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<document id="FFACBC8FBC07BC7933648D598E4C9617" ID="10.11646/zootaxa.4268.3.6SLASH11064" ID-CLB-Dataset="33032" ID-DOI="10.11646/zootaxa.4268.3.6" ID-GBIF-Dataset="aaeff7f4-9d74-4834-ad84-54045b3ab444" ID-ISSN="1175-5326" ID-Zenodo-Dep="580536" ID-ZooBank="6532AE05-DB08-4742-BEDF-DE0CB81B1F64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495010266095" checkinUser="plazi" docAuthor="Zanol, Joana, Da Silva, Thauane Dos S. C. &amp; Hutchings, Pat" docDate="2017" docId="CC4187C9716F3B32FF4D15FD239118FE" docLanguage="en" docName="zootaxa.4268.3.6SLASH11064.pdf" docOrigin="Zootaxa 4268 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marphysa pseudosessiloa Zanol, Silva &amp; Hutchings, 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="420" masterDocId="3078FFB171693B3BFFDA172A21601F10" masterDocTitle="One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia" masterLastPageNumber="426" masterPageNumber="411" pageNumber="417" updateTime="1698439664845" updateUser="ExternalLinkService">
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<mods:title id="6DA2C52DE11A03D30642FDE3BFBA15A9">One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia</mods:title>
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<mods:name id="5D9E416F803B73AB834A2E3B23426FF0" type="personal">
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<mods:namePart id="B78844052DE612D7342BD9E89065E213">Zanol, Joana</mods:namePart>
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<mods:namePart id="8A99EB44F6897DDB0539A470BF6C617A">Da Silva, Thauane Dos S. C.</mods:namePart>
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<mods:namePart id="4CBF03636D04E00F3F67AF9F353DB4B0">Hutchings, Pat</mods:namePart>
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<mods:title id="C36E2BC3BFB0F9B4126F87F13A2EEC59">Zootaxa</mods:title>
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<mods:date id="D06FD789DE6A65BB94B1B7FAB55B968D">2017</mods:date>
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<mods:identifier id="0C2E10B87220DDEDD1ECC013AF22624B" type="DOI">10.11646/zootaxa.4268.3.6</mods:identifier>
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<treatment id="CC4187C9716F3B32FF4D15FD239118FE" ID-DOI="http://doi.org/10.5281/zenodo.6039585" ID-GBIF-Taxon="130810149" ID-Zenodo-Dep="6039585" LSID="urn:lsid:plazi:treatment:CC4187C9716F3B32FF4D15FD239118FE" httpUri="http://treatment.plazi.org/id/CC4187C9716F3B32FF4D15FD239118FE" lastPageId="9" lastPageNumber="420" pageId="6" pageNumber="417">
<subSubSection id="0CF26554716F3B3DFF4D15FD206A1C03" pageId="6" pageNumber="417" type="nomenclature">
<paragraph id="445736DF716F3B3DFF4D15FD23731DE2" blockId="6.[151,531,727,787]" box="[151,531,727,754]" pageId="6" pageNumber="417">
<heading id="1F1F81B3716F3B3DFF4D15FD23731DE2" bold="true" box="[151,531,727,754]" fontSize="11" level="1" pageId="6" pageNumber="417" reason="1">
<emphasis id="769CEACD716F3B3DFF4D15FD23731DE2" bold="true" box="[151,531,727,754]" pageId="6" pageNumber="417">
<taxonomicName id="83E84D5C716F3B3DFF4D15FD20AA1DE1" ID-CoL="72TQG" box="[151,458,727,753]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="417" phylum="Annelida" rank="species" species="pseudosessiloa" status="sp. nov.">
<emphasis id="769CEACD716F3B3DFF4D15FD20AA1DE1" bold="true" box="[151,458,727,753]" italics="true" pageId="6" pageNumber="417">Marphysa pseudosessiloa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DAF57B6716F3B3DFE0B15F223731DE2" box="[465,531,728,754]" pageId="6" pageNumber="417" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="445736DF716F3B3DFF4D15D0206A1C03" blockId="6.[151,531,727,787]" box="[151,266,762,787]" pageId="6" pageNumber="417">
(
<figureCitation id="DCD32A5A716F3B3DFF4515D021871C03" box="[159,231,762,787]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Figs 3</figureCitation>
,
<figureCitation id="DCD32A5A716F3B3DFF2915D120621C03" box="[243,258,763,787]" captionStart="FIGURE 4" captionStartId="9.[151,250,1234,1256]" captionTargetBox="[445,1142,205,1196]" captionTargetId="figure@9.[436,1143,193,1207]" captionTargetPageId="9" captionText="FIGURE 4. Neighbor joining tree based on cytochrome oxidase I (COI) sequences and Kimura- 2 - parameters model; bootstrap branch support above 70 % shown on branches." httpUri="https://zenodo.org/record/580540/files/figure.png" pageId="6" pageNumber="417">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0CF26554716F3B3DFF4D1468232E1CB3" pageId="6" pageNumber="417" type="materials_examined">
<paragraph id="445736DF716F3B3DFF4D1468232E1CB3" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">
<materialsCitation id="F4803C82716F3B3DFF4D1468237E1C6F" ID-GBIF-Occurrence="1503335923" collectingDate="2011-02-17" country="Australia" county="Material" latitude="-33.614998" location="Careel Bay" longLatPrecision="19" longitude="151.32333" municipality="Pittwater" pageId="6" pageNumber="417" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="769CEACD716F3B3DFF4D1468201D1C4B" bold="true" box="[151,381,834,859]" pageId="6" pageNumber="417">
<collectingCounty id="AD364E53716F3B3DFF4D1468219F1C4B" box="[151,255,834,859]" pageId="6" pageNumber="417">Material</collectingCounty>
examined
</emphasis>
.
<typeStatus id="9B53887D716F3B3DFE54146923461C4B" box="[398,550,835,859]" pageId="6" pageNumber="417" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="3CFF764F716F3B3DFDEF146823FE1C4B" box="[565,670,834,859]" name="Australia" pageId="6" pageNumber="417">Australia</collectingCountry>
,
<collectingRegion id="862CF83D716F3B3DFD77146922E01C4B" box="[685,896,834,859]" country="Australia" name="New South Wales" pageId="6" pageNumber="417">
New
<collectingRegion id="862CF83D716F3B3DFD37146822E01C4B" box="[749,896,834,859]" inferredBySuffix="true" pageId="6" pageNumber="417">
<collectingRegion id="862CF83D716F3B3DFD371468224F1C4B" box="[749,815,834,859]" country="Tanzania" name="Kusini Unguja" pageId="6" pageNumber="417">South</collectingRegion>
Wales
</collectingRegion>
</collectingRegion>
,
<collectingMunicipality id="A433ACA5716F3B3DFC54146822951C4B" box="[910,1013,834,859]" pageId="6" pageNumber="417">Pittwater</collectingMunicipality>
,
<location id="41376004716F3B3DFBDE146825E61C4B" LSID="urn:lsid:plazi:treatment:CC4187C9716F3B32FF4D15FD239118FE:41376004716F3B3DFBDE146825E61C4B" box="[1028,1158,834,859]" country="Australia" county="Material" latitude="-33.614998" longLatPrecision="19" longitude="151.32333" municipality="Pittwater" name="Careel Bay" pageId="6" pageNumber="417" stateProvince="New South Wales">Careel Bay</location>
,
<geoCoordinate id="21DC5018716F3B3DFB4F1469244B1C4B" box="[1173,1323,834,859]" direction="south" orientation="latitude" pageId="6" pageNumber="417" precision="15" value="-33.614998">33° 36' 54&quot;S</geoCoordinate>
,
<geoCoordinate id="21DC5018716F3B3DFAE1146821B21C6F" direction="east" orientation="longitude" pageId="6" pageNumber="417" precision="15" value="151.32333">151° 19' 24&quot;E</geoCoordinate>
,
<date id="3056101F716F3B3DFF07144D20081C6F" box="[221,360,870,895]" pageId="6" pageNumber="417" value="2011-02-17">
<collectingDate id="2012E9F7716F3B3DFF07144D20081C6F" box="[221,360,870,895]" pageId="6" pageNumber="417" value="2011-02-17">17 Feb 2011</collectingDate>
</date>
(AM W.37249)
</materialsCitation>
.
<materialsCitation id="F4803C82716F3B3DFDF2144D232A1CB3" ID-GBIF-Occurrence="1503335921" collectingDate="2011-02-17" country="Australia" county="Material" latitude="-33.614998" location="Careel Bay" longLatPrecision="19" longitude="151.32333" municipality="Pittwater" pageId="6" pageNumber="417" specimenCount="4" stateProvince="New South Wales" typeStatus="paratype">
<typeStatus id="9B53887D716F3B3DFDF2144D23A71C6F" box="[552,711,871,895]" pageId="6" pageNumber="417" type="paratype">PARATYPES</typeStatus>
: same locality as for
<typeStatus id="9B53887D716F3B3DFC68144C25771C6F" box="[946,1047,870,895]" pageId="6" pageNumber="417" type="holotype">holotype</typeStatus>
, AM W.37089, AM W.37088, AM W.37246, AM W.37250 (
<specimenCount id="52EEFD56716F3B3DFE6814A123221CB3" box="[434,578,906,931]" pageId="6" pageNumber="417" type="generic">4 specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF26554716F3B3CFF1D1484224B18A9" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417" type="description">
<paragraph id="445736DF716F3B3DFF1D148422A81BD3" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">
<emphasis id="769CEACD716F3B3DFF1D1484201D1CD7" bold="true" box="[199,381,942,967]" pageId="6" pageNumber="417">Measurements.</emphasis>
<typeStatus id="9B53887D716F3B3DFE56148420971CD7" box="[396,503,942,967]" pageId="6" pageNumber="417" type="holotype">Holotype</typeStatus>
, complete female with regenerating pygidium,
<quantity id="83109B3A716F3B3DFBE1148425C01CD6" box="[1083,1184,942,967]" metricMagnitude="-1" metricUnit="m" metricValue="1.64" pageId="6" pageNumber="417" unit="mm" value="164.0">164 mm</quantity>
in total length, 379 chaetigers, in four fragments, 132 anteriormost chaetigers, 53 median and 174 posteriormost chaetigers fixed in 8% formalin, 20 median chaetigers fixed in 95% ethanol,
<quantity id="83109B3A716F3B3DFD2314DC22301B1E" box="[761,848,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="417" unit="mm" value="6.5">6.5 mm</quantity>
length through chaetiger 10 and about
<quantity id="83109B3A716F3B3DFAD614DC24031B1E" box="[1292,1379,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="417" unit="mm" value="3.5">3.5 mm</quantity>
with parapodia (
<quantity id="83109B3A716F3B3DFECC133120381B22" box="[278,344,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="417" unit="mm" value="2.0">2 mm</quantity>
without parapodia) in width at chaetiger 10 (maximal width). One
<typeStatus id="9B53887D716F3B3DFB8E133125D41B23" box="[1108,1204,1051,1075]" pageId="6" pageNumber="417" type="paratype">paratype</typeStatus>
complete specimen, with
<quantity id="83109B3A716F3B3DFF0B131520501B46" box="[209,304,1087,1111]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="417" unit="mm" value="100.0">100 mm</quantity>
in total length, 239 chaetigers,
<quantity id="83109B3A716F3B3DFD43131423901B46" box="[665,752,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="417" unit="mm" value="5.2">5.2 mm</quantity>
in length through chaetiger 10 and
<quantity id="83109B3A716F3B3DFB53131525AC1B46" box="[1161,1228,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="417" unit="mm" value="2.0">2 mm</quantity>
with parapodia (
<quantity id="83109B3A716F3B3DFA4A131521A31B6A" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="417" unit="mm" value="1.0">1 mm</quantity>
without parapodia) in width at chaetiger 10. Three
<typeStatus id="9B53887D716F3B3DFCD51349221B1B6B" box="[783,891,1123,1147]" pageId="6" pageNumber="417" type="paratype">paratypes</typeStatus>
incomplete specimens or regenerating posterior end ranging from
<quantity id="83109B3A716F3B3DFEBC13AC20811B8E" box="[358,481,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="6" pageNumber="417" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">5055 mm</quantity>
in total length, 85249 chaetigers,
<quantity id="83109B3A716F3B3DFCA213AC228B1B8E" box="[888,1003,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="6" pageNumber="417" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
in length through chaetiger 10,
<quantity id="83109B3A716F3B3DFA8713AC21A31BD2" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="417" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
with parapodia (
<quantity id="83109B3A716F3B3DFE5A138020B71BD2" box="[384,471,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="417" unit="mm" value="1.6">1.6 mm</quantity>
without parapodia) in width at chaetiger 10.
</paragraph>
<paragraph id="445736DF716F3B3DFF1D13E422F61BF7" blockId="6.[151,1437,834,2011]" box="[199,918,1230,1255]" pageId="6" pageNumber="417">
<emphasis id="769CEACD716F3B3DFF1D13E420391BF7" bold="true" box="[199,345,1230,1255]" pageId="6" pageNumber="417">Description.</emphasis>
(Of
<typeStatus id="9B53887D716F3B3DFE5713E420921BF7" box="[397,498,1230,1255]" pageId="6" pageNumber="417" type="holotype">holotype</typeStatus>
, values in parentheses for
<typeStatus id="9B53887D716F3B3DFCC013E522EA1BF7" box="[794,906,1231,1255]" pageId="6" pageNumber="417" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="445736DF716F3B3DFF1D13D8204D1A3E" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">Live specimens whitish translucent with pink to red hues due to red blood. Fixed specimens white, lacking color pattern.</paragraph>
<paragraph id="445736DF716F3B3DFF1D121022CB1A8B" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">Body long, slender, abruptly tapering towards anterior end from around chaetiger 5, evenly tapering towards posterior end; cross section dorsoventrally flattened at anterior region and rounded at median and posterior regions. Chaetigers around 35 times wider than long at widest region of body.</paragraph>
<paragraph id="445736DF716F3B3DFF1D128C22ED1917" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">
Prostomium as long and wide as peristomium, two-thirds (half) as deep as peristomium; dorsally flat with anterior end higher (completely flat), anteriorly triangular (round); not bilobed; median sulcus absent at anterior and dorsal sides, conspicuously present at ventral side (
<figureCitation id="DCD32A5A716F3B3DFCD912C422241917" box="[771,836,1518,1543]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
AD).
</paragraph>
<paragraph id="445736DF716F3B3DFF1D113824FC185B" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">
Median and lateral antennae in straight line, palps inserted slightly more anteriorly (
<figureCitation id="DCD32A5A716F3B3DFB7011382590193B" box="[1194,1264,1554,1579]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
A, B). Median antenna isolated by gap from lateral antennae and palps; longest prostomial appendage, as long as prostomium, folding back to posterior margin of second peristomium ring (mid second peristomial ring). Lateral antennae around two thirds length of prostomium, folding back to mid of second peristomium ring (posterior margin of first to anterior margin of second peristomium ring). Palps shortest, around as long as half of prostomium, folding back to mid of first peristomial ring. Antennal styles and palpostyle tapering (palpostyle digitiform), all irregularly articulated and sensory papillae distributed in rings, last visible only under SEM, under stereomicroscope prostomial appendages appearing wrinkled (
<figureCitation id="DCD32A5A716F3B3DFD50102423AD1837" box="[650,717,1806,1831]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
C, E). Antennophores and palpophores ring shaped. One pair of inconspicuous eyes present, in shape of dark line at base of palpophores (lateral antennae antennophores) (
<figureCitation id="DCD32A5A716F3B3DFAEC10182418185B" box="[1334,1400,1842,1867]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="445736DF716F3B3DFF1D107C245918A7" blockId="6.[151,1437,834,2011]" pageId="6" pageNumber="417">
Separation between peristomial rings distinct all around. Second peristomial ring about as long as 1/31/2 of complete peristomium. Peristomial ventro-lateral lips distinct laterally just as elevated surface. Peristomial anterior margin of dorsal and ventral sides in shallow arc, lateral margins longer; shortest on dorsal side (
<figureCitation id="DCD32A5A716F3B3DFB0810B4247418A7" box="[1234,1300,1950,1975]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="445736DF716F3B3CFF1D10E8220A1E75" blockId="6.[151,1437,834,2011]" lastBlockId="7.[151,1437,151,1977]" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417">
Posterior end of muscularized pharynx at chaetiger 4. Mandibles missing calcareous cutting plates. MxI twice as long as carriers and six times longer than locking system. MxIII at least in part located ventral to MxII; attachment lamella upside down drop-shaped at anterior end of median margin of MxIII (
<figureCitation id="DCD32A5A716E3B3CFBA6179625DE1FC4" box="[1148,1214,188,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
F). Left MxIV plate semi-circular; attachment lamella shorter than plate, almost as wide as plate, along most of anterior edge, missing on lateral edge of plate (
<figureCitation id="DCD32A5A716E3B3CFE6E162E20991E0C" box="[436,505,260,285]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
F). Right MxIV attachment lamella starting between lateralmost teeth 23, along around 2/3 of plate, longest at lateral side and evenly shorter towards opposite side (
<figureCitation id="DCD32A5A716E3B3CFBE5160D25E11E50" box="[1087,1153,295,320]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
F). Maxillary formula: I= 1+1, II= 5(6)+6, III= 6(5)+0, IV= 6(7)+9(8), V= 1+1, VI absent.
</paragraph>
<paragraph id="445736DF716E3B3CFF1D164522281DE0" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="418">
Pre-neurochaetal lobe shorter than neurochaetal lobe along whole body. Post-neurochaetal lobe longer than neurochaetal lobe in anterior chaetigers, decreasing in length along body, becoming as long as or shorter than neurochaetal lobe in median and posterior chaetigers. Anterior post-neurochaetal lobe wide, distally truncate with longer dorsal edge (
<figureCitation id="DCD32A5A716E3B3CFE5816F620A61EE4" box="[386,454,476,501]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
G, H). Neurochaetal lobe round in anteriormost region (P1) (
<figureCitation id="DCD32A5A716E3B3CFB4916F625BA1EE4" box="[1171,1242,476,501]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">
Fig.
<date id="3056101F716E3B3CFB1616F625BA1EE4" box="[1228,1242,476,500]" pageId="7" pageNumber="418">3</date>
</figureCitation>
I), tapering from parapodia P2P6 (
<figureCitation id="DCD32A5A716E3B3CFEBE16D520C51D08" box="[356,421,511,536]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
H, J). Anterior notopodial cirri fusiform (tapering), longer than neurochaetal lobe (
<figureCitation id="DCD32A5A716E3B3CFAE516D524E21D08" box="[1343,1410,511,536]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">
Fig.
<date id="3056101F716E3B3CFAAE152A24E21D08" box="[1396,1410,512,536]" pageId="7" pageNumber="418">3</date>
</figureCitation>
I); median and posterior cirri slender and slightly shorter than anterior ones, but still longer than neurochaetal lobes (
<figureCitation id="DCD32A5A716E3B3CFF45156D21831D70" box="[159,227,583,608]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
J) (shorter than neurochaetal lobe in branchiated region). Lateral sense organs slightly elevated in P1, as conspicuously ciliated bump in remaining of body (
<figureCitation id="DCD32A5A716E3B3CFCDE1546222D1D94" box="[772,845,620,645]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
H). Ventral cirri tapering with round wide tips at chaetigers 1 to 4, around 2/3 as long as notopodial cirri (
<figureCitation id="DCD32A5A716E3B3CFCCA15A522331DB8" box="[784,851,655,680]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">
Fig.
<date id="3056101F716E3B3CFC9F15BA22331DB8" box="[837,851,656,680]" pageId="7" pageNumber="418">3</date>
</figureCitation>
I); basally inflated from chaetiger 5, inflated base of round shape with round tip (
<figureCitation id="DCD32A5A716E3B3CFE22159E23211DDC" box="[504,577,692,717]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
J), gradually decreasing from chaetiger 61 (35, 43) to 152 (85); round (triangular) tapering with distinct tip from chaetiger 153 (86).
</paragraph>
<paragraph id="445736DF716E3B3CFF1D15D623491CB4" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="418">
Branchiae palmate (
<figureCitation id="DCD32A5A716E3B3CFE7215D6208B1C04" box="[424,491,764,789]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
J, K), with 2 (1) filaments from chaetiger 28 (2126), reaching maximum of 6 (4, 5) filaments at chaetiger 129 (71106; first 1/3 of body), terminating at last chaetiger (21 chaetigers before pygidium). Best developed branchial filaments around 4 times longer than notopodial cirri and 8 times longer than branchial stems, length of filaments decreasing at median region of body. Branchial stems distally loose, basally attached to notopodial cirri and body (
<figureCitation id="DCD32A5A716E3B3CFE1814A623641CB4" box="[450,516,908,933]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
K).
</paragraph>
<paragraph id="445736DF716E3B3CFF1D1485252119AC" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="418">
Notopodial aciculae yellow, present in notopodial cirri along whole body. Neurochaetae in two distinct bundles; supra-acicular with limbate chaetae and pectinate chaetae, subacicular with bidentate compound falciger chaetae and subacicular hooks. Neuroaciculae blunt to tapering, brown in anterior chaetigers, becoming yellow at posterior most chaetigers, dorsal to midline in P1 and in midline thereafter; distributed in oblique row, anteriormost neuroaciculae also dorsalmost in parapodium (
<figureCitation id="DCD32A5A716E3B3CFD7F131523881B48" box="[677,744,1087,1112]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
L). Number of neuroaciculae, limbate chaetae and compound falciger chaetae decreasing towards posterior end. Three neuroaciculae present in P1 and P2, in posterior chaetiger mostly one present. Around 15 limbate chaetae at anteriormost region (P1),
<quantity id="83109B3A716E3B3CFC3113A225441BB0" box="[1003,1060,1159,1184]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="418" unit="in" value="10.0">10 in</quantity>
P2 and more posterior chaetigers with as few as 6. Limbate chaetae longer than all other chaetae, strongly serrated, serration increases towards flat distal end (spike like serration surround middle of its length). Compound falciger chaetae numerous in P1, around 40 (30) present, 15 (10) by P2 and
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more posterior parapodia; shaft and appendage of compound falciger chaetae strongly serrated (shaft serration spike like toward proximal end of chaetae); appendage bidentate with guards symmetrically blunt, marginally serrated, without mucros; both teeth directed laterally, distal tooth curved, proximal tooth straight, perpendicular to length of chaetae, about twice as long as distal tooth (
<figureCitation id="DCD32A5A716E3B3CFB131275246B1A68" box="[1225,1291,1375,1400]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
MO). One two curved thin pectinate chaetae present at anterior edge of supra-acicular bundle from P1P3; around 1116 teeth present (
<figureCitation id="DCD32A5A716E3B3CFF20128D205E1AD0" box="[250,318,1447,1472]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
H, Q, R); inner teeth equal in length; outer teeth not equal in length and longer than inner teeth. Curved thick pectinate chaetae in median to posterior regions with around 1214 coarse teeth with about same length at distal end but with increasingly deeper proximal ends from margins to center, teeth with deepest proximal end off center (
<figureCitation id="DCD32A5A716E3B3CFE99113E20E5193C" box="[323,389,1556,1581]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
M, S); 23 present in P4, 2
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P5 and
<quantity id="83109B3A716E3B3CFC95113E22F7193D" box="[847,919,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="7" pageNumber="418" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
P6. Pseudocompound chaetae and compound spiniger chaetae absent. Subacicular hook first present from chaetiger 37 (2736), present in all chaetigers thereafter, always single; yellow, clearly bidentate, thinner than aciculae; distal tooth erect tapering directed distally and much smaller than proximal tooth; proximal tooth perpendicular to length of hook, directed laterally to distally, tip tapering to round; guards truncate present on both sides of teeth (
<figureCitation id="DCD32A5A716E3B3CFC4F118E22B619AC" box="[917,982,1700,1725]" captionStart="FIGURE 3" captionStartId="8.[151,250,1670,1692]" captionTargetBox="[303,1275,207,1637]" captionTargetId="figure@8.[303,1275,207,1637]" captionTargetPageId="8" captionText="FIGURE 3. Marphysa pseudosessiloa n. sp. A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W. 37249. B, D, F, H J, M, N, Q. AM W. 37089. E, G, K, O, P, R- T. AM W. 37250. Scale bars: A D. 500 µm. E L. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri." httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
M, Q, T).
</paragraph>
<paragraph id="445736DF716E3B3CFF1D11ED23B51838" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="418">
Pygidium longer on ventral side, with two pairs of pygidial cirri attached to ventral margin; pygidial cirri triangular, about as long as pygidium, dorsal cirri as long as ventral (tapering, longer than pygidium, dorsal 3 times longer than ventral cirri in
<typeStatus id="9B53887D716E3B3CFE1E103B23441839" box="[452,548,1809,1833]" pageId="7" pageNumber="418" type="paratype">paratype</typeStatus>
AM W.37089).
</paragraph>
<paragraph id="445736DF716E3B3CFF1D101E224B18A9" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="418">
<emphasis id="769CEACD716E3B3CFF1D101E205F185D" bold="true" box="[199,319,1844,1869]" pageId="7" pageNumber="418">Variation.</emphasis>
Calcareous cutting plates of mandibles are probably dissolved due to formalin fixation. Thin pectinate is inconspicuous in P1 under light microscopy. It was only clearly observed in the
<typeStatus id="9B53887D716E3B3CFB04107D24231860" box="[1246,1347,1879,1904]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
, largest specimen. Spike-like serration present in some chaetae may be artifact of preparation. The appearance of pygidial cirri in the
<typeStatus id="9B53887D716E3B3CFEC9108A201518A9" box="[275,373,1952,1977]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
is probably due to recent regeneration.
</paragraph>
</subSubSection>
<caption id="1097665771613B33FF4D11AC205B18FD" httpUri="https://zenodo.org/record/580539/files/figure.png" pageId="8" pageNumber="419" targetBox="[303,1275,207,1637]" targetPageId="8">
<paragraph id="445736DF71613B33FF4D11AC205B18FD" blockId="8.[151,1436,1670,2029]" pageId="8" pageNumber="419">
<emphasis id="769CEACD71613B33FF4D11AC2075198B" bold="true" box="[151,277,1670,1692]" pageId="8" pageNumber="419">FIGURE 3.</emphasis>
<taxonomicName id="83E84D5C71613B33FEC411AC2346198C" box="[286,550,1670,1692]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="419" phylum="Annelida" rank="species" species="pseudosessiloa" status="sp. nov.">
<emphasis id="769CEACD71613B33FEC411AC2346198C" box="[286,550,1670,1692]" italics="true" pageId="8" pageNumber="419">Marphysa pseudosessiloa</emphasis>
</taxonomicName>
<emphasis id="769CEACD71613B33FDF511AD230B198C" bold="true" box="[559,619,1671,1692]" pageId="8" pageNumber="419">
<taxonomicNameLabel id="6DAF57B671613B33FDF511AD230B198C" box="[559,619,1671,1692]" pageId="8" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A. Anterior end of holotype, dorsal view. B. Anterior end of paratype, dorsal view. C. Anterior end of holotype, lateral view. D. Anterior end of paratype, lateral view. E. Anterior end of paratype, anterior view. F. Maxillary apparatus of paratype, dorsal view. G. Right parapodium 15 of paratype, anterior view. H. Left parapodium 31 of paratype, anterior view. I. Parapodium 3 of paratype. J. Parapodium 45 of paratype. K. Branchiae palmate, posterior view, left parapodium of chaetiger between 143 and 163 of paratype. L. Anterior parapodia, lateral view. M. Neurochaetae, acicula and subacicular hook, parapodium 185; arrow to shaft of wide pectinate chaeta. N. Compound falciger chaetae, parapodium 3. O. Compound falciger chaeta of paratype, parapodium 15. P. Compound falciger chaetae strongly serrated, parapodium 218 of paratype. Q. Neurochaetae, acicula, subacicular hook, parapodium 45 of paratype; arrow to shaft of thin pectinate chaeta. R. Thin pectinate chaeta, parapodium 81 of paratype. S. Curved thick pectinate chaetae, parapodium 218 of paratype. T. Subacicular hook bidentate, parapodium 143. A, C, L. AM W.37249. B, D, F, HJ, M, N, Q. AM W.37089. E, G, K, O, P, R- T. AM W.37250. Scale bars: AD. 500 µm. EL. 100 µm. M, N, P, Q, S, T. 10 µm. O, R. 2 µm. BrSt, branchial stem; NC, notopodial cirri.
</paragraph>
</caption>
<caption id="1097665771603B32FF4D13F823151A17" httpUri="https://zenodo.org/record/580540/files/figure.png" pageId="9" pageNumber="420" targetBox="[445,1142,205,1196]" targetPageId="9">
<paragraph id="445736DF71603B32FF4D13F823151A17" blockId="9.[151,1435,1234,1287]" pageId="9" pageNumber="420">
<emphasis id="769CEACD71603B32FF4D13F820721BF7" bold="true" box="[151,274,1234,1256]" pageId="9" pageNumber="420">FIGURE 4.</emphasis>
Neighbor joining tree based on cytochrome oxidase I (COI) sequences and Kimura-2-parameters model; bootstrap branch support above 70% shown on branches.
</paragraph>
</caption>
<subSubSection id="0CF2655471603B32FF1D1267246C1829" pageId="9" pageNumber="420" type="discussion">
<paragraph id="445736DF71603B32FF1D126722F219E2" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="420">
<emphasis id="769CEACD71603B32FF1D126720541A76" bold="true" box="[199,308,1357,1382]" pageId="9" pageNumber="420">Remarks</emphasis>
. Among the species of the Aenea-group,
<taxonomicName id="83E84D5C71603B32FCFB126422A71A75" box="[801,967,1358,1381]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="mortenseni">
<emphasis id="769CEACD71603B32FCFB126422A71A75" box="[801,967,1358,1381]" italics="true" pageId="9" pageNumber="420">M. mortenseni</emphasis>
</taxonomicName>
,
<emphasis id="769CEACD71603B32FC03126424081A75" box="[985,1384,1357,1381]" italics="true" pageId="9" pageNumber="420">
<taxonomicName id="83E84D5C71603B32FC031264250B1A75" box="[985,1131,1357,1381]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="bifurcata">M. bifurcata</taxonomicName>
,
<taxonomicName id="83E84D5C71603B32FBA0126424081A75" box="[1146,1384,1357,1381]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="sessilobranchiata">M. sessilobranchiata</taxonomicName>
</emphasis>
and
<taxonomicName id="83E84D5C71603B32FF4D125B23541A99" authority="Orensanz, 1990" authorityName="Orensanz" authorityYear="1990" box="[151,564,1393,1417]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="galluccii">
<emphasis id="769CEACD71603B32FF4D125B20151A99" box="[151,373,1393,1417]" italics="true" pageId="9" pageNumber="420">Marphysa galluccii</emphasis>
<bibRefCitation id="20794B2E71603B32FE5A125B23541A99" author="Orensanz" box="[384,564,1393,1417]" pageId="9" pageNumber="420" refString="Orensanz, J. M. (1990) The eunicemorph polychaete annelids from Antarctic and subantarctic seas with addenda to the Eunicemorpha of Argentina, Chile, New Zealand, Australia, and the Southern Indian Ocean. Biology of the Antarctic Seas XXI. Antarctic Research Series, 52, 1 ‾ 183." type="journal article" year="1990">Orensanz, 1990</bibRefCitation>
</taxonomicName>
are the most similar to this new species in having a tapering non-bilobed prostomium.
<taxonomicName id="83E84D5C71603B32FEF612BF23401ABD" box="[300,544,1429,1453]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="mortenseni">
<emphasis id="769CEACD71603B32FEF612BF23401ABD" box="[300,544,1429,1453]" italics="true" pageId="9" pageNumber="420">Marphysa mortenseni</emphasis>
</taxonomicName>
,
<taxonomicName id="83E84D5C71603B32FDF712BC23D81ABD" box="[557,696,1429,1453]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="bifurcata">
<emphasis id="769CEACD71603B32FDF712BC23D81ABD" box="[557,696,1429,1453]" italics="true" pageId="9" pageNumber="420">M. bifurcata</emphasis>
</taxonomicName>
and
<taxonomicName id="83E84D5C71603B32FD3612BC22131ABD" box="[748,883,1429,1453]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="galluccii">
<emphasis id="769CEACD71603B32FD3612BC22131ABD" box="[748,883,1429,1453]" italics="true" pageId="9" pageNumber="420">M. galluccii</emphasis>
</taxonomicName>
are clearly distinct from
<taxonomicName id="83E84D5C71603B32FB5012BC24371ABD" box="[1162,1367,1429,1453]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="pseudosessiloa" status="sp. nov.">
<emphasis id="769CEACD71603B32FB5012BC24371ABD" box="[1162,1367,1429,1453]" italics="true" pageId="9" pageNumber="420">M. pseudosessiloa</emphasis>
</taxonomicName>
<emphasis id="769CEACD71603B32FA8712BF24FC1ABE" bold="true" box="[1373,1436,1429,1454]" pageId="9" pageNumber="420">
<taxonomicNameLabel id="6DAF57B671603B32FA8712BF24FC1ABE" box="[1373,1436,1429,1454]" pageId="9" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in having prostomial appendages shorter than half the length of the prostomium; the first two species may also have bifurcated notopodial cirri, which is simple in the current species.
<taxonomicName id="83E84D5C71603B32FCA512F7253A1AE5" box="[895,1114,1501,1525]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="galluccii">
<emphasis id="769CEACD71603B32FCA512F7253A1AE5" box="[895,1114,1501,1525]" italics="true" pageId="9" pageNumber="420">Marphysa galluccii</emphasis>
</taxonomicName>
also differs from the current species in lacking thin pectinate chaetae, having only single filament branchiae and unidentate subacicular hooks. General features are similar among the specimens examined here,
<typeStatus id="9B53887D71603B32FCA2110C2283192E" box="[888,995,1574,1598]" pageId="9" pageNumber="420" type="paratype">paratypes</typeStatus>
and non-types of
<taxonomicName id="83E84D5C71603B32FB71110C24F4192D" box="[1195,1428,1573,1597]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="sessilobranchiata">
<emphasis id="769CEACD71603B32FB71110C24F4192D" box="[1195,1428,1573,1597]" italics="true" pageId="9" pageNumber="420">M. sessilobranchiata</emphasis>
</taxonomicName>
. The main divergences among them are the shape of the left MxIV plate, left and right MxIV attachment lamellae, teeth of subacicular hooks and position of branchial stem. In the current species branchiae are attached to a loose stem connected only basally to notopodial cirri and body wall, while in
<taxonomicName id="83E84D5C71603B32FC2D11B8258819B9" box="[1015,1256,1681,1705]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="sessilobranchiata">
<emphasis id="769CEACD71603B32FC2D11B8258819B9" box="[1015,1256,1681,1705]" italics="true" pageId="9" pageNumber="420">M. sessilobranchiata</emphasis>
</taxonomicName>
, as well as
<taxonomicName id="83E84D5C71603B32FAA711B8219D19DD" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="bifurcata">
<emphasis id="769CEACD71603B32FAA711B8219D19DD" italics="true" pageId="9" pageNumber="420">M. bifurcata</emphasis>
</taxonomicName>
, branchial stems are completely attached to the body wall, giving the appearance that branchial filaments are individually connected to the body wall as previously described.
</paragraph>
<paragraph id="445736DF71603B32FF1D11D7246C1829" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="420">
Molecular identification clearly distinguishes this species from other species of the genus (K2P= 1825%) and from
<taxonomicName id="83E84D5C71603B32FF0E100820001829" box="[212,352,1825,1849]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="bifurcata">
<emphasis id="769CEACD71603B32FF0E100820001829" box="[212,352,1825,1849]" italics="true" pageId="9" pageNumber="420">M. bifurcata</emphasis>
</taxonomicName>
(K2P= 17%), the only species of the Aenea-group included in the analyses (
<figureCitation id="DCD32A5A71603B32FB63100B259E1829" box="[1209,1278,1825,1850]" captionStart="FIGURE 4" captionStartId="9.[151,250,1234,1256]" captionTargetBox="[445,1142,205,1196]" captionTargetId="figure@9.[436,1143,193,1207]" captionTargetPageId="9" captionText="FIGURE 4. Neighbor joining tree based on cytochrome oxidase I (COI) sequences and Kimura- 2 - parameters model; bootstrap branch support above 70 % shown on branches." httpUri="https://zenodo.org/record/580540/files/figure.png" pageId="9" pageNumber="420">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0CF2655471603B32FF1D106F23AC1891" pageId="9" pageNumber="420" type="etymology">
<paragraph id="445736DF71603B32FF1D106F23AC1891" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="420">
<emphasis id="769CEACD71603B32FF1D106F202E184E" bold="true" box="[199,334,1861,1886]" pageId="9" pageNumber="420">Etymology.</emphasis>
It is very similar to
<taxonomicName id="83E84D5C71603B32FDEF106C2241184D" box="[565,801,1861,1885]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="sessilobranchiata">
<emphasis id="769CEACD71603B32FDEF106C2241184D" box="[565,801,1861,1885]" italics="true" pageId="9" pageNumber="420">M. sessilobranchiata</emphasis>
</taxonomicName>
in general morphology but it lacks the sessile branchiae and hence the specific name of
<taxonomicName id="83E84D5C71603B32FE22104023A51891" box="[504,709,1897,1921]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="420" phylum="Annelida" rank="species" species="pseudosessiloa">
<emphasis id="769CEACD71603B32FE221040236F1891" box="[504,527,1898,1921]" italics="true" pageId="9" pageNumber="420">M</emphasis>
.
<emphasis id="769CEACD71603B32FDC7104323A51891" box="[541,709,1897,1921]" italics="true" pageId="9" pageNumber="420">pseudosessiloa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF2655471603B32FF1D10A722EB18D9" pageId="9" pageNumber="420" type="materials_examined">
<paragraph id="445736DF71603B32FF1D10A722CC18B6" blockId="9.[151,1437,1357,2030]" box="[199,940,1933,1958]" pageId="9" pageNumber="420">
<materialsCitation id="F4803C8271603B32FF1D10A722CC18B6" ID-GBIF-Occurrence="1503335926" box="[199,940,1933,1958]" country="Australia" location="Careel Bay" municipality="Pittwater" pageId="9" pageNumber="420" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="769CEACD71603B32FF1D10A7200518B6" bold="true" box="[199,357,1933,1958]" pageId="9" pageNumber="420">
<typeStatus id="9B53887D71603B32FF1D10A7206218B6" box="[199,258,1933,1958]" pageId="9" pageNumber="420">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="3CFF764F71603B32FEB610A720B618B6" box="[364,470,1933,1958]" name="Australia" pageId="9" pageNumber="420">Australia</collectingCountry>
,
<collectingRegion id="862CF83D71603B32FE3B10A423CC18B6" box="[481,684,1933,1958]" country="Australia" name="New South Wales" pageId="9" pageNumber="420">
New
<collectingRegion id="862CF83D71603B32FDC710A723CC18B6" box="[541,684,1933,1958]" inferredBySuffix="true" pageId="9" pageNumber="420">
<collectingRegion id="862CF83D71603B32FDC710A7233E18B6" box="[541,606,1933,1958]" country="Tanzania" name="Kusini Unguja" pageId="9" pageNumber="420">South</collectingRegion>
Wales
</collectingRegion>
</collectingRegion>
,
<collectingMunicipality id="A433ACA571603B32FD6C10A7227D18B6" box="[694,797,1933,1958]" pageId="9" pageNumber="420">Pittwater</collectingMunicipality>
,
<location id="4137600471603B32FCF210A722C818B6" LSID="urn:lsid:plazi:treatment:CC4187C9716F3B32FF4D15FD239118FE:4137600471603B32FCF210A722C818B6" box="[808,936,1933,1958]" country="Australia" municipality="Pittwater" name="Careel Bay" pageId="9" pageNumber="420" stateProvince="New South Wales">Careel Bay</location>
.
</materialsCitation>
</paragraph>
<paragraph id="445736DF71603B32FF1D109B22EB18D9" blockId="9.[151,1437,1357,2030]" box="[199,907,1969,1994]" pageId="9" pageNumber="420">
<emphasis id="769CEACD71603B32FF1D109B204A18DA" bold="true" box="[199,298,1969,1994]" pageId="9" pageNumber="420">Habitat.</emphasis>
Intertidal mud in seagrass beds of
<taxonomicName id="83E84D5C71603B32FD6B109822E318D9" box="[689,899,1970,1993]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="9" pageNumber="420" phylum="Tracheophyta" rank="species" species="capricorni">
<emphasis id="769CEACD71603B32FD6B109822E318D9" box="[689,899,1970,1993]" italics="true" pageId="9" pageNumber="420">Zostera capricorni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF2655471603B32FF1D10FF239118FE" box="[199,753,2005,2030]" pageId="9" pageNumber="420" type="distribution">
<paragraph id="445736DF71603B32FF1D10FF239118FE" blockId="9.[151,1437,1357,2030]" box="[199,753,2005,2030]" pageId="9" pageNumber="420">
<emphasis id="769CEACD71603B32FF1D10FF200018FE" bold="true" box="[199,352,2005,2030]" pageId="9" pageNumber="420">Distribution.</emphasis>
Only known from the
<typeStatus id="9B53887D71603B32FDBA10FC23F018FE" box="[608,656,2006,2030]" pageId="9" pageNumber="420">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>