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<document ID-CLB-Dataset="176497" ID-DOI="http://dx.doi.org/10.3897/zse.98.93851" ID-GBIF-Dataset="cf500229-a971-4afd-8eb6-42aa34a380d1" ID-Pensoft-Pub="1860-0743-2-427" ID-Pensoft-UUID="367FFEC1B6A856EEB610C78A492EE8B3" ID-ZooBank="08CD0830C6574D7AA8C48049BBA3E7A5" ModsDocID="1860-0743-98-2-427" checkinTime="1667237297416" checkinUser="pensoft" docAuthor="Koehler, Frank &amp; Shea, Michael" docDate="2022" docId="BEAB224A3FAE57139E364A2F6570ABDD" docLanguage="en" docName="ZoosystEvol 98(2): 427-433" docOrigin="Zoosystematics and Evolution 98 (2)" docPubDate="2022-10-31" docSource="http://dx.doi.org/10.3897/zse.98.93851" docTitle="Parglogenia pelodes" docType="treatment" docVersion="5" id="367FFEC1B6A856EEB610C78A492EE8B3" lastPageNumber="427" masterDocId="367FFEC1B6A856EEB610C78A492EE8B3" masterDocTitle="A new cryptic species of land snail from the Northern Territory, Australia (Stylommatophora, Camaenidae, Parglogenia)" masterLastPageNumber="433" masterPageNumber="427" pageNumber="427" updateTime="1732938673549" updateUser="ExternalLinkService">
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<mods:title id="7FAC79893B56700D18C08AA3B3C966E3">A new cryptic species of land snail from the Northern Territory, Australia (Stylommatophora, Camaenidae, Parglogenia)</mods:title>
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<mods:namePart id="1FD96CF0B3966ADB221D9D397F8431CA">Koehler, Frank</mods:namePart>
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<mods:affiliation id="F6697817ECD6662A144F05F10FA38900">Australian Museum, Sydney, Australia</mods:affiliation>
<mods:nameIdentifier id="4485CAD7678063ADED9BD4123970A7CC" type="email">frank.koehler@austmus.gov.au</mods:nameIdentifier>
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<mods:namePart id="A6A361A094498B7B772E34E43A330E26">Shea, Michael</mods:namePart>
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<mods:affiliation id="AF00D1DBF6E4539025862E6E9EBF5F2E">Australian Museum, Sydney, Australia</mods:affiliation>
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<mods:title id="C9B4A0D8BB2FAB4A4E25D26B8567B5B9">Zoosystematics and Evolution</mods:title>
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<mods:number id="40BF9FF7259974143FBB0BCFAC06644E">98</mods:number>
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<treatment id="BEAB224A3FAE57139E364A2F6570ABDD" ID-GBIF-Taxon="203712419" LSID="urn:lsid:plazi:treatment:BEAB224A3FAE57139E364A2F6570ABDD" httpUri="http://treatment.plazi.org/id/BEAB224A3FAE57139E364A2F6570ABDD" lastPageNumber="427" pageId="0" pageNumber="427" scope_family="Camaenidae" scope_order="Stylommatophora">
<subSubSection id="20EA47C33E8570809823FEBDDF9CFD8B" pageId="0" pageNumber="427" type="nomenclature">
<paragraph id="A07E861EC9B37B485B6F6F4AAA1F266B" pageId="0" pageNumber="427">
<taxonomicName id="F94401CE6C0B58C827E57FFBE0D52679" ID-CoL="6TRJP" LSID="BEAB224A-3FAE-5713-9E36-4A2F6570ABDD" authority="(Pfeiffer, 1846)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1846" class="Gastropoda" family="Camaenidae" genus="Parglogenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parglogenia pelodes" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pelodes">Parglogenia pelodes (Pfeiffer, 1846)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="011E1732D5CE0028B40DBF4B37EEF593" pageId="0" pageNumber="427" type="reference_group">
<paragraph id="D41BD75199056FB90879D7CA8F5142BA" pageId="0" pageNumber="427">
<taxonomicName id="D475871CB2B8AFCC373F78FB27AA469E" authorityName="L. Pfeiffer" authorityYear="1846" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix pelodes" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pelodes">Helix pelodes</taxonomicName>
Pfeiffer, 1846: 126 ('north coast of
<normalizedToken id="26DB8104569C6E95533A6B0B9B977347" originalValue="Australia’">Australia'</normalizedToken>
). Neotype FMNH 182450, Darwin, Dudley Point (
<geoCoordinate id="95D3538B0DE22D3DF984E006E37E07AB" degrees="12" direction="south" minutes="24" orientation="latitude" precision="925" value="-12.4">12°24'S</geoCoordinate>
,
<geoCoordinate id="54E04AC26B0FFB7D3E040794E4C54DBF" degrees="130" direction="east" minutes="49" orientation="longitude" precision="925" value="130.81667">130°49'E</geoCoordinate>
), designated by
<bibRefCitation id="A025717FEE48D45BC8F3C019AAC21778" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="37CE58D981765ED4DE4AE0077CB6D32E" originalValue="Köhler">Koehler</normalizedToken>
(2012)
</bibRefCitation>
.
</paragraph>
<paragraph id="8A4EF54C15CCF3B2801D22EEAB0EB145" pageId="0" pageNumber="427">
<taxonomicName id="CBD9733FCBD8C0AA058424BF00854110" authorityName="Ferussac" authorityYear="1821" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix prunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="prunum">Helix prunum</taxonomicName>
- Pfeiffer 1851 [in 1849-1853]: 54, pl. 74, figs 11-13; Reeve 1852 [in 1851-1854]: pl. 68, fig. 353;
<bibRefCitation id="31C376017FAFF4A1A81E5529A5981D07" DOI="https://doi.org/10.5962/bhl.title.1292" author="Cox, JC" journalOrPublisher="Maddock, Sydney" pageId="0" pageNumber="427" refId="B2" refString="Cox, JC, 1868. A monograph of Australian land shells. Maddock, Sydney, DOI: https://doi.org/10.5962/bhl.title.1292" title="A monograph of Australian land shells." url="https://doi.org/10.5962/bhl.title.1292" year="1868">Cox 1868</bibRefCitation>
: 43, pl. 4, fig. 6 (not
<taxonomicName id="08F79ABC1F9D3C0091B8CE999A0E6DD8" authorityName="Ferussac" authorityYear="1821" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix prunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="prunum">Helix prunum</taxonomicName>
<normalizedToken id="5072C7DD5C6836809A18D03386B33924" originalValue="Férussac">Ferussac</normalizedToken>
, 1821).
</paragraph>
<paragraph id="99753E7D8891F66889C20F94B384E8BB" pageId="0" pageNumber="427">
<taxonomicName id="3A680638465FBE5A5F8A061A8FAF897B" authorityName="Pilsbry" authorityYear="1893" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis pseudoprunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pseudoprunum">Chloritis pseudoprunum</taxonomicName>
Pilsbry, 1893: 271 ('Port
<normalizedToken id="3BBE45A0FCC43307F0C5D7809C342A1F" originalValue="Darwin’">Darwin'</normalizedToken>
; Holotype ANSP 62411).
</paragraph>
<paragraph id="B02343C647548A478A4762882AAF014C" pageId="0" pageNumber="427">
<taxonomicName id="80A26E2746F534C62C376A1BAB67B9E4" baseAuthorityName="Pfeiffer" baseAuthorityYear="1846" class="Gastropoda" family="Camaenidae" genus="Parglogenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parglogenia pelodes" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pelodes">Parglogenia pelodes</taxonomicName>
-
<bibRefCitation id="EE04D34E4603304E4C2DC63E8D6042DA" author="Iredale, T" journalOrPublisher="Australian Zoologist" pageId="0" pageNumber="427" pagination="83 - 124" refId="B6" refString="Iredale, T, 1938. A basic list of the land Mollusca of Australia. Part III. Australian Zoologist 9: 83 - 124" title="A basic list of the land Mollusca of Australia. Part III." volume="9" year="1938">Iredale 1938</bibRefCitation>
: 99;
<bibRefCitation id="F5AE3A8EB40F9147F7A6981C06DBD645" author="Solem, A" journalOrPublisher="7. Reeve, London" pageId="0" pageNumber="427" refId="B21" refString="Solem, A, 1979. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5-142." title="Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5 - 142." year="1979">Solem 1979</bibRefCitation>
: 128-133, fig. 32, pls 11a, b.
</paragraph>
</subSubSection>
<subSubSection id="2C7628D48E532BBE6A2E8FFF8C7C60D7" pageId="0" pageNumber="427" type="taxonomic remarks">
<paragraph id="FF2B18BFCF8716CA59B3872D513CDE6D" pageId="0" pageNumber="427">Taxonomic remarks.</paragraph>
<paragraph id="E792C25AEDEFEDD05D2896DF568FBCD6" pageId="0" pageNumber="427">
Since its original description, the name
<taxonomicName id="8354C929669C9E2393A6BE1C8EF0B487" authorityName="L. Pfeiffer" authorityYear="1846" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix pelodes" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pelodes">
<emphasis id="083875A5BA93F68ABE905F995DD5546B" italics="true" pageId="0" pageNumber="427">Helix pelodes</emphasis>
</taxonomicName>
has been shrouded in uncertainty until relatively recently. Pfeiffer (1851 [in 1849-1853]), subsequently treated
<taxonomicName id="12A09970E817355C95783C0ACA939627" lsidName="H. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="C7A45B87090548A635BD624AC9CBEB1B" italics="true" pageId="0" pageNumber="427">H. pelodes</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="266EA5437C70421269E956BDBFC4782F" authorityName="Ferussac" authorityYear="1821" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix prunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="prunum">
<emphasis id="A11906DB0FE6A2CE062DA175665D696B" italics="true" pageId="0" pageNumber="427">Helix prunum</emphasis>
</taxonomicName>
<normalizedToken id="D08CC7CCD0F5A33BE0EDD25A6330F54D" originalValue="Férussac">Ferussac</normalizedToken>
, 1821. Subsequently,
<bibRefCitation id="0D9CBA7D1B1432110A5178315ED9431F" author="Pilsbry, HA" editor="Tryon, W" journalOrPublisher="Second series, Pulmonata. Vol. 6. Conchological Section, Academy of Natural Sciences, Philadelphia" pageId="0" pageNumber="427" pagination="1 - 324" refId="B18" refString="Pilsbry, HA, 1890. Helicidae. Vol. IV. In: Tryon, W, Pilsbry, HA, Eds., Manual of conchology, structural and systematic with illustrations of the species. Second series, Pulmonata. Vol. 6. Conchological Section, Academy of Natural Sciences, Philadelphia: 1 - 324" title="Helicidae. Vol. IV." volumeTitle="Manual of conchology, structural and systematic with illustrations of the species." year="1890">Pilsbry (1890</bibRefCitation>
: 135) agreed with this synonymy, but accepted
<taxonomicName id="24D6C429383CAE81D1A56AAC7C34AFE1" lsidName="H. prunum" pageId="0" pageNumber="427" rank="species" species="prunum">
<emphasis id="BEDAB6631FF5203DEB22A7911957FC20" italics="true" pageId="0" pageNumber="427">H. prunum</emphasis>
</taxonomicName>
as an eastern Australian species with uncertain systematic relationships. Based on this presumption, he considered that all previous authors had misapplied the name
<taxonomicName id="BEE47FB730450D8716B495F5BAE7792C" lsidName="H. prunum" pageId="0" pageNumber="427" rank="species" species="prunum">
<emphasis id="BF5426A3FE32742265A0DAE13156C51F" italics="true" pageId="0" pageNumber="427">H. prunum</emphasis>
</taxonomicName>
for an unnamed species from Arnhem Land, for which
<bibRefCitation id="2EF9EED2079B6C484BA683881EC79F91" author="Pilsbry, HA" editor="Tryon, W" journalOrPublisher="Second series: Pulmonata. Vol. 8. Conchological Section, Academy of Natural Sciences, Philadelphia" pageId="0" pageNumber="427" pagination="1 - 314" refId="B19" refString="Pilsbry, HA, 1893. Helicidae. Vol. VI. In: Tryon, W, Pilsbry, HA, Eds., Manual of Conchology, structural and systematic, with illustrations of the species. Second series: Pulmonata. Vol. 8. Conchological Section, Academy of Natural Sciences, Philadelphia: 1 - 314" title="Helicidae. Vol. VI." volumeTitle="Manual of Conchology, structural and systematic, with illustrations of the species." year="1893">Pilsbry (1893)</bibRefCitation>
introduced the new name
<taxonomicName id="40EFF6DBFEC7B4BA4A6FD0559142CAB6" authorityName="Pilsbry" authorityYear="1893" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis pseudoprunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pseudoprunum">
<emphasis id="12DA48399A3354AD05E96F239BF9217C" italics="true" pageId="0" pageNumber="427">Chloritis pseudoprunum</emphasis>
</taxonomicName>
. By contrast,
<bibRefCitation id="0874E3109187AB282A109B6C62E5F260" author="Iredale, T" journalOrPublisher="Australian Zoologist" pageId="0" pageNumber="427" pagination="83 - 124" refId="B6" refString="Iredale, T, 1938. A basic list of the land Mollusca of Australia. Part III. Australian Zoologist 9: 83 - 124" title="A basic list of the land Mollusca of Australia. Part III." volume="9" year="1938">Iredale (1938)</bibRefCitation>
and
<bibRefCitation id="7F5CE2EFA2205CC0EFF02F3E80831933" author="Solem, A" journalOrPublisher="7. Reeve, London" pageId="0" pageNumber="427" refId="B21" refString="Solem, A, 1979. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5-142." title="Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5 - 142." year="1979">Solem (1979)</bibRefCitation>
considered that the supposed synonymy of
<taxonomicName id="2374899D86F065B0FBE151109881C92A" lsidName="H. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="B91935A01FB574F2A97EA8FDC728E225" italics="true" pageId="0" pageNumber="427">H. pelodes</emphasis>
</taxonomicName>
with
<taxonomicName id="99B644FB96E40F25847D4211FD8889BA" lsidName="H. prunum" pageId="0" pageNumber="427" rank="species" species="prunum">
<emphasis id="F1B6E7606F4AA75B78522A4D9906473F" italics="true" pageId="0" pageNumber="427">H. prunum</emphasis>
</taxonomicName>
was in error. While
<taxonomicName id="59A1F2A000575437435542BBC400E0BE" lsidName="H. prunum" pageId="0" pageNumber="427" rank="species" species="prunum">
<emphasis id="3E3AE21FE4A522D588662A61B64B0A0D" italics="true" pageId="0" pageNumber="427">H. prunum</emphasis>
</taxonomicName>
continues to be a taxonomic enigma to this date (
<bibRefCitation id="889414C5C5F9A1BF13A29D39A639DCDC" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="5D2FE08F53D026C584F1511159145AB2" originalValue="Köhler">Koehler</normalizedToken>
2012
</bibRefCitation>
),
<taxonomicName id="61AC76AD8E3D401724C65BBC1B55CB4A" authorityName="L. Pfeiffer" authorityYear="1846" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix pelodes" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pelodes">
<emphasis id="3471DCFA5BA9A36B6E38928F3D2F59D8" italics="true" pageId="0" pageNumber="427">Helix pelodes</emphasis>
</taxonomicName>
was removed from its synonymy and treated as an accepted species, for which
<bibRefCitation id="05F847F1F4ECEA6B4A685DCC06E09080" author="Iredale, T" journalOrPublisher="Australian Zoologist" pageId="0" pageNumber="427" pagination="83 - 124" refId="B6" refString="Iredale, T, 1938. A basic list of the land Mollusca of Australia. Part III. Australian Zoologist 9: 83 - 124" title="A basic list of the land Mollusca of Australia. Part III." volume="9" year="1938">Iredale (1938)</bibRefCitation>
described the genus
<taxonomicName id="A72A8AA74968616D1DC1C0C112096153" authorityName="Iredale" authorityYear="1938" class="Gastropoda" family="Camaenidae" genus="Parglogenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parglogenia" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="genus">
<emphasis id="FB42398A7EDCBE1B48254C9C60681AC1" italics="true" pageId="0" pageNumber="427">Parglogenia</emphasis>
</taxonomicName>
. This treatment rendered
<taxonomicName id="352BBCBA9022F38BA357269FFD1C1CA4" lsidName="C. pseudoprunum" pageId="0" pageNumber="427" rank="species" species="pseudoprunum">
<emphasis id="6BF02ACD3083FAF7C70DEBF9D824AAA5" italics="true" pageId="0" pageNumber="427">C. pseudoprunum</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="5FE3EF0FF9E1F55B1D07F69831F53349" lsidName="P. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="8E4F2723E8C5F4E838A5DB71D7182843" italics="true" pageId="0" pageNumber="427">P. pelodes</emphasis>
</taxonomicName>
.
<bibRefCitation id="78943CBC449F3523B573E9EDF7A487FE" author="Iredale, T" journalOrPublisher="Australian Zoologist" pageId="0" pageNumber="427" pagination="83 - 124" refId="B6" refString="Iredale, T, 1938. A basic list of the land Mollusca of Australia. Part III. Australian Zoologist 9: 83 - 124" title="A basic list of the land Mollusca of Australia. Part III." volume="9" year="1938">Iredale (1938)</bibRefCitation>
included a second species in this genus,
<taxonomicName id="ACFE116D8D2D2CB786E463E2C5AB793A" lsidName="P. subgranosa" pageId="0" pageNumber="427" rank="species" species="subgranosa">
<emphasis id="AB7243127812910FCCEED3E25D5ADD1C" italics="true" pageId="0" pageNumber="427">P. subgranosa</emphasis>
</taxonomicName>
(Le Guillou, 1842).
<bibRefCitation id="C9F81331ACB705FFF5801D2A5999877B" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="83F696024EAB59FAED1BA47A286A3320" originalValue="Köhler">Koehler</normalizedToken>
(2012)
</bibRefCitation>
rejected this treatment, arguing that the type of
<taxonomicName id="07F50885170677B5FD6BF4AFBBB7235A" authorityName="Le Guillou" authorityYear="1842" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix subgranosa" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="subgranosa">
<emphasis id="E9246CDEB70C2425B7E3CF4B9F0B2331" italics="true" pageId="0" pageNumber="427">Helix subgranosa</emphasis>
</taxonomicName>
represented a juvenile shell of
<taxonomicName id="549D0135103C4892F1EEF34195CD5B44" baseAuthorityName="Hombron &amp; Jacquinot" baseAuthorityYear="1841" class="Gastropoda" family="Camaenidae" genus="Xanthomelon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xanthomelon durvillii" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="durvillii">
<emphasis id="DD8730D48CBE2925E886491364B4EDAF" italics="true" pageId="0" pageNumber="427">Xanthomelon durvillii</emphasis>
</taxonomicName>
(Hombron &amp; Jacquinot, 1841). However,
<bibRefCitation id="61E68F58B966E76FD4C072A311FE0512" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="26942FFA302B39CE7F5B68EE865F5157" originalValue="Köhler">Koehler</normalizedToken>
(2012)
</bibRefCitation>
maintained
<taxonomicName id="10C4CC12424F7AA661959842A93D778E" lsidName="H. subgranosa" pageId="0" pageNumber="427" rank="species" species="subgranosa">
<emphasis id="ED47158E94BD92D992EEB3FFA3F43A73" italics="true" pageId="0" pageNumber="427">H. subgranosa</emphasis>
</taxonomicName>
as a
<emphasis id="B21D9E0FD26CC40C84459698E960FF31" italics="true" pageId="0" pageNumber="427">nomen inquirendum</emphasis>
. To remove the remaining ambiguity,
<taxonomicName id="D9D75E47587680D44ABDAFF17624F03E" authorityName="Le Guillou" authorityYear="1842" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix subgranosa" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="subgranosa">
<emphasis id="75A07626CEBAB3326817E507364B5691" italics="true" pageId="0" pageNumber="427">Helix subgranosa</emphasis>
</taxonomicName>
is herewith placed in the synonymy of
<taxonomicName id="E2A8C4AC2059552DD056269F2444EE47" lsidName="X. durvillii" pageId="0" pageNumber="427" rank="species" species="durvillii">
<emphasis id="79D253D1B1FD1E545BCF99EFA68CCB05" italics="true" pageId="0" pageNumber="427">X. durvillii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BAA5A9ED58D58BB18FF48354DFB52EFD" pageId="0" pageNumber="427" type="diagnosis">
<paragraph id="45527A7300D38F797BF68D32A968CC75" pageId="0" pageNumber="427">Diagnosis.</paragraph>
<paragraph id="6FB42CC1377D749C5422C0435C406556" pageId="0" pageNumber="427">
Shell relatively large (D = 14.3-31.5 mm, H = 10.8-20.3 mm; n = 92), with strongly and almost evenly elevated spire, comprising between 4.5 and 5.8 whorls. Apical sculpture of anastomosing ridges initially, becoming pustulated after first half whorl. Postapical whorls with microsculpture of very small, rather widely spaced setae and extremely fine ridgelets with weak radial ribs appearing on spire and body whorl. Umbilicus very narrow, partly covered by reflected lip, internally with crowded pustules. Body whorl globose, rounded, only slightly descending behind strongly reflected, thin, white lip. Shell light yellow brown, uniform. Vagina and penis very long, atrium short, bursa copulatrix very short, without enlarged head. Free oviduct with glandular, convoluted walls. Vas deferens with bifurcated caecum on ascending arm and entering the epiphallus near insertion of penial retractor muscle without differentiation. A narrow, raised ridge separates the epiphallus from the penis; inner penial walls with longitudinal corrugated ridges; no penial sheath, verge, or epiphallic appendages present (
<bibRefCitation id="19742EDE2CD287CE4E50CB4ED9F78F94" author="Solem, A" journalOrPublisher="7. Reeve, London" pageId="0" pageNumber="427" refId="B21" refString="Solem, A, 1979. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5-142." title="Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5 - 142." year="1979">Solem 1979</bibRefCitation>
;
<bibRefCitation id="E7824EDB35A333D0A115A53ECF54F407" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="8D2BC18C733D88664478DF656AF2D923" originalValue="Köhler">Koehler</normalizedToken>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1EA6A9D0CED5F2105EC29FA8FF62F490" pageId="0" pageNumber="427" type="comparative remarks">
<paragraph id="F732D57984F64675A25A31B23154F890" pageId="0" pageNumber="427">Comparative remarks.</paragraph>
<paragraph id="28A22FD5385978DFA880736D594E7FBF" pageId="0" pageNumber="427">
<bibRefCitation id="B44183685810D40BFA8D78EC9A9744C2" author="Solem, A" journalOrPublisher="7. Reeve, London" pageId="0" pageNumber="427" refId="B21" refString="Solem, A, 1979. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5-142." title="Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). I. Taxa with trans-Australian distribution. Records of the Western Australian Museum (Suppl. 10): 5 - 142." year="1979">Solem (1979)</bibRefCitation>
provided a detailed description of the shell and reproductive anatomy of this species based on examination of specimens from near Darwin. He remarked that the periostracal setae in
<taxonomicName id="5C05EC1BDBD66A88985947C15D62A570" lsidName="P. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="06239A28D86AAEEBA62E9BA14FFDCA23" italics="true" pageId="0" pageNumber="427">P. pelodes</emphasis>
</taxonomicName>
were similar to those in
<taxonomicName id="573374427FB1D3DD5D116254093DE41F" class="Gastropoda" family="Camaenidae" genus="Semotrachia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Semotrachia" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="genus">
<emphasis id="E980593EA7976DDD5EAF1228C3F99A51" italics="true" pageId="0" pageNumber="427">Semotrachia</emphasis>
</taxonomicName>
,
<taxonomicName id="1947415BDE3CF7CB776C667EFB1280B3" authorityName="Pilsbry" authorityYear="1891" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" lsidName="Austrochloritis" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="genus">
<emphasis id="37CC734B1B9F556B458630DC4C5FEDBC" italics="true" pageId="0" pageNumber="427">Austrochloritis</emphasis>
</taxonomicName>
and &quot;
<taxonomicName id="DF4FBAA8E12B8B5096B03187BF268CFD" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis argilacea" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="argilacea">
<emphasis id="4C36371DE64933DFE3BE6BA05AF5E571" italics="true" pageId="0" pageNumber="427">Chloritis argilacea</emphasis>
</taxonomicName>
&quot; (presently
<taxonomicName id="63C106873370BDC9E166F2DD4499DC28" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Camaenidae" genus="Parachloritis" kingdom="Animalia" lsidName="Parachloritis argilacea" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="argilacea">
<emphasis id="68B1B3BC05042039DF95D3D017B030C8" italics="true" pageId="0" pageNumber="427">Parachloritis argilacea</emphasis>
</taxonomicName>
; see
<bibRefCitation id="A354E602585EF88BD3A1E2371B68CAA2" DOI="https://doi.org/10.1163/18759866-08301001" author="Koehler, F" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="427" pagination="1 - 40" refId="B13" refString="Koehler, F, Kessner, V, 2014. Mitochondrial and morphological differentiation in a previously unrecognized radiation of land snails on Timor (Gastropoda, Camaenidae, Parachloritis Ehrmann, 1912). Contributions to Zoology 83 (1): 1 - 40, DOI: https://doi.org/10.1163/18759866-08301001" title="Mitochondrial and morphological differentiation in a previously unrecognized radiation of land snails on Timor (Gastropoda, Camaenidae, Parachloritis Ehrmann, 1912)." url="https://doi.org/10.1163/18759866-08301001" volume="83" year="2014">
<normalizedToken id="1FE089D001EE9B265FBBDE4BF9D722AF" originalValue="Köhler">Koehler</normalizedToken>
and Kessner 2014
</bibRefCitation>
), but that the genital anatomy of
<taxonomicName id="C6731FF56D3438550CB31A93BD3977BD" authorityName="Iredale" authorityYear="1938" class="Gastropoda" family="Camaenidae" genus="Parglogenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parglogenia" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="genus">
<emphasis id="14EEB89D21FA9516EFA49A469F59BA3B" italics="true" pageId="0" pageNumber="427">Parglogenia</emphasis>
</taxonomicName>
was highly distinctive.
<bibRefCitation id="DFE9C248AD3D9B17D30598F6DA460B97" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="FB8A7B5F885CD87F5D55FB423D55BBD1" originalValue="Köhler">Koehler</normalizedToken>
(2012)
</bibRefCitation>
illustrated the reproductive anatomy of a specimen of
<taxonomicName id="7546DD35F99ADD996D3F6076E9F4CDCF" lsidName="P. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="59F8B389546B4F2B2FDB449B253081E4" italics="true" pageId="0" pageNumber="427">P. pelodes</emphasis>
</taxonomicName>
from Melville Island, which closely resembled that of specimens from Darwin. Shells of
<taxonomicName id="EB38C88062A8B163699E692C8F6924B2" lsidName="P. pelodes" pageId="0" pageNumber="427" rank="species" species="pelodes">
<emphasis id="1AB2DEC0711AFEAB21649EB4ADD4FE18" italics="true" pageId="0" pageNumber="427">P. pelodes</emphasis>
</taxonomicName>
, including the holotype of
<taxonomicName id="89A460B45AF266D0C6C2C3C6E75DCC9E" authorityName="Pilsbry" authorityYear="1893" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis pseudoprunum" order="Stylommatophora" pageId="0" pageNumber="427" phylum="Mollusca" rank="species" species="pseudoprunum">
<emphasis id="39EBE7777CB4CEE3E1F48AEF0AABB3AF" italics="true" pageId="0" pageNumber="427">Chloritis pseudoprunum</emphasis>
</taxonomicName>
, have been figured by
<bibRefCitation id="A0C0BE8CB2E2CCC25C0A53BB8789FFFC" DOI="https://doi.org/10.1002/zoos.201200006" author="Koehler, F" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="427" pagination="53 - 62" refId="B10" refString="Koehler, F, 2012. Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae). Zoosystematics and Evolution 88 (1): 53 - 62, DOI: https://doi.org/10.1002/zoos.201200006" title="Taxonomic revision of two endemic land snail genera from the Top End of Northern Australia with remarks on two problematic species named by de Ferussac and Le Guillou (Eupulmonata, Camaenidae)." url="https://doi.org/10.1002/zoos.201200006" volume="88" year="2012">
<normalizedToken id="7E46653A9C338D11323C35FFB8C2A933" originalValue="Köhler">Koehler</normalizedToken>
(2012)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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