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<document id="391FDD5A7A3581E7BCA732C273D038E7" ID-DOI="10.3897/asp.82.e114351" ID-ZooBank="8070F22C-9721-418D-B82F-9333A0076455" ID-publisher-id="114351" URI-arpha="7B065A72-C3DA-5CEF-9F0E-0B883828FCF8" URI-zoobank="http://zoobank.org/8070F22C-9721-418D-B82F-9333A0076455" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1722960088179" checkinUser="pensoft" docAuthor="Ren, Mengjiao, Tian, Chunpo, Grossi, Alexandra A., Zou, Fasheng &amp; Gustafsson, Daniel R." docDate="2024" docId="BEABE1D94E44581BBD9D065390D913E2" docLanguage="en" docName="Arthropod Systematics &amp; Phylogeny 82: 585-605" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 82" docSource="https://arthropod-systematics.arphahub.com/article/114351/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Coronedax longiceps Ren, Tian, Grossi, Zou &amp; Gustafsson 2024, sp. n." docType="treatment" docUuid="794224B7-CE8B-4581-A03A-3E082A3C268B" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="605" masterDocId="7B065A72C3DA5CEF9F0E0B883828FCF8" masterDocTitle="Cryptic genera, cryptic species: phylogeny of the genus Philopteroides Mey, 2004, sensu lato, with descriptions of two new genera and one new species" masterLastPageNumber="605" masterPageNumber="585" pageNumber="585" updateTime="1722963898131" updateUser="ExternalLinkService">
<mods:mods id="CC7044C849D5646BC6C162B6B30F342A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1C077E003DEF65CF34D1217314B533BB">Cryptic genera, cryptic species: phylogeny of the genus Philopteroides Mey, 2004, sensu lato, with descriptions of two new genera and one new species</mods:title>
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<mods:namePart id="418D7ABDC0D2D3309AB07418609933AC">Ren, Mengjiao</mods:namePart>
<mods:nameIdentifier id="2901DA8DD0485D9B1B8D87E19A0368BB" type="ORCID">0000-0002-0108-6104</mods:nameIdentifier>
<mods:affiliation id="705D0A4291250397C4975F57BB840CF8">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China</mods:affiliation>
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<mods:namePart id="DA0586E7E05D73469C31CAA96663DC79">Tian, Chunpo</mods:namePart>
<mods:nameIdentifier id="1DF8BC4B4C5B95505BB72E5B83983FAA" type="ORCID">0000-0001-8681-4475</mods:nameIdentifier>
<mods:affiliation id="52010DE9719713F821B4C8BDD4297FB2">College of Life Sciences, Shaanxi Normal University, 620 West Chang’an Street, Chang’an District, Xi’an City, 710119, Shaanxi Province, China</mods:affiliation>
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<mods:namePart id="582A6B0AF4446D6ABE997C2B798BC4C6">Grossi, Alexandra A.</mods:namePart>
<mods:nameIdentifier id="38B8414DC18F084924EED20D5E7C94ED" type="ORCID">0000-0002-1397-4221</mods:nameIdentifier>
<mods:affiliation id="53EC0951D50391AF0072B52FCBDB4604">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China</mods:affiliation>
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<mods:namePart id="56823C4B9D026C1E9AA0FC5F22473DDA">Zou, Fasheng</mods:namePart>
<mods:nameIdentifier id="8E8ED4CC7B14B1D06B3338ABA21A3257" type="ORCID">0000-0002-8913-5651</mods:nameIdentifier>
<mods:affiliation id="04E2D64875C7443C57DC90D520501F34">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China</mods:affiliation>
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<mods:namePart id="CD140AF97B0A719FFDCB944B9A4FFB1D">Gustafsson, Daniel R.</mods:namePart>
<mods:nameIdentifier id="DCFA6AE949367AB117BA072AA3C769E3" type="ORCID">0000-0001-8868-8310</mods:nameIdentifier>
<mods:affiliation id="2149CF6EBBA2C4D45A73711F66551983">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Library of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang West Road, Haizhu District, Guangzhou, 510260, Guangdong Province, China</mods:affiliation>
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<mods:title id="0D5CA628EB671FD436B41E920CC4D65D">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="2CA9706F8EB65A502CC85B761AE5AE2B">2024</mods:date>
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<mods:number id="C7DAD21C20936130E13D0FE732A49044">82</mods:number>
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<treatment id="BEABE1D94E44581BBD9D065390D913E2" ID-DOI="http://doi.org/10.5281/zenodo.13240743" ID-Zenodo-Dep="13240743" ID-arpha="BEABE1D9-4E44-581B-BD9D-065390D913E2" ID-zoobank="https://zoobank.org/794224B7-CE8B-4581-A03A-3E082A3C268B" LSID="urn:lsid:zoobank.org:act:794224B7-CE8B-4581-A03A-3E082A3C268B" LSID-ZBK="urn:lsid:zoobank.org:act:794224B7-CE8B-4581-A03A-3E082A3C268B" httpUri="http://treatment.plazi.org/id/BEABE1D94E44581BBD9D065390D913E2">
<subSubSection id="59722ED12BCA6801838911B702D5C2A9" type="nomenclature">
<paragraph id="661650DA5003870A8E8647B2B7CB141B">
<taxonomicName id="093E16FEB74902A82A461767BEC7AB5B" authority="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityName="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Phthiraptera" phylum="Arthropoda" rank="species" species="longiceps" status="sp. nov.">
<emphasis id="017EC58979406C8EA6BCC7EE0890E232" italics="true">Coronedax longiceps</emphasis>
Ren, Tian, Grossi, Zou &amp; Gustafsson
</taxonomicName>
<taxonomicNameLabel id="FD07F5EDB2BF01C749805D41D6185DF2" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5ED24C1499419C3011CFD040DD83850F" type="description">
<paragraph id="1A82F34A9E14ECA2AAD8873E3E04F0F9">
<figureCitation id="AB555E905D1A0F7669273BC78AED48B3" captionStart="Figures 2–17" captionStartId="F2" captionText="Figures 2–17. Comparison of the male genitalia of some species of Philopteroides Mey, 2004, Coronedax gen. n., and Stasiasticopterus gen. n. All figures redrawn from their original descriptions (Tandan 1955; Mey 2004; Valim &amp; Palma 2013), unless otherwise noted. Illustrations have been rescaled to be roughly the same size. Illustrations are in ventral view, unless otherwise noted. No illustrations of the genitalia have ever been published for Philopteroides lineatus (Giebel, 1874), Philopteroides mitsusui (Uchida, 1948), or Stasiasticopterus kayanobori (Uchida, 1948). 2 Philopteroides novaezelandiae Mey, 2004. 3 Philopteroides xenicus Mey, 2004. 4 Philopteroides fuliginosus Valim &amp; Palma, 2013. 5 Philopteroides macrocephalus Valim &amp; Palma, 2013. 6 Philopteroides gigas Najer et al., 2016. 7 Philopteroides sinancorellus Najer et al., 2016. 8 Philopteroides sclerotifrons (Tandan, 1955); no scale in original. 9 Philopteroides pilgrimi Valim &amp; Palma, 2013. 10 Philopteroides beckeri (Mey, 2004) (redrawn from Valim &amp; Palma 2013). 11 Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 a). 12 Coronedax longiceps sp. n. 13 Stasiasticopterus longiclypeatus (Gustafsson et al., 2022 b). 14 Stasiasticopterus holosternus (Gustafsson et al., 2022 b). 15 Stasiasticopterus haerixos (Gustafsson et al., 2022 b). 16 Stasiasticopterus flavala (Najer &amp; Sychra [in Najer et al.], 2012 a. 17 Stasiasticopterus cucphuongensis (Mey, 2004) (redrawn from Gustafsson et al. 2022 b). Bold grey lines signify the generic divisions used here; narrow grey lines signify groups that are morphologically different from the type species of the respective genera, but where there is insufficient data to propose addition (e. g., subgeneric) limits. The two species here considered incerta sedis are placed in separate groups, pending further investigations. — Abbreviations used: BA = basal apodeme; GP = gonopore; MS = mesosome; lpmes = lateral posterios mesosomal setae; PM = parameres; pst 1–2 = parameral setae 1–2 (2 distal to 1)." figureDoi="10.3897/asp.82.e114351.figures2-17" httpUri="https://binary.pensoft.net/fig/1108929">Figures 12</figureCitation>
,
<figureCitation id="2545E7CEB3D5FB3D3E75764B6CC9FE12" captionStart="Figures 18, 19" captionStartId="F3" captionText="Figures 18, 19. Coronedax longiceps sp. n. 18 male habitus, dorsal and ventral views. 19 female habitus, dorsal and ventral views. — Abbreviations used: mms = marginal mesometanotal setae; ppss = pronotal post-spiracular seta; ps = paratergal setae; psps = principal post-spiracular setae; ss = sutural setae; sts = sternal setae; tps = tergal posterior setae." figureDoi="10.3897/asp.82.e114351.figures18-19" httpUri="https://binary.pensoft.net/fig/1108930">18–19</figureCitation>
,
<figureCitation id="4534E32A85E0F4C4A75BB2CD8614A971" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">20–23</figureCitation>
,
<figureCitation id="78DA1B834971313DD9235511073056E3" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELTAK" type="Type host">
<paragraph id="9A207E48DAED736FBF55A2BA1534B8F8">
<heading id="08898E508043727C07A97C53A99A5F02" reason="title">Type host.</heading>
</paragraph>
<paragraph id="EFC9F01FA60C5E62DF800B3CDCD14814">
<taxonomicName id="A0608DFDD6EBEF28338ACA5C1DE368EE" authority="(Gould, 1852)" baseAuthorityName="Gould" baseAuthorityYear="1852" class="Aves" family="Monarchidae" genus="Terpsiphone" kingdom="Animalia" order="Passeriformes" phylum="Chordata" rank="species" species="incei">
<emphasis id="57DD3D88F2DA23E79C94E764ED519BDE" italics="true">Terpsiphone incei</emphasis>
(Gould, 1852)
</taxonomicName>
– Amur paradise flycatcher.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3TAK" type="diagnosis">
<paragraph id="5FD5C2F72EFFDAA41E198FB7FC533728">
<heading id="A5A23E14E5513CCB283D8A2BD0EE0BC1" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D45506B739BB0027C446F996ABC92B5D">
Morphologically similar to
<taxonomicName id="3F709BBFB7361BD2A46DCEAC95432FC2" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="BD9114D0FF9DE03BACBFEB39814E464F" italics="true">Coronedax terpsiphoni</emphasis>
</taxonomicName>
(Najer &amp; Sychra [in
<bibRefCitation id="3867B9107525598F6E588806FDA1263A" DOI="10.2478/s11686-012-0005-x" author="Najer T &amp; Sychra O &amp; Literák I &amp; Procházka P &amp; Capek M &amp; Koubek P" journalOrPublisher="Acta Parasitologica" pagination="90-98" refId="B45" refString="Najer T, Sychra O, Literák I, Procházka P, Capek M, Koubek P (2012 b) Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides. Acta Parasitologica 57: 90–98. https://doi.org/10.2478/s11686-012-0005-x" title="Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides." volume="57">Najer et al.], 2012 b</bibRefCitation>
), but can be separated by the following characters: 1) head proportionately longer and narrower in
<taxonomicName id="3D4221096F4D4D858E127E816AFE169E" authorityName="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="AC0DBBECDFE2718A6B48AFF6E1BD70CA" italics="true">C. longiceps</emphasis>
</taxonomicName>
than in
<taxonomicName id="4BA9964D3A380E92A57B9E3D402F8C7C" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="BE1CD511BAFB88674FE195ECD3C88317" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
(Figs
<figureCitation id="48B8925D8200299D096300C18D8ECB59" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">24</figureCitation>
,
<figureCitation id="704854C5C0E9A6F2091041D5DCCD48CB" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">25</figureCitation>
); 2) dorsal anterior plate more slender in
<taxonomicName id="CC4D2D11B52B60BBB51C0DB64391D06C" authorityName="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="C191C020556DAAF12B23D5F3B797FFC8" italics="true">C. longiceps</emphasis>
</taxonomicName>
than in
<taxonomicName id="5BEA775310640CF3AA4F7B5DE0DE3AA5" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="C6B67EA09811B32237C1D46BE4648D8B" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
(Figs
<figureCitation id="99DC5F01D445A07B7B678F98715BC2D7" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">24</figureCitation>
,
<figureCitation id="69CD43556CFE1CCE882F8D908BBFCFE2" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">25</figureCitation>
); 3) female vulval margin of
<taxonomicName id="F37E40F8CC3D98BA6DE5D4B856BF807B" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="A35E9FFFE5BE086C784A707B10443883" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
with short
<abbrev id="ABBRID0EZWAK" xlink_title="vulval submarginal setae">
<emphasis id="860A9070061FE159F011733CE0B979DD" italics="true">vss</emphasis>
</abbrev>
, but these are absent in
<taxonomicName id="3252B0E92CFCF358D8BFA352E29EDEF9" authorityName="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="3AD5A771A79B4DCC43179A54FD837F50" italics="true">C. longiceps</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="51EA3653ED2DD2805E5FF64E78525DF8" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">23</figureCitation>
); 4) male subgenital plate with only 1 macroseta on each side (on segment VII) in
<taxonomicName id="F45004F4E111FB8E0F514AF2E0A52F94" authorityName="Ren, Tian, Grossi, Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="69E1DD3FF271B96FCB8F1A577CE98C84" italics="true">C. longiceps</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="909A57DD2689E91097DC8846FA5758B0" captionStart="Figures 18, 19" captionStartId="F3" captionText="Figures 18, 19. Coronedax longiceps sp. n. 18 male habitus, dorsal and ventral views. 19 female habitus, dorsal and ventral views. — Abbreviations used: mms = marginal mesometanotal setae; ppss = pronotal post-spiracular seta; ps = paratergal setae; psps = principal post-spiracular setae; ss = sutural setae; sts = sternal setae; tps = tergal posterior setae." figureDoi="10.3897/asp.82.e114351.figures18-19" httpUri="https://binary.pensoft.net/fig/1108930">18</figureCitation>
), but with 2 macrosetae on each side (on segments VII – VIII) in
<taxonomicName id="036D7FC2FE305FAFBCF30683FE7BD01D" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="EB64EFC05BCD7373C0D0766DCF86BD62" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="8F429EAE1A47D628CE55CA36979EDCBF" ID-DOI="10.3897/asp.82.e114351.figures24-25" ID-arpha="CC964376-F665-561F-8876-C7DAF54EB75F" httpUri="https://binary.pensoft.net/fig/1108932" startId="F5">
<paragraph id="429746C0F734C5B1D5F0742CC11F4266">
<label id="51C80EF9857C7E708FDF35CC3D98919A">Figures 24, 25.</label>
</paragraph>
<paragraph id="B68A4EBB540F88D46409FE811EB2195F">
Comparison in shape and proportions of head in species of
<taxonomicName id="E456007850ED12CDF1CDFE291FBB2C89" authorityName="Ren &amp; Tian &amp; Grossi &amp; Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6E7A5460F2F0104A44BF53BF083CD423" italics="true">Coronedax</emphasis>
</taxonomicName>
<emphasis id="D7CA0CABCA895EC96029D7CE01AB6CBA" bold="true">
<taxonomicNameLabel id="E9C0FEED3C237A5AB2CE8355B649E314" rank="genus">gen. n.</taxonomicNameLabel>
24
</emphasis>
male head of
<taxonomicName id="9035F1DFAA2A50EC5A644626C2ED20A4" authorityName="Ren &amp; Tian &amp; Grossi &amp; Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps" status="sp. nov.">
<emphasis id="AA7352BD7DCF36455FA4FAA1E952A8E6" italics="true">Coronedax longiceps</emphasis>
</taxonomicName>
<emphasis id="CA226AF6C8FE7437E5719F8F70389A7A" bold="true">
<taxonomicNameLabel id="D2F7162E4FEA302D3AAAA999DD0E7211" rank="species">sp. n.</taxonomicNameLabel>
25
</emphasis>
female head of
<taxonomicName id="8DB7048D556E2356840EEF768D760EA7" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="53294406EB293B8CA50B9B028975DAD3" italics="true">Coronedax terpsiphoni</emphasis>
</taxonomicName>
(Najer &amp; Sychra [in
<bibRefCitation id="98BCB00939AC191696F6E7923890E02E" DOI="10.2478/s11686-012-0005-x" author="Najer T &amp; Sychra O &amp; Literák I &amp; Procházka P &amp; Capek M &amp; Koubek P" journalOrPublisher="Acta Parasitologica" pagination="90-98" refId="B45" refString="Najer T, Sychra O, Literák I, Procházka P, Capek M, Koubek P (2012 b) Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides. Acta Parasitologica 57: 90–98. https://doi.org/10.2478/s11686-012-0005-x" title="Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides." volume="57">Najer et al.], 2012 b</bibRefCitation>
) (redrawn from original description to same scale as Fig.
<figureCitation id="D0021250BD971B595E23615E5CED57F8" captionStart="Figures 24, 25" captionStartId="F5" captionText="Figures 24, 25. Comparison in shape and proportions of head in species of Coronedax gen. n. 24 male head of Coronedax longiceps sp. n. 25 female head of Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 b) (redrawn from original description to same scale as Fig. 24). Setae and some other characters illustrated by Najer et al. (2012 b) have been omitted for clarity." figureDoi="10.3897/asp.82.e114351.figures24-25" httpUri="https://binary.pensoft.net/fig/1108932">24</figureCitation>
). Setae and some other characters illustrated by
<bibRefCitation id="2BCD4954D7B4AF0ACD7CD62FE23E8E2E" DOI="10.2478/s11686-012-0005-x" author="Najer T &amp; Sychra O &amp; Literák I &amp; Procházka P &amp; Capek M &amp; Koubek P" journalOrPublisher="Acta Parasitologica" pagination="90-98" refId="B45" refString="Najer T, Sychra O, Literák I, Procházka P, Capek M, Koubek P (2012 b) Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides. Acta Parasitologica 57: 90–98. https://doi.org/10.2478/s11686-012-0005-x" title="Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides." volume="57">Najer et al. (2012 b</bibRefCitation>
) have been omitted for clarity.
</paragraph>
</caption>
<paragraph id="7BC679EA6C1FBDE20ED4DAFB617A1DC2">
Possibly, head sensillum
<emphasis id="A4E94F400E2AE679042522A0BB965494" italics="true">s 4</emphasis>
is absent in
<taxonomicName id="00D7508B24D7FCD5AF0EE443BF7616B5" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="ED9F8558D20B9053E7FF4F7055080DA6" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
, but these sensilla are easily overlooked and as no specimen of
<taxonomicName id="36ED6364EC34D2C745F3C2D988BA4D9B" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="terpsiphoni">
<emphasis id="908A89D9DEF9BDA4E847DE53EE2E9DED" italics="true">C. terpsiphoni</emphasis>
</taxonomicName>
were examined, this cannot be verified;
<emphasis id="03A572F7BA550520D417D058AF835EBA" italics="true">s 4</emphasis>
is present in
<taxonomicName id="E9BD90C35E8BCF3B4DD9B68CBE837A77" authorityName="Ren &amp; Tian &amp; Grossi &amp; Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="771A3D25F1DEDC0D03F10EC1C4846373" italics="true">C. longiceps</emphasis>
</taxonomicName>
. Differences in the male genitalia between
<taxonomicName id="0EE5F3D31FCD38DCB2B6C4AFE5679F25" authorityName="Ren &amp; Tian &amp; Grossi &amp; Zou &amp; Gustafsson" authorityYear="2024" class="Insecta" family="Philopteridae" genus="Coronedax" kingdom="Animalia" order="Psocodea" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="61EB1F00FEFC1B41D8DE5D01216DC3A3" italics="true">C. longiceps</emphasis>
</taxonomicName>
and the illustrations of
<bibRefCitation id="85CF9A42480994A6E4689439ABAB648A" DOI="10.2478/s11686-012-0005-x" author="Najer T &amp; Sychra O &amp; Literák I &amp; Procházka P &amp; Capek M &amp; Koubek P" journalOrPublisher="Acta Parasitologica" pagination="90-98" refId="B45" refString="Najer T, Sychra O, Literák I, Procházka P, Capek M, Koubek P (2012 b) Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides. Acta Parasitologica 57: 90–98. https://doi.org/10.2478/s11686-012-0005-x" title="Chewing lice (Phthiraptera) from wild birds in Senegal, with descriptions of three new species of the genera Brueelia and Philopteroides." volume="57">Najer et al. (2012 b</bibRefCitation>
) (Figs
<figureCitation id="9A3A7B7AC4E79B194BF95FED3765771A" captionStart="Figures 2–17" captionStartId="F2" captionText="Figures 2–17. Comparison of the male genitalia of some species of Philopteroides Mey, 2004, Coronedax gen. n., and Stasiasticopterus gen. n. All figures redrawn from their original descriptions (Tandan 1955; Mey 2004; Valim &amp; Palma 2013), unless otherwise noted. Illustrations have been rescaled to be roughly the same size. Illustrations are in ventral view, unless otherwise noted. No illustrations of the genitalia have ever been published for Philopteroides lineatus (Giebel, 1874), Philopteroides mitsusui (Uchida, 1948), or Stasiasticopterus kayanobori (Uchida, 1948). 2 Philopteroides novaezelandiae Mey, 2004. 3 Philopteroides xenicus Mey, 2004. 4 Philopteroides fuliginosus Valim &amp; Palma, 2013. 5 Philopteroides macrocephalus Valim &amp; Palma, 2013. 6 Philopteroides gigas Najer et al., 2016. 7 Philopteroides sinancorellus Najer et al., 2016. 8 Philopteroides sclerotifrons (Tandan, 1955); no scale in original. 9 Philopteroides pilgrimi Valim &amp; Palma, 2013. 10 Philopteroides beckeri (Mey, 2004) (redrawn from Valim &amp; Palma 2013). 11 Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 a). 12 Coronedax longiceps sp. n. 13 Stasiasticopterus longiclypeatus (Gustafsson et al., 2022 b). 14 Stasiasticopterus holosternus (Gustafsson et al., 2022 b). 15 Stasiasticopterus haerixos (Gustafsson et al., 2022 b). 16 Stasiasticopterus flavala (Najer &amp; Sychra [in Najer et al.], 2012 a. 17 Stasiasticopterus cucphuongensis (Mey, 2004) (redrawn from Gustafsson et al. 2022 b). Bold grey lines signify the generic divisions used here; narrow grey lines signify groups that are morphologically different from the type species of the respective genera, but where there is insufficient data to propose addition (e. g., subgeneric) limits. The two species here considered incerta sedis are placed in separate groups, pending further investigations. — Abbreviations used: BA = basal apodeme; GP = gonopore; MS = mesosome; lpmes = lateral posterios mesosomal setae; PM = parameres; pst 1–2 = parameral setae 1–2 (2 distal to 1)." figureDoi="10.3897/asp.82.e114351.figures2-17" httpUri="https://binary.pensoft.net/fig/1108929">11</figureCitation>
,
<figureCitation id="C753E6F0711E42B7536DF322B0F6B52C" captionStart="Figures 2–17" captionStartId="F2" captionText="Figures 2–17. Comparison of the male genitalia of some species of Philopteroides Mey, 2004, Coronedax gen. n., and Stasiasticopterus gen. n. All figures redrawn from their original descriptions (Tandan 1955; Mey 2004; Valim &amp; Palma 2013), unless otherwise noted. Illustrations have been rescaled to be roughly the same size. Illustrations are in ventral view, unless otherwise noted. No illustrations of the genitalia have ever been published for Philopteroides lineatus (Giebel, 1874), Philopteroides mitsusui (Uchida, 1948), or Stasiasticopterus kayanobori (Uchida, 1948). 2 Philopteroides novaezelandiae Mey, 2004. 3 Philopteroides xenicus Mey, 2004. 4 Philopteroides fuliginosus Valim &amp; Palma, 2013. 5 Philopteroides macrocephalus Valim &amp; Palma, 2013. 6 Philopteroides gigas Najer et al., 2016. 7 Philopteroides sinancorellus Najer et al., 2016. 8 Philopteroides sclerotifrons (Tandan, 1955); no scale in original. 9 Philopteroides pilgrimi Valim &amp; Palma, 2013. 10 Philopteroides beckeri (Mey, 2004) (redrawn from Valim &amp; Palma 2013). 11 Coronedax terpsiphoni (Najer &amp; Sychra [in Najer et al.], 2012 a). 12 Coronedax longiceps sp. n. 13 Stasiasticopterus longiclypeatus (Gustafsson et al., 2022 b). 14 Stasiasticopterus holosternus (Gustafsson et al., 2022 b). 15 Stasiasticopterus haerixos (Gustafsson et al., 2022 b). 16 Stasiasticopterus flavala (Najer &amp; Sychra [in Najer et al.], 2012 a. 17 Stasiasticopterus cucphuongensis (Mey, 2004) (redrawn from Gustafsson et al. 2022 b). Bold grey lines signify the generic divisions used here; narrow grey lines signify groups that are morphologically different from the type species of the respective genera, but where there is insufficient data to propose addition (e. g., subgeneric) limits. The two species here considered incerta sedis are placed in separate groups, pending further investigations. — Abbreviations used: BA = basal apodeme; GP = gonopore; MS = mesosome; lpmes = lateral posterios mesosomal setae; PM = parameres; pst 1–2 = parameral setae 1–2 (2 distal to 1)." figureDoi="10.3897/asp.82.e114351.figures2-17" httpUri="https://binary.pensoft.net/fig/1108929">12</figureCitation>
) may not be diagnostic as much of the distal genitalia are largely soft, and thus affected by mounting.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E52AK" type="description">
<paragraph id="2194EE39BF6AE323904E8DDE8337ADBB">
<heading id="4454D950B09400734BFAB766E540630A" reason="title">Description.</heading>
</paragraph>
<paragraph id="3EE7B12C14AECA6E823AA07EAA147F29">
Head structure and chaetotaxy as in Fig.
<figureCitation id="7959A6F0DCBDABB31F69C8AFE77B4518" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">20</figureCitation>
; head long and slender; dorsal anterior plate slender (Fig.
<figureCitation id="18935E156C3B18FE17E1C1729BDC4C9F" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">20</figureCitation>
); head sensillum
<emphasis id="542C60A62D202399473FB2F787F6D2D3" italics="true">s 4</emphasis>
present;
<abbrev id="ABBRID0EP3AK" xlink_title="ocular seta">
<emphasis id="57E49AAB435E64B1E09EEE511E6B32BB" italics="true">os</emphasis>
</abbrev>
much shorter than
<emphasis id="9D3F445DB04000BB4E0D77393DAA6747" italics="true">mts 3</emphasis>
;
<abbrev id="ABBRID0EW3AK" xlink_title="marginal temporal setae 1, 3">
<emphasis id="D032A7A4797BB39D3A720283E4E8A798" italics="true">mts 1</emphasis>
</abbrev>
intermediate in length between
<abbrev id="ABBRID0E23AK" xlink_title="ocular seta">
<emphasis id="05B0617F935F5EEE929AB09A4C688774" italics="true">os</emphasis>
</abbrev>
and
<emphasis id="E99ACD2265785CCC9EE7762CCF438A3D" italics="true">mts 3</emphasis>
. Thoracic and abdominal segments and chaetotaxy as in Figs
<figureCitation id="6E00D1F91DBF7E69924755D263DD3AFC" captionStart="Figures 18, 19" captionStartId="F3" captionText="Figures 18, 19. Coronedax longiceps sp. n. 18 male habitus, dorsal and ventral views. 19 female habitus, dorsal and ventral views. — Abbreviations used: mms = marginal mesometanotal setae; ppss = pronotal post-spiracular seta; ps = paratergal setae; psps = principal post-spiracular setae; ss = sutural setae; sts = sternal setae; tps = tergal posterior setae." figureDoi="10.3897/asp.82.e114351.figures18-19" httpUri="https://binary.pensoft.net/fig/1108930">18</figureCitation>
,
<figureCitation id="0D98394116B52BA325BDA0EE2A4D81A2" captionStart="Figures 18, 19" captionStartId="F3" captionText="Figures 18, 19. Coronedax longiceps sp. n. 18 male habitus, dorsal and ventral views. 19 female habitus, dorsal and ventral views. — Abbreviations used: mms = marginal mesometanotal setae; ppss = pronotal post-spiracular seta; ps = paratergal setae; psps = principal post-spiracular setae; ss = sutural setae; sts = sternal setae; tps = tergal posterior setae." figureDoi="10.3897/asp.82.e114351.figures18-19" httpUri="https://binary.pensoft.net/fig/1108930">19</figureCitation>
. Male abdominal chaetotaxy:
<emphasis id="7FB166B07CED0E7CC4D70822AD959BB9" italics="true">ss</emphasis>
present on segments II – VIII;
<abbrev id="ABBRID0EM4AK" xlink_title="tergal posterior seta">
<emphasis id="EF50F0CFFC01AF0689CF472B4D2D6EF3" italics="true">tps</emphasis>
</abbrev>
present on segments II – VIII (2 on each side on II – VI, 1 on each side on VII – VII);
<abbrev id="ABBRID0ER4AK" xlink_title="principal postspiracular seta">
<emphasis id="1B989BF534E711D0678490EB330CC5B4" italics="true">psps</emphasis>
</abbrev>
present on segments III – VII;
<abbrev id="ABBRID0EW4AK" xlink_title="accessory postspiracular seta">
<emphasis id="3F59A85E8831A8BE16EC37B1F451397A" italics="true">aps</emphasis>
</abbrev>
absent;
<abbrev id="ABBRID0E24AK" xlink_title="paratergal seta">
<emphasis id="CD00492D5E6ED8143E5BB339569C2C69" italics="true">ps</emphasis>
</abbrev>
present on segments IV – VIII. Female abdominal chaetotaxy:
<emphasis id="D7D10C50457C22DD0DD1B27E1769C741" italics="true">ss</emphasis>
present on segments II – VIII;
<abbrev id="ABBRID0EC5AK" xlink_title="tergal posterior seta">
<emphasis id="DD552E7822F7C26801935A03E06151CB" italics="true">tps</emphasis>
</abbrev>
present on segments II – VIII (1 on each side);
<abbrev id="ABBRID0EH5AK" xlink_title="principal postspiracular seta">
<emphasis id="C9E6B3019E3B73C215196A90E9699EDA" italics="true">psps</emphasis>
</abbrev>
present on segments III – VII;
<abbrev id="ABBRID0EM5AK" xlink_title="accessory postspiracular seta">
<emphasis id="8C3D47C1E10E752F3A904DCE41E72DAB" italics="true">aps</emphasis>
</abbrev>
absent;
<abbrev id="ABBRID0ER5AK" xlink_title="paratergal seta">
<emphasis id="CA938AC397D8349C04D6DEDD3294335A" italics="true">ps</emphasis>
</abbrev>
present on segments IV – VIII. Central sternal plates present on segments II – VI in both sexes; accessory sternal plates absent in male (but some central sternal plates with modified lateral ends) and present on segments III – VI in female. Each of sternal plates II – VI in both sexes with 1 thorn-like and 1 normal seta on each side; on segments II – V thorn-like setae median to normal setae, but on VI thorn-like setae lateral to normal setae. Basal apodeme slender, not conspicuously thickened laterally, bulging distally (Fig.
<figureCitation id="4776F0E7F7C9500C4E069D72CB6A3EC6" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">21</figureCitation>
). Mesosome as mainly dorsal thickening of distal mesosome centrally; 2
<abbrev id="ABBRID0E15AK" xlink_title="lateral posterior mesosomal seta">
<emphasis id="D678929A008D736F58D60395E554D3A9" italics="true">lpmes</emphasis>
</abbrev>
microsetae visible on each side. Gonopore reduced, extending slightly beyond distal margin of basal apodeme, with slight nodi distally (Fig.
<figureCitation id="13527DF66AA9D5F539EFC58B01CD3F9B" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">22</figureCitation>
). Parameres completely fused to basal apodeme proximally, soft and flexible, may be displaced dorsally;
<abbrev id="ABBRID0ED6AK" xlink_title="parameral setae 1–2">
<emphasis id="C4AEE44D35E7A8ABFF4F09606B4668D4" italics="true">pst 1</emphasis>
</abbrev>
sensillum,
<emphasis id="CEA5A074BCF25D66EA0CC6AA67EBE704" italics="true">pst 2</emphasis>
distal seta, as long as paramere. Female subgenital plate with triangular extension distally (Fig.
<figureCitation id="6D98027A50919E8FA6F65AA5A939BA3D" captionStart="Figures 20–23" captionStartId="F4" captionText="Figures 20–23. Coronedax longiceps sp. n. 20 male head, dorsal and ventral views. 21 male genitalia, dorsal view. 22 male genitalia, ventral view. 23 female subgenital plate, vulval margin, and post-vulval area, ventral view. Male genitalia are illustrated asymmetrically as in holotype specimen, to indicate the “ looseness ” of the parameres. — Abbreviations used: as 1 = anterior seta 1; lpmes = lateral posterior mesosomal seta; mts 1, 3 = marginal temporal setae 1, 3; os = ocular seta; pst 1–2 = parameral setae 1–2; pts = posttemporal seta; s 1–4 = sensilla 1–4 of dorsal postantennal head; vms = vulval marginal setae." figureDoi="10.3897/asp.82.e114351.figures20-23" httpUri="https://binary.pensoft.net/fig/1108931">23</figureCitation>
). Vulval margin bulging, with 2
<abbrev id="ABBRID0EO6AK" xlink_title="vulval marginal setae">
<emphasis id="7C4A2DDA779695E9AED974F3077286C4" italics="true">vms</emphasis>
</abbrev>
mesosetae on each side. Subvulval plates roughly triangular.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES6AK" type="description">
<paragraph id="C1CAC381F301A648FB52570F2F4D7ACF">
<heading id="3166A0562AF710B66E24521E8450CAE6" reason="title">Measurements.</heading>
</paragraph>
<paragraph id="A9723ADDDE48ED53D85B444C756407A7">
Male (n = 3, except for
<abbrev id="ABBRID0EY6AK" xlink_title="total length">TL</abbrev>
, where n = 1 and PW where n = 2):
<abbrev id="ABBRID0E36AK" xlink_title="total length">TL</abbrev>
= 1.26;
<abbrev id="ABBRID0EAABK" xlink_title="head length">HL</abbrev>
= 0.39–0.41;
<abbrev id="ABBRID0EEABK" xlink_title="head width">HW</abbrev>
= 0.35–0.37;
<abbrev id="ABBRID0EIABK" xlink_title="prothoracic width">PRW</abbrev>
= 0.22–0.26;
<abbrev id="ABBRID0EMABK" xlink_title="pterothoracic width">PTW</abbrev>
= 0.31–0.35;
<abbrev id="ABBRID0EQABK" xlink_title="abdominal width">AW</abbrev>
= 0.41–0.48. Female (n = 1, total length not measured due to breakage in pterothorax):
<abbrev id="ABBRID0EUABK" xlink_title="head length">HL</abbrev>
= 0.42;
<abbrev id="ABBRID0EYABK" xlink_title="head width">HW</abbrev>
= 0.37;
<abbrev id="ABBRID0E3ABK" xlink_title="prothoracic width">PRW</abbrev>
= 0.25;
<abbrev id="ABBRID0EABBK" xlink_title="pterothoracic width">PTW</abbrev>
= 0.30;
<abbrev id="ABBRID0EEBBK" xlink_title="abdominal width">AW</abbrev>
= 0.41.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIBBK" type="etymology">
<paragraph id="E8EE0081CBE8472490FB0FDD9D1D3DA1">
<heading id="24436EEC5CAAEBE223941AD09B38A0A3" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C82286839976D9304EC556BC3BF915E9">
Specific name derived from “
<emphasis id="7D7E1777456DFFA540BAE1131794E129" italics="true">longus</emphasis>
”, Latin for “ long ”, and “ -
<emphasis id="98AB763C69DA1474A465CC71706D54F6" italics="true">ceps</emphasis>
”, Latin for “ - headed ”, referring to the relatively long head of this species compared to the only other known member of the genus.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESBBK" type="Type specimens">
<paragraph id="C71405586E1EFC346ED4ED2FD5A0B079">
<heading id="B543C3B3FCF2734D8BB577FC15D026DF" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="DECC902E2042F7013767E90094EA22B5">
<materialsCitation id="8AE05B6E7A136D8692C8B8F6ADFFBE94" collectedFrom="primary forest" collectingDate="2013-06-06" collectionCode="IZGAS" collectorName="D. Su &amp; Y. Zhao" country="China" county="Mengla County" location="Xinhuikuan and Manpa villages" municipality="Banna Prefecture" specimenCode="GD-PHTH- 901" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="93003415ED2D3D9D61DBC03F5B191354" bold="true">
<typeStatus id="11B1D1F7F7FA9408CABE0A4804237E0F" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="786C257E0D86384A46EE246331C0DB80" count="1" type="male">♂</specimenCount>
,
<collectingCountry id="9ECB19A4C82C2D4323E4DA2BB4192E3A" name="China">CHINA</collectingCountry>
:
<collectingRegion id="D81C2AF964129B47960C19B68716A52A" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="475600008DE807FDD900A65F127560FE">Banna Prefecture</collectingMunicipality>
,
<collectingCounty id="3553F17221769915F176985BC8A14360">Mengla County</collectingCounty>
,
<collectedFrom id="6551C92763D6B1CD7F40B18B14A991E9">primary forest</collectedFrom>
near
<location id="6C854AD086A6FBA2BE13AEDBC4B10727" LSID="urn:lsid:plazi:treatment:BEABE1D94E44581BBD9D065390D913E2:6C854AD086A6FBA2BE13AEDBC4B10727" country="China" county="Mengla County" municipality="Banna Prefecture" name="Xinhuikuan and Manpa villages" stateProvince="Yunnan Province">Xinhuikuan and Manpa villages</location>
,
<date id="A78734A5B93AA073E88B895F0F1DF97B" value="2013-06-06">
<collectingDate id="9EDF748D589DFE62EAC37DBA61A9361A" value="2013-06-06">6 Jun. 2013</collectingDate>
</date>
, coll.
<collectorName id="5B194E4BEA965F411003B4389DE7A6A9">D. Su</collectorName>
&amp;
<collectorName id="6F26817B6C2AF648BA6B0AA9DF4DAF1A">Y. Zhao</collectorName>
, bird ID: J 1124, louse ID:
<specimenCode id="9F852F869C97DA9A107202D4066F9A63">GD-PHTH- 901</specimenCode>
(
<collectionCode id="CF70993E9FFE9E02D995EF59E880F340" country="Georgia" httpUri="http://grbio.org/cool/ahe1-tsbj" name="Georgian Academy of Sciences, Insititute of Zoology" type="">IZGAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="CB35D512810D64A97F0437458F40FD07" collectedFrom="primary forest" collectingDate="2013-06-06" collectionCode="IZGAS" collectorName="D. Su &amp; Y. Zhao" country="China" county="Mengla County" location="Xinhuikuan and Manpa villages" municipality="Banna Prefecture" specimenCode="GD-PHTH-00899" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan Province" typeStatus="paratype">
<emphasis id="BB450E4BBA2D39F5657A2634AF1A560C" bold="true">
<typeStatus id="A0CDAF1ECA474CB2F45C2688E600726E" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="6DB7E3B1638371E6911E8A708E2D6780" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="F0457DDA22555565ECB8A449CE1A89BE" count="1" type="female">1 ♀</specimenCount>
, same data as holotype, louse ID:
<specimenCode id="AC824E82301BEA4228AA18C8BEDFEF6D">GD-PHTH-00899</specimenCode>
–900, 902 (
<collectionCode id="2521E7C805592FC945FF8CD09516FC5B" country="Georgia" httpUri="http://grbio.org/cool/ahe1-tsbj" name="Georgian Academy of Sciences, Insititute of Zoology" type="">IZGAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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