1F" lastPageId="219" lastPageNumber="664" pageId="216" pageNumber="661" type="diagnosis">
|
||
<paragraph id="8BB4E372CAC1FF32EEBDFAFE4317FA9F" box="[159,530,1299,1325]" pageId="216" pageNumber="661">
|
||
<heading id="D0FC541ECAC1FF32EEBDFAFE4317FA9F" bold="true" box="[159,530,1299,1325]" fontSize="11" level="3" pageId="216" pageNumber="661" reason="6">
|
||
<emphasis id="B97F3F60CAC1FF32EEBDFAFE4317FA9F" bold="true" box="[159,530,1299,1325]" pageId="216" pageNumber="661">Male morphological diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CAC1FF31EEBDFAD54213FE1F" lastPageId="219" lastPageNumber="664" pageId="216" pageNumber="661">
|
||
Belongs to the
|
||
<taxonomicName id="4C0B98F1CAC1FF32EF7EFAD540CDFAE0" box="[348,456,1336,1362]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC1FF32EF7EFAD540CDFAE0" box="[348,456,1336,1362]" italics="true" pageId="216" pageNumber="661">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
-complex by (a) the modest size, Hw
|
||
<quantity id="4CF34E97CAC1FF32EDBDFAD54509FAE0" box="[927,1036,1336,1362]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="216" pageNumber="661" unit="mm" value="36.5">36.5 mm</quantity>
|
||
(n = 1); (b) the moderately dense venation, with 13 ½– 14 Ax in Fw; (c) the largely black hind legs; (d) the pointed and forward-curved apex of the anterior lamina (
|
||
<figureCitation id="1330FFF7CAC1FF32EF69FA4540A5FA70" box="[331,416,1448,1474]" captionStart="Figure 35" captionStartId="219.[744,812,1322,1347]" captionTargetBox="[158,706,638,1549]" captionTargetId="figure@219.[156,709,634,1552]" captionTargetPageId="219" captionText="Figure 35. Secondary genitalia (left) and male appendages (right) in lateral view of the flavicosta - complex of Zygonyx; illustrations of Z. flavicosta are based on a specimen from Sierra Leone." pageId="216" pageNumber="661">Fig. 35</figureCitation>
|
||
); and (e) the pale dorsal band on S 7 that stands out against the largely dark S 5–6. Differs from the sympatric
|
||
<taxonomicName id="4C0B98F1CAC1FF32ED75FA2342E0FA5A" box="[855,997,1485,1512]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="216" pageNumber="661" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC1FF32ED75FA2342E0FA5A" box="[855,997,1485,1512]" italics="true" pageId="216" pageNumber="661">Z. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
by (1)the dorsum of the eyes being deep reddish brown, contrasting with their greyish underside, rather than uniformly grey-blue; and (2) the shorter hook of hamule, which is less than a third as long as the hamule is wide (
|
||
<figureCitation id="1330FFF7CAC0FF33EE8AFEA541FFFED0" box="[168,250,328,354]" captionStart="Figure 35" captionStartId="219.[744,812,1322,1347]" captionTargetBox="[158,706,638,1549]" captionTargetId="figure@219.[156,709,634,1552]" captionTargetPageId="219" captionText="Figure 35. Secondary genitalia (left) and male appendages (right) in lateral view of the flavicosta - complex of Zygonyx; illustrations of Z. flavicosta are based on a specimen from Sierra Leone." pageId="217" pageNumber="662">Fig. 35</figureCitation>
|
||
). Also differs from
|
||
<taxonomicName id="4C0B98F1CAC0FF33EFFAFEA44364FED1" box="[472,609,328,355]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="217" pageNumber="662" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC0FF33EFFAFEA44364FED1" box="[472,609,328,355]" italics="true" pageId="217" pageNumber="662">Z. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
from Sierra Leone, Liberia and Gha- na by (3) the Hw membrane being darkly stained and not clear beside the membranule; (4) the more continuous pale thoracic markings, the stripe on the metepisternum not being broken between the metastigma and wing base, while that on the metepimeron is not invaded by black posteriorly to appear T-shaped; and (5) the straighter cerci (
|
||
<figureCitation id="1330FFF7CAC2FF31EC96FE7F420DFE1F" box="[692,776,402,429]" captionStart="Figure 35" captionStartId="219.[744,812,1322,1347]" captionTargetBox="[158,706,638,1549]" captionTargetId="figure@219.[156,709,634,1552]" captionTargetPageId="219" captionText="Figure 35. Secondary genitalia (left) and male appendages (right) in lateral view of the flavicosta - complex of Zygonyx; illustrations of Z. flavicosta are based on a specimen from Sierra Leone." pageId="219" pageNumber="664">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF74B3FACAC0FF33EEBDFA704288F9BC" pageId="217" pageNumber="662" targetBox="[159,1032,380,1396]" targetPageId="217">
|
||
<paragraph id="8BB4E372CAC0FF33EEBDFA704288F9BC" pageId="217" pageNumber="662">
|
||
<emphasis id="B97F3F60CAC0FF33EEBDFA7041F3FA07" bold="true" box="[159,246,1437,1461]" pageId="217" pageNumber="662">Tree 10.</emphasis>
|
||
COI gene tree from Bayesian inference analysis for the
|
||
<taxonomicName id="4C0B98F1CAC0FF33ED60FA7042ADFA07" box="[834,936,1437,1461]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="217" pageNumber="662" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC0FF33ED60FA7042ADFA07" box="[834,936,1437,1461]" italics="true" pageId="217" pageNumber="662">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
-complex of
|
||
<taxonomicName id="4C0B98F1CAC0FF33EE9FFA574010FA61" box="[189,277,1466,1491]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="217" pageNumber="662" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97F3F60CAC0FF33EE9FFA574010FA61" box="[189,277,1466,1491]" italics="true" pageId="217" pageNumber="662">Zygonyx</emphasis>
|
||
</taxonomicName>
|
||
and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50% majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF74B3FACAC3FF30EEBDFBDE403FFBDB" pageId="218" pageNumber="663" targetBox="[158,1035,249,1035]" targetPageId="218">
|
||
<paragraph id="8BB4E372CAC3FF30EEBDFBDE403FFBDB" pageId="218" pageNumber="663">
|
||
<emphasis id="B97F3F60CAC3FF30EEBDFBDE4047FBF9" bold="true" box="[159,322,1075,1100]" pageId="218" pageNumber="663">
|
||
Type
|
||
<figureCitation id="1330FFF7CAC3FF30EEFBFBDE403BFBF9" box="[217,318,1075,1099]" captionStart="Photo 58" captionStartId="169.[159,225,1500,1525]" captionTargetBox="[158,1035,891,1460]" captionTargetId="figure@169.[156,1038,888,1462]" captionTargetPageId="169" captionText="Photo 58. Eleuthemis eogaster, male; Lumanie River near Uíge, Angola. Photo: KDBD (03 - x- 2013)" pageId="218" pageNumber="663">Photo 58</figureCitation>
|
||
.
|
||
</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C0B98F1CAC3FF30EFE9FBDE4374FBF9" box="[459,625,1075,1099]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="218" pageNumber="663" phylum="Arthropoda" rank="species" species="annika" status="sp. nov.">
|
||
<emphasis id="B97F3F60CAC3FF30EFE9FBDE4374FBF9" box="[459,625,1075,1099]" italics="true" pageId="218" pageNumber="663">Zygonyx annika</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C821BCAC3FF30EC54FBDE43C9FBF9" box="[630,716,1075,1099]" pageId="218" pageNumber="663" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, RMNH (horizontally mirrored from original).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF74B3FACAC3FF30EEBDFAC74026F9A3" pageId="218" pageNumber="663">
|
||
<paragraph id="8BB4E372CAC3FF30EEBDFAC74026F9A3" pageId="218" pageNumber="663">
|
||
<emphasis id="B97F3F60CAC3FF30EEBDFAC7400DFAF1" bold="true" box="[159,264,1322,1347]" pageId="218" pageNumber="663">Photo 75.</emphasis>
|
||
Rocky stretch of the Moa River at Tiwai Island, Sierra Leone. Type locality of
|
||
<taxonomicName id="4C0B98F1CAC3FF30EEBDFA4D405FFA0A" box="[159,346,1440,1464]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="218" pageNumber="663" phylum="Arthropoda" rank="species" species="annika">
|
||
<emphasis id="B97F3F60CAC3FF30EEBDFA4D405FFA0A" box="[159,346,1440,1464]" italics="true" pageId="218" pageNumber="663">Zygonyx annika</emphasis>
|
||
</taxonomicName>
|
||
and habitat of
|
||
<taxonomicName id="4C0B98F1CAC3FF30EF34FA5041E3FA40" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="218" pageNumber="663" phylum="Arthropoda" rank="species" species="pacale">
|
||
<emphasis id="B97F3F60CAC3FF30EF34FA5041E3FA40" italics="true" pageId="218" pageNumber="663">Pseudagrion pacale</emphasis>
|
||
</taxonomicName>
|
||
. Photo: KDBD (14-iii-2011)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C311B0F9CAC2FF31EEBDFE33409FFDDA" pageId="219" pageNumber="664" type="etymology">
|
||
<paragraph id="8BB4E372CAC2FF31EEBDFE334021FE4A" box="[159,292,478,504]" pageId="219" pageNumber="664">
|
||
<heading id="D0FC541ECAC2FF31EEBDFE334021FE4A" bold="true" box="[159,292,478,504]" fontSize="11" level="3" pageId="219" pageNumber="664" reason="6">
|
||
<emphasis id="B97F3F60CAC2FF31EEBDFE334021FE4A" bold="true" box="[159,292,478,504]" pageId="219" pageNumber="664">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CAC2FF31EEBDFDEF409FFDDA" pageId="219" pageNumber="664">
|
||
Named in honour of Dr
|
||
<taxonomicName id="4C0B98F1CAC2FF31EFF6FDEF432EFDAF" box="[468,555,514,541]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="219" pageNumber="670" phylum="Arthropoda" rank="species" species="annika">Annika</taxonomicName>
|
||
Hillers, who advanced research on the Gola rainforest ecosystem and enabled the author to make his contribution (noun in apposition).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF74B3FACAC2FF31ECCAFAC74273F9A2" pageId="219" pageNumber="664" targetBox="[158,706,638,1549]" targetPageId="219">
|
||
<paragraph id="8BB4E372CAC2FF31ECCAFAC74273F9A2" pageId="219" pageNumber="664">
|
||
<emphasis id="B97F3F60CAC2FF31ECCAFAC74253FAF0" bold="true" box="[744,854,1322,1347]" pageId="219" pageNumber="664">Figure 35.</emphasis>
|
||
Secondary genitalia (left) and male appendages (right) in lateral view of the
|
||
<taxonomicName id="4C0B98F1CAC2FF31ED44FA6F42C8FA28" box="[870,973,1410,1434]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="219" pageNumber="664" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC2FF31ED44FA6F42C8FA28" box="[870,973,1410,1434]" italics="true" pageId="219" pageNumber="664">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
-complex of
|
||
<taxonomicName id="4C0B98F1CAC2FF31ED69FA4D42A6FA0A" box="[843,931,1440,1464]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="219" pageNumber="664" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97F3F60CAC2FF31ED69FA4D42A6FA0A" box="[843,931,1440,1464]" italics="true" pageId="219" pageNumber="664">Zygonyx</emphasis>
|
||
</taxonomicName>
|
||
; illustrations of
|
||
<taxonomicName id="4C0B98F1CAC2FF31ED73FA5042DFFA67" box="[849,986,1469,1493]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="219" pageNumber="664" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC2FF31ED73FA5042DFFA67" box="[849,986,1469,1493]" italics="true" pageId="219" pageNumber="664">Z. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
are based on a specimen from Sierra Leone.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C311B0F9CAC5FF36EEBDFF13406BFE1F" pageId="220" pageNumber="665" type="distribution">
|
||
<paragraph id="8BB4E372CAC5FF36EEBDFF134081FEAA" box="[159,388,254,280]" pageId="220" pageNumber="665">
|
||
<heading id="D0FC541ECAC5FF36EEBDFF134081FEAA" bold="true" box="[159,388,254,280]" fontSize="11" level="3" pageId="220" pageNumber="665" reason="6">
|
||
<emphasis id="B97F3F60CAC5FF36EEBDFF134081FEAA" bold="true" box="[159,388,254,280]" pageId="220" pageNumber="665">Range and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB4E372CAC5FF36EEBDFECF406BFE1F" pageId="220" pageNumber="665">
|
||
Rocky fast-flowing section of the large Moa River in Sierra Leone, flanked by gallery forest and about
|
||
<quantity id="4CF34E97CAC5FF36EFC3FEA54329FED0" box="[481,556,328,354]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="220" pageNumber="665" unit="m" value="120.0">120 m</quantity>
|
||
a.s.l. Similar adults and possible exuviae were seen near Kenema
|
||
<quantity id="4CF34E97CAC5FF36EF9EFE834303FE3A" box="[444,518,365,392]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="220" pageNumber="665" unit="km" value="35.0">35 km</quantity>
|
||
upstream, but
|
||
<taxonomicName id="4C0B98F1CAC5FF36EC9AFE834246FE3A" box="[696,835,365,392]" class="Insecta" family="Libellulidae" genus="Zygonyx" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="220" pageNumber="665" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B97F3F60CAC5FF36EC9AFE834246FE3A" box="[696,835,365,392]" italics="true" pageId="220" pageNumber="665">Z. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
is sympatric and possibly syntopic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |