277 lines
35 KiB
XML
277 lines
35 KiB
XML
<document id="89D672667C80573B2A9FAE87BA29A909" ID-CLB-Dataset="54605" ID-DOI="10.37828/em.2017.10.5" ID-GBIF-Dataset="1f58e33f-ed07-4f7c-8c2a-61e7caafe9cf" ID-ISSN="2336-9744" ID-Zenodo-Dep="8027941" ID-ZooBank="urn:lsid:zoobank.org:pub:476BC71C-E055-47A4-8280-205F37A6A981" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1686314400675" checkinUser="felipe" docAuthor="Grego, Jozef & Szekeres, Miklós" docDate="2017" docId="03DD87F50708FFFFFF4BFC59ED39FCF9" docLanguage="en" docName="EcolMontenegrina.10.26-30.pdf" docOrigin="Ecologica Montenegrina 10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.1:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="1" docTitle="Cotyorica nemethi Grego & Szekeres 2017, n. sp." docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FFE4FF8D0709FFFCFFDCFFDFEF45FFC4" masterDocTitle="Cotyorica nemethi n. gen. n. sp., a remarkable Tertiary relict of the subfamily Phaedusinae (Gastropoda: Pulmonata: Clausiliidae) from northern Turkey" masterLastPageNumber="30" masterPageNumber="26" pageNumber="27" updateTime="1686575673261" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="5406F8213B6BB4672FF237C06CD033EC">
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<mods:title id="6F3D8DDB1A2E949558B0257CCCF8547D">Cotyorica nemethi n. gen. n. sp., a remarkable Tertiary relict of the subfamily Phaedusinae (Gastropoda: Pulmonata: Clausiliidae) from northern Turkey</mods:title>
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<mods:name id="09921628B7246DCC0BC1A6B8210BAA77" type="personal">
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<mods:role id="6628A8CAF674CC6C1FBD25F7051D19C9">
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<mods:roleTerm id="B77195D05C7298560A9D38A06006F608">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F8582D2E6AB1A3876F236B375990E9D">Grego, Jozef</mods:namePart>
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<mods:affiliation id="06B48919B34F149F6396991B5FE3A4A5">Horná Mičiná 219, SK- 97401 Banská Bystrica, Slovakia, e-mail: jozef. grego @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="23DF6DF3E9E8E709C08060736E762A6D" type="personal">
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<mods:role id="3267498A12717E665FB57543090D0D94">
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<mods:roleTerm id="99C953F9CA3B0BD53D8349FE58F8163B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7B381B78903D178B977C06522B2716CF">Szekeres, Miklós</mods:namePart>
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<mods:affiliation id="D1C419DE34B18B1D05E02A8832C3C155">Horná Mičiná 219, SK- 97401 Banská Bystrica, Slovakia, e-mail: jozef. grego @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="11E37AC8521FBBACCF68E90A12A57847">text</mods:typeOfResource>
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<mods:title id="CAEE363BCC7EEB89A3102D7E3C79E9C4">Ecologica Montenegrina</mods:title>
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</mods:titleInfo>
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<mods:part id="61D0BD963B9C2DA539C51568C1A269F0">
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<mods:date id="DDB1D288C17559AD07C52911B7C70A53">2017</mods:date>
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<mods:number id="5D5A29D6B913762F18F69A3040687D62">2017-03-17</mods:number>
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<mods:number id="44039563CBB1934424BE31911BEA9723">10</mods:number>
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<mods:start id="D4BC8000B86D42635C1D3D174202C057">26</mods:start>
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<mods:end id="FAC455E94EC16D0A66605ECDD7865700">30</mods:end>
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<mods:classification id="23518E2542677084887A7DACB4B68A54">journal article</mods:classification>
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<mods:identifier id="56512CF79BE5446A5BA25F4E60981262" type="DOI">10.37828/em.2017.10.5</mods:identifier>
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<treatment id="03DD87F50708FFFFFF4BFC59ED39FCF9" ID-DOI="http://doi.org/10.5281/zenodo.8027951" ID-Zenodo-Dep="8027951" LSID="urn:lsid:plazi:treatment:03DD87F50708FFFFFF4BFC59ED39FCF9" httpUri="http://treatment.plazi.org/id/03DD87F50708FFFFFF4BFC59ED39FCF9" lastPageId="3" lastPageNumber="29" pageId="1" pageNumber="27">
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<subSubSection id="C36E65680708FFFDFF4BFC59EE85FC64" box="[151,448,902,928]" pageId="1" pageNumber="27" type="nomenclature">
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<paragraph id="8BCB36E30708FFFDFF4BFC59EE85FC64" blockId="1.[151,448,902,961]" box="[151,448,902,928]" pageId="1" pageNumber="27">
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<heading id="D083818F0708FFFDFF4BFC59EE85FC64" box="[151,448,902,928]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="27" reason="2">
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<emphasis id="B900EAF10708FFFDFF4BFC59EE85FC64" bold="true" box="[151,448,902,928]" pageId="1" pageNumber="27">
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<taxonomicName id="4C744D600708FFFDFF4BFC59EE30FC64" ID-CoL="YZD6" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[151,373,902,928]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="27" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
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<emphasis id="B900EAF10708FFFDFF4BFC59EE30FC64" bold="true" box="[151,373,902,928]" italics="true" pageId="1" pageNumber="27">Cotyorica nemethi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A233578A0708FFFDFEA1FC59EE85FC64" box="[381,448,902,928]" pageId="1" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E65680708FFFDFF4BFC79EE54FC05" box="[151,273,934,961]" pageId="1" pageNumber="27" type="description">
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<paragraph id="8BCB36E30708FFFDFF4BFC79EE54FC05" blockId="1.[151,448,902,961]" box="[151,273,934,961]" pageId="1" pageNumber="27">
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(
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<figureCitation id="134F2A660708FFFDFF43FC79EFAFFC05" box="[159,234,934,961]" captionStart="Figure 1" captionStartId="1.[151,226,2020,2044]" captionTargetBox="[176,1411,1109,2002]" captionTargetId="figure-195@1.[176,1411,1109,2002]" captionTargetPageId="1" captionText="Figure 1. Cotyorica nemethi n. sp. Holotype, 11.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8027943" httpUri="https://zenodo.org/record/8027943/files/figure.png" pageId="1" pageNumber="27">Figs 1</figureCitation>
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,
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<figureCitation id="134F2A660708FFFDFF24FC79EE43FC05" box="[248,262,934,961]" captionStart="Figure 2" captionStartId="2.[151,227,1562,1586]" captionTargetBox="[408,1140,825,1503]" captionTargetId="figure-260@2.[344,1243,763,1556]" captionTargetPageId="2" captionText="Figure 2. Lamella positions in the aperture of Cotyorica nemethi n. sp. The lamellae superior (Lsu), inferior (Lin) and subcolumellaris (Lsc) are marked." figureDoi="http://doi.org/10.5281/zenodo.8027945" httpUri="https://zenodo.org/record/8027945/files/figure.png" pageId="1" pageNumber="27">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E65680708FFFDFF4BFC35EECBFB8C" pageId="1" pageNumber="27" type="materials_examined">
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<paragraph id="8BCB36E30708FFFDFF4BFC35EECBFB8C" blockId="1.[151,1436,1002,1096]" pageId="1" pageNumber="27">
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<emphasis id="B900EAF10708FFFDFF4BFC35EE0DFBC0" bold="true" box="[151,328,1002,1028]" pageId="1" pageNumber="27">Type material</emphasis>
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:
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<materialsCitation id="3B1C3CBE0708FFFDFE84FC35EECBFB8C" collectingDate="1991-06-30" collectionCode="HNHM" collectorName="L. Nemeth" country="Turkey" elevation="20" latitude="41.116665" location="Persembe" longLatPrecision="1158" longitude="37.783333" pageId="1" pageNumber="27" specimenCode="HNHM 100140" specimenCount="1" stateProvince="Ordu" typeStatus="holotype">
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<collectingCountry id="F36376730708FFFDFE84FC35EEF5FBC1" box="[344,432,1002,1029]" name="Turkey" pageId="1" pageNumber="27">Turkey</collectingCountry>
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,
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<collectingRegion id="49B0F8010708FFFDFE60FC35ED29FBC1" box="[444,620,1002,1029]" country="Turkey" name="Ordu" pageId="1" pageNumber="27">Ordu Province</collectingRegion>
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, coastal slope between
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<location id="8EAB60380708FFFDFC5CFC35ECB7FBC1" LSID="urn:lsid:plazi:treatment:03DD87F50708FFFFFF4BFC59ED39FCF9:8EAB60380708FFFDFC5CFC35ECB7FBC1" box="[896,1010,1002,1029]" country="Turkey" latitude="41.116665" longLatPrecision="1158" longitude="37.783333" name="Persembe" pageId="1" pageNumber="27" stateProvince="Ordu">Perşembe</location>
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and Mersin (
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<geoCoordinate id="EE4050240708FFFDFB4CFC35EBBCFBC1" box="[1168,1273,1002,1029]" degrees="41" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="27" precision="925" value="41.116665">41°07' N</geoCoordinate>
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<geoCoordinate id="EE4050240708FFFDFADCFC35EA2CFBC1" box="[1280,1385,1002,1029]" degrees="37" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="27" precision="925" value="37.783333">37°47' E</geoCoordinate>
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),
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<quantity id="4C8C9B060708FFFDFAA3FC35EFF5FBE2" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="27" unit="m" value="20.0">
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<elevation id="0059D1D00708FFFDFAA3FC35EFF5FBE2" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="27" unit="m" value="20.0">20 m</elevation>
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</quantity>
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, leg.
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<collectorName id="268153350708FFFDFF2EFBD3EE34FBE2" box="[242,369,1035,1062]" pageId="1" pageNumber="27">L. Németh</collectorName>
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<date id="FFCA10230708FFFDFEA6FBD4ED47FBE2" box="[378,514,1035,1062]" pageId="1" pageNumber="27" value="1991-06-30">
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<collectingDate id="EF8EE9CB0708FFFDFEA6FBD4ED47FBE2" box="[378,514,1035,1062]" pageId="1" pageNumber="27" value="1991-06-30">30.06.1991</collectingDate>
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</date>
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.
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<typeStatus id="54CF88410708FFFDFDD2FBD4ED3BFBE2" box="[526,638,1035,1062]" pageId="1" pageNumber="27">Holotype</typeStatus>
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,
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<specimenCode id="DBD29E980708FFFDFD50FBD3EC03FBE2" box="[652,838,1035,1062]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="27">HNHM 100140</specimenCode>
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, to be deposited at the Hungarian Natural History Museum (Budapest).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<caption id="DF0B666B0708FFFDFF4BF83BEDA3F83F" ID-DOI="http://doi.org/10.5281/zenodo.8027943" ID-Zenodo-Dep="8027943" box="[151,742,2020,2044]" httpUri="https://zenodo.org/record/8027943/files/figure.png" pageId="1" pageNumber="27" startId="1.[151,226,2020,2044]" targetBox="[176,1411,1109,2002]" targetPageId="1" targetType="figure">
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<paragraph id="8BCB36E30708FFFDFF4BF83BEDA3F83F" blockId="1.[151,742,2020,2044]" box="[151,742,2020,2044]" pageId="1" pageNumber="27">
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<emphasis id="B900EAF10708FFFDFF4BF83BEFB3F838" bold="true" box="[151,246,2020,2044]" pageId="1" pageNumber="27">Figure 1</emphasis>
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.
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<taxonomicName id="4C744D600708FFFDFED8F83BEE8FF838" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[260,458,2020,2044]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="27" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
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<emphasis id="B900EAF10708FFFDFED8F83BEE8FF838" box="[260,458,2020,2044]" italics="true" pageId="1" pageNumber="27">Cotyorica nemethi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A233578A0708FFFDFE0DF83BED4FF83F" box="[465,522,2020,2043]" pageId="1" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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Holotype, 11.5 mm.
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</paragraph>
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</caption>
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<subSubSection id="C36E6568070BFFFEFF4BFF1BEBF0FEE6" pageId="2" pageNumber="28" type="diagnosis">
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<paragraph id="8BCB36E3070BFFFEFF4BFF1BEBF0FEE6" blockId="2.[151,1436,195,290]" pageId="2" pageNumber="28">
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<emphasis id="B900EAF1070BFFFEFF4BFF1BEE55FF1A" bold="true" box="[151,272,196,222]" pageId="2" pageNumber="28">Diagnosis</emphasis>
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: Non-decollated small species of the
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<taxonomicName id="4C744D60070BFFFEFD01FF1CEC35FF1A" authorityName="A. J. Wagner" authorityYear="1922" box="[733,880,195,222]" class="Gastropoda" family="Clausiliidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="30" phylum="Mollusca" rank="subFamily" subFamily="Phaedusinae">Phaedusinae</taxonomicName>
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with strongly costate neck, all around serrate peristome, high positioned, deep divided lamella inferior ending in two parallel branches, uninterrupted lamella subcolumellaris, as well as short upper and longer lower plicae on the lateral side.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6568070BFFFEFF4BFE94EEEFFD3E" pageId="2" pageNumber="28" type="description">
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<paragraph id="8BCB36E3070BFFFEFF4BFE94EDA9FD73" blockId="2.[151,1436,324,762]" pageId="2" pageNumber="28">
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<emphasis id="B900EAF1070BFFFEFF4BFE94EE6DFEA1" bold="true" box="[151,296,331,357]" pageId="2" pageNumber="28">Description</emphasis>
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: The colourless shell with slightly bent spire consists of 9
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<superScript id="7C019BAB070BFFFEFC34FE9BECB4FE91" attach="none" box="[1000,1009,324,341]" fontSize="7" pageId="2" pageNumber="28">1</superScript>
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/
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<subScript id="17F034A6070BFFFEFC25FE89EB47FEA3" attach="left" box="[1017,1026,342,359]" fontSize="7" pageId="2" pageNumber="28">3</subScript>
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flat whorls. The dense ribs of the surface become stronger, sharper and widely spaced at the neck. The ovoid peristome is projected, at the basis not backward-bent. Its wide, reflexed margin has strong plicae all around, except at the sinulus. The lamella superior is moderately emerged, in front view not more conspicuous than the interlamellar plicae. Inward it is smoothly continuous with the lamella spiralis. Deeper than visible through the aperture the lamella inferior divides in two parallel branches, which descend along the lamella spiralis and terminate marginally as strong plicae above half height of the aperture. The uninterrupted lamella subcolumellaris also ends marginally, separated from the lower branch of the inferior by a strong plica. The plica principalis starts ventrolaterally and, gradually converging toward the suture, reaches close to the aperture. On the lateral side it is accompanied by a short upper and a farther forward reaching, twice longer lower palatal plica. The deep clausilium plate is not visible through the aperture.
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</paragraph>
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<paragraph id="8BCB36E3070BFFFEFF2AFD62EEEFFD3E" blockId="2.[151,1436,324,762]" pageId="2" pageNumber="28">
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Dimensions of the
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<typeStatus id="54CF8841070BFFFEFE3DFD62ED09FD1C" box="[481,588,701,728]" pageId="2" pageNumber="28">holotype</typeStatus>
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: shell height 11.5 mm, shell width 2.9 mm, aperture height 2.2 mm, aperture width 1.7 mm.
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</paragraph>
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</subSubSection>
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<caption id="DF0B666B070BFFFEFF4BF9C5ED41F994" ID-DOI="http://doi.org/10.5281/zenodo.8027945" ID-Zenodo-Dep="8027945" httpUri="https://zenodo.org/record/8027945/files/figure.png" pageId="2" pageNumber="28" startId="2.[151,227,1562,1586]" targetBox="[408,1140,825,1503]" targetPageId="2" targetType="figure">
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<paragraph id="8BCB36E3070BFFFEFF4BF9C5ED41F994" blockId="2.[151,1436,1561,1616]" pageId="2" pageNumber="28">
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<emphasis id="B900EAF1070BFFFEFF4BF9C5EFBDF9F6" bold="true" box="[151,248,1562,1586]" pageId="2" pageNumber="28">Figure 2</emphasis>
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. Lamella positions in the aperture of
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<taxonomicName id="4C744D60070BFFFEFD4DF9C5EC1DF9F5" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[657,856,1561,1585]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="28" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
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<emphasis id="B900EAF1070BFFFEFD4DF9C5EC1DF9F5" box="[657,856,1561,1585]" italics="true" pageId="2" pageNumber="28">Cotyorica nemethi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A233578A070BFFFEFCBDF9C5ECDEF9F5" box="[865,923,1562,1585]" pageId="2" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
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The lamellae superior (Lsu), inferior (Lin) and subcolumellaris (Lsc) are marked.
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</paragraph>
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</caption>
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<subSubSection id="C36E6568070BFFFEFF4BF944EB0EF807" pageId="2" pageNumber="28" type="etymology">
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<paragraph id="8BCB36E3070BFFFEFF4BF944EA2CF913" blockId="2.[151,1436,1691,1751]" pageId="2" pageNumber="28">
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<emphasis id="B900EAF1070BFFFEFF4BF944EE5BF971" bold="true" box="[151,286,1691,1717]" pageId="2" pageNumber="28">Etymology</emphasis>
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: The new species is named after László Németh, the collector of the
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<typeStatus id="54CF8841070BFFFEFB52F943EB85F972" box="[1166,1216,1692,1718]" pageId="2" pageNumber="28">type</typeStatus>
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specimen and an esteemed friend of the authors, who substantially contributed to the research of the Turkish malacofauna.
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</paragraph>
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<paragraph id="8BCB36E3070BFFFEFF4BF8DFEB0EF807" blockId="2.[151,1436,1792,1987]" pageId="2" pageNumber="28">
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<emphasis id="B900EAF1070BFFFEFF4BF8DFEFB2F8DE" bold="true" box="[151,247,1792,1818]" pageId="2" pageNumber="28">Habitat</emphasis>
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:
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<taxonomicName id="4C744D60070BFFFEFED6F8DEEEA3F8DF" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[266,486,1792,1819]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="28" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
|
|
<emphasis id="B900EAF1070BFFFEFED6F8DEEEA3F8DF" box="[266,486,1792,1819]" italics="true" pageId="2" pageNumber="28">Cotyorica nemethi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A233578A070BFFFEFE2CF8DEED77F8DF" box="[496,562,1793,1819]" pageId="2" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
|
was recovered as an empty shell from a litter-soil sample collected at a cliff. As its closest relatives
|
|
<taxonomicName id="4C744D60070BFFFEFE30F8FDECF4F8F9" authority="Rahle, 1982" authorityName="Rahle" authorityYear="1982" box="[492,945,1826,1853]" class="Gastropoda" family="Clausiliidae" genus="Graecophaedusa" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="28" phylum="Mollusca" rank="species" species="sperrlei">
|
|
<emphasis id="B900EAF1070BFFFEFE30F8FDEC5DF8F9" box="[492,792,1826,1853]" italics="true" pageId="2" pageNumber="28">Graecophaedusa sperrlei</emphasis>
|
|
<bibRefCitation id="EFE54B12070BFFFEFCFDF8FDECF4F8F9" author="Rahle, W." box="[801,945,1826,1853]" pageId="2" pageNumber="28" pagination="45 - 48" refId="ref1892" refString="Rahle, W. (1982) Graecophaedusa sperrlei n. gen., n. sp.: ein bemerkenswertes Tertiarrelikt aus Griechisch- Mazedonien. Archiv fur Molluskenkunde, 112, 45 - 48." type="journal article" year="1982">Rähle, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C744D60070BFFFEFC31F8FDEE2BF89A" authority="Nemeth & Szekeres, 1995" authorityName="Nemeth & Szekeres" authorityYear="1995" class="Gastropoda" family="Clausiliidae" genus="Nothoserrulina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="28" phylum="Mollusca" rank="species" species="subterranea">
|
|
<emphasis id="B900EAF1070BFFFEFC31F8FDEA72F8F9" box="[1005,1335,1826,1853]" italics="true" pageId="2" pageNumber="28">Nothoserrulina subterranea</emphasis>
|
|
<bibRefCitation id="EFE54B12070BFFFEFA9CF8FDEE2BF89A" author="Nemeth, L. & Szekeres, M." pageId="2" pageNumber="28" pagination="93 - 96" refId="ref1700" refString="Nemeth, L. & Szekeres, M. (1995) New and little known species of the Serrulina group from northern Turkey (Gastropoda: Pulmonata: Clausiliidae). Archiv fur Molluskenkunde, 124, 93 - 96." type="journal article" year="1995">Németh & Szekeres, 1995</bibRefCitation>
|
|
</taxonomicName>
|
|
were found in narrow underground passages between rocks (
|
|
<bibRefCitation id="EFE54B12070BFFFEFB87F89CEAD2F89A" author="Nemeth, L. & Szekeres, M." box="[1115,1431,1859,1886]" pageId="2" pageNumber="28" pagination="93 - 96" refId="ref1700" refString="Nemeth, L. & Szekeres, M. (1995) New and little known species of the Serrulina group from northern Turkey (Gastropoda: Pulmonata: Clausiliidae). Archiv fur Molluskenkunde, 124, 93 - 96." type="journal article" year="1995">Németh & Szekeres 1995</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFE54B12070BFFFEFF4BF8BAEEE1F844" author="Reischutz, A. & Reischutz, P. L. & Fischer, W." box="[151,420,1893,1920]" pageId="2" pageNumber="28" pagination="11 - 13" refId="ref1928" refString="Reischutz, A., Reischutz, P. L. & Fischer, W. (2008) Hellenika pantoia, 15: Kein Ende abzusehen - Zur Kenntnis der Gattung Balcanodiscus Riedel & Urbanski 1964 (Gastropoda: Pulmonata: Zonitidae). Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft, 15, 11 - 13." type="journal article" year="2008">
|
|
Reischütz
|
|
<emphasis id="B900EAF1070BFFFEFEC4F8B9EE14F844" box="[280,337,1893,1920]" italics="true" pageId="2" pageNumber="28">et al</emphasis>
|
|
. 2008
|
|
</bibRefCitation>
|
|
), it is assumed that this species may have similar habitat preference. This is also suggested by the highly local occurrence of the new species in a zoogeographically well researched region. The
|
|
<typeStatus id="54CF8841070BFFFEFF10F877EE76F807" box="[204,307,1960,1987]" pageId="2" pageNumber="28">holotype</typeStatus>
|
|
may have been washed out from crevices of the cliff by rain water.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36E6568070AFFFFFF4BFF1BED39FCF9" pageId="3" pageNumber="29" type="discussion">
|
|
<paragraph id="8BCB36E3070AFFFFFF4BFF1BEE4CFF1A" blockId="3.[151,1437,196,829]" box="[151,265,196,222]" pageId="3" pageNumber="29">
|
|
<emphasis id="B900EAF1070AFFFFFF4BFF1BEE4CFF1A" bold="true" box="[151,265,196,222]" pageId="3" pageNumber="29">Remarks</emphasis>
|
|
</paragraph>
|
|
<paragraph id="8BCB36E3070AFFFFFF4BFF3AEDB0FDB7" blockId="3.[151,1437,196,829]" pageId="3" pageNumber="29">
|
|
Among non-decollated western
|
|
<taxonomicName id="4C744D60070AFFFFFDFEFF3AEDF0FEC4" authorityName="A. J. Wagner" authorityYear="1922" box="[546,693,229,256]" class="Gastropoda" family="Clausiliidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="30" phylum="Mollusca" rank="subFamily" subFamily="Phaedusinae">Phaedusinae</taxonomicName>
|
|
<taxonomicName id="4C744D60070AFFFFFD1FFF39EC0CFEC4" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[707,841,229,256]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
|
|
<emphasis id="B900EAF1070AFFFFFD1FFF39EC0CFEC4" box="[707,841,229,256]" italics="true" pageId="3" pageNumber="29">C. nemethi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A233578A070AFFFFFC8AFF39ECD9FEC4" box="[854,924,230,256]" pageId="3" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
|
|
shows superficial similarity to
|
|
<taxonomicName id="4C744D60070AFFFFFAF2FF3AEE81FEE6" authority="(Pfeiffer, 1847)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1847" class="Gastropoda" family="Clausiliidae" genus="Serrulina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="species" species="serrulata">
|
|
<emphasis id="B900EAF1070AFFFFFAF2FF3AEE46FEE6" italics="true" pageId="3" pageNumber="29">Serrulina serrulata</emphasis>
|
|
(Pfeiffer, 1847)
|
|
</taxonomicName>
|
|
that differs markedly by its steeply descending and low ending lamella inferior, a feature shared between members of the genera
|
|
<taxonomicName id="4C744D60070AFFFFFCDFFEF7EB06FE87" authority="Mousson, 1873" authorityName="Mousson" authorityYear="1873" box="[771,1091,296,323]" class="Gastropoda" family="Clausiliidae" genus="Serrulina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFCDFFEF7EC37FE87" box="[771,882,296,323]" italics="true" pageId="3" pageNumber="29">Serrulina</emphasis>
|
|
Mousson, 1873
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C744D60070AFFFFFB85FEF7EADDFE87" authority="Nordsieck, 1973" authorityName="Nordsieck" authorityYear="1973" box="[1113,1432,296,323]" class="Gastropoda" family="Clausiliidae" genus="Dobatia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFB85FEF7EBFCFE87" box="[1113,1209,296,323]" italics="true" pageId="3" pageNumber="29">Dobatia</emphasis>
|
|
Nordsieck, 1973
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C744D60070AFFFFFF4BFE95EEA7FEA1" authority="Boettger, 1935" authorityName="Boettger" authorityYear="1935" box="[151,482,330,357]" class="Gastropoda" family="Clausiliidae" genus="Sciocochlea" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFF4BFE95EE60FEA1" box="[151,293,330,357]" italics="true" pageId="3" pageNumber="29">Sciocochlea</emphasis>
|
|
Boettger, 1935
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="4C744D60070AFFFFFDFAFE95ECE7FEA1" authority="Gittenberger, 2000" authorityName="Gittenberger" authorityYear="2000" box="[550,930,330,357]" class="Gastropoda" family="Clausiliidae" genus="Tsoukatosia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFDFAFE95EDF1FEA1" box="[550,692,330,357]" italics="true" pageId="3" pageNumber="29">Tsoukatosia</emphasis>
|
|
<bibRefCitation id="EFE54B12070AFFFFFD61FE95ECE7FEA1" author="Gittenberger, E." box="[701,930,330,357]" pageId="3" pageNumber="29" pagination="81 - 87" refId="ref1616" refString="Gittenberger, E. (2000) Serrulininae in Greece, there may be more (Gastropoda, Pulmonata, Clausiliidae). Basteria, 64, 81 - 87." type="journal article" year="2000">Gittenberger, 2000</bibRefCitation>
|
|
</taxonomicName>
|
|
. By contrast, the high positioned lamellae inferior and subcolumellaris of the new species (
|
|
<figureCitation id="134F2A66070AFFFFFCDCFEB3EC0BFE43" box="[768,846,364,391]" captionStart="Figure 2" captionStartId="2.[151,227,1562,1586]" captionTargetBox="[408,1140,825,1503]" captionTargetId="figure-260@2.[344,1243,763,1556]" captionTargetPageId="2" captionText="Figure 2. Lamella positions in the aperture of Cotyorica nemethi n. sp. The lamellae superior (Lsu), inferior (Lin) and subcolumellaris (Lsc) are marked." figureDoi="http://doi.org/10.5281/zenodo.8027945" httpUri="https://zenodo.org/record/8027945/files/figure.png" pageId="3" pageNumber="29">Fig. 2</figureCitation>
|
|
) resemble those seen in the monotypic genera
|
|
<taxonomicName id="4C744D60070AFFFFFF4BFE51EEB3FE6D" authority="Rahle, 1982" authorityName="Rahle" authorityYear="1982" box="[151,502,398,425]" class="Gastropoda" family="Clausiliidae" genus="Graecophaedusa" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFF4BFE51EE25FE6D" box="[151,352,398,425]" italics="true" pageId="3" pageNumber="29">Graecophaedusa</emphasis>
|
|
<bibRefCitation id="EFE54B12070AFFFFFEBBFE51EEB3FE6D" author="Rahle, W." box="[359,502,398,425]" pageId="3" pageNumber="29" pagination="45 - 48" refId="ref1892" refString="Rahle, W. (1982) Graecophaedusa sperrlei n. gen., n. sp.: ein bemerkenswertes Tertiarrelikt aus Griechisch- Mazedonien. Archiv fur Molluskenkunde, 112, 45 - 48." type="journal article" year="1982">Rähle, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C744D60070AFFFFFDF3FE51EB66FE6D" authority="Nemeth & Szekeres, 1995" authorityName="Nemeth & Szekeres" authorityYear="1995" box="[559,1059,398,425]" class="Gastropoda" family="Clausiliidae" genus="Nothoserrulina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFDF3FE51EDA7FE6D" box="[559,738,398,425]" italics="true" pageId="3" pageNumber="29">Nothoserrulina</emphasis>
|
|
<bibRefCitation id="EFE54B12070AFFFFFD37FE51EB66FE6D" author="Nemeth, L. & Szekeres, M." box="[747,1059,398,425]" pageId="3" pageNumber="29" pagination="93 - 96" refId="ref1700" refString="Nemeth, L. & Szekeres, M. (1995) New and little known species of the Serrulina group from northern Turkey (Gastropoda: Pulmonata: Clausiliidae). Archiv fur Molluskenkunde, 124, 93 - 96." type="journal article" year="1995">Németh & Szekeres, 1995</bibRefCitation>
|
|
</taxonomicName>
|
|
. Nevertheless,
|
|
<taxonomicName id="4C744D60070AFFFFFB0AFE51EA10FE6D" authority="Grego & Szekeres, 2017" authorityName="Grego & Szekeres" authorityYear="2017" box="[1238,1365,398,425]" class="Gastropoda" family="Clausiliidae" genus="Cotyorica" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="species" species="nemethi" status="n. sp.">
|
|
<emphasis id="B900EAF1070AFFFFFB0AFE51EA10FE6D" box="[1238,1365,398,425]" italics="true" pageId="3" pageNumber="29">C. nemethi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A233578A070AFFFFFA81FE50EAD9FE6D" box="[1373,1436,399,425]" pageId="3" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
|
|
is well distinguishable from
|
|
<taxonomicName id="4C744D60070AFFFFFE2CFE70EC5BFE0E" authorityName="Rahle" authorityYear="1982" box="[496,798,431,458]" class="Gastropoda" family="Clausiliidae" genus="Graecophaedusa" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="species" species="sperrlei">
|
|
<emphasis id="B900EAF1070AFFFFFE2CFE70EC5BFE0E" box="[496,798,431,458]" italics="true" pageId="3" pageNumber="29">Graecophaedusa sperrlei</emphasis>
|
|
</taxonomicName>
|
|
by the narrow apex and less retracted palatal plicae, from
|
|
<taxonomicName id="4C744D60070AFFFFFF07FE0EED62FE28" authorityName="Nemeth & Szekeres" authorityYear="1995" box="[219,551,465,492]" class="Gastropoda" family="Clausiliidae" genus="Nothoserrulina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="species" species="subterranea">
|
|
<emphasis id="B900EAF1070AFFFFFF07FE0EED62FE28" box="[219,551,465,492]" italics="true" pageId="3" pageNumber="29">Nothoserrulina subterranea</emphasis>
|
|
</taxonomicName>
|
|
by the spindle-shaped shell and the absence of minor folds between the upper and lower plicae, and from both species by the projected aperture with non-recessed basal rim, and the strongly serrate parietal peristome margin. Considering these differences relative to the morphological coherence of species in the non-monotypic genera (
|
|
<emphasis id="B900EAF1070AFFFFFD24FDE8EC59FD94" box="[760,796,567,592]" italics="true" pageId="3" pageNumber="29">e.g</emphasis>
|
|
.,
|
|
<taxonomicName id="4C744D60070AFFFFFCEEFDE9ECE4FD95" authorityName="Mousson" authorityYear="1873" box="[818,929,566,593]" class="Gastropoda" family="Clausiliidae" genus="Serrulina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFCEEFDE9ECE4FD95" box="[818,929,566,593]" italics="true" pageId="3" pageNumber="29">Serrulina</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C744D60070AFFFFFC6CFDE9EB97FD95" authority=", Tsoukatosa" authorityName="Tsoukatosa" box="[944,1234,566,593]" class="Gastropoda" family="Clausiliidae" genus="Sciocochlea" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B900EAF1070AFFFFFC6CFDE9EB7BFD95" box="[944,1086,566,593]" italics="true" pageId="3" pageNumber="29">Sciocochlea</emphasis>
|
|
,
|
|
<emphasis id="B900EAF1070AFFFFFB91FDE9EB97FD95" box="[1101,1234,566,593]" italics="true" pageId="3" pageNumber="29">Tsoukatosa</emphasis>
|
|
</taxonomicName>
|
|
), they justify the classification of the new species in a distinct genus.
|
|
</paragraph>
|
|
<paragraph id="8BCB36E3070AFFFFFF2AFDA5ED39FCF9" blockId="3.[151,1437,196,829]" pageId="3" pageNumber="29">
|
|
Ample all year round rainfall and the narrow, within 15°C range of monthly average temperatures of the southern and eastern Black Sea coast provide ideal climatic conditions for the survival of
|
|
<taxonomicName id="4C744D60070AFFFFFAD5FD43EADEFD73" authorityName="A. J. Wagner" authorityYear="1922" box="[1289,1435,668,695]" class="Gastropoda" family="Clausiliidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="30" phylum="Mollusca" rank="subFamily" subFamily="Phaedusinae">Phaedusinae</taxonomicName>
|
|
species, which were abundant elements of the European
|
|
<taxonomicName id="4C744D60070AFFFFFC88FD62ECA5FD1C" authorityName="J. E. Gray" authorityYear="1855" box="[852,992,701,728]" class="Gastropoda" family="Clausiliidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="29" phylum="Mollusca" rank="family">Clausiliidae</taxonomicName>
|
|
fauna before the Pleistocene period (
|
|
<bibRefCitation id="EFE54B12070AFFFFFF43FD00EE25FD3E" author="Nordsieck, H." box="[159,352,735,762]" pageId="3" pageNumber="29" pagination="1 - 15" refId="ref1853" refString="Nordsieck, H. (2000) Annotated check-list of the fossil (pre-Pleistocene) Clausiliidae (Gastropoda: Stylommatophora) from central and western Europe. Mitteilungen der Deutschen Malakozoologischen Gesellschaft, 65, 1 - 15." type="journal article" year="2000">Nordsieck 2000</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFE54B12070AFFFFFEB1FD00ED29FD3E" author="Reischutz, A. & Reischutz, P. L. & Szekeres, M." box="[365,620,735,762]" pageId="3" pageNumber="29" pagination="93 - 117" refId="ref1986" refString="Reischutz, A., Reischutz, P. L. & Szekeres, M. (2016) The Clausiliidae subfamily Phaedusinae (Gastropoda, Pulmonata) in the Balkans. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft, 23, 93 - 117." type="journal article" year="2016">
|
|
Reischütz
|
|
<emphasis id="B900EAF1070AFFFFFE34FD3FED58FD3E" box="[488,541,735,762]" italics="true" pageId="3" pageNumber="29">et al</emphasis>
|
|
. 2016
|
|
</bibRefCitation>
|
|
). This temperate region is home to four highly endemic species (
|
|
<figureCitation id="134F2A66070AFFFFFAB1FD00EFE2FCD8" captionStart="Figure 3" captionStartId="3.[151,226,1408,1432]" captionTargetBox="[154,1441,845,1385]" captionTargetId="figure-294@3.[151,1444,842,1389]" captionTargetPageId="3" captionText="Figure 3. Occurrence of highly endemic Phaedusinae species along the northeastern Aegean and the Black Sea coast. Graecophaedusa sperrlei Rähle, 1982 (1); Nothoserrulina subterranea Németh & Szekeres, 1995 (2); Cotyorica nemethi n. sp. (3); Pontophaedusella ofensis Nordsieck, 1994 (4); Truncatophaedusa evae Majoros, Németh & Szili- Kovács, 1994 (5)." figureDoi="http://doi.org/10.5281/zenodo.8027947" httpUri="https://zenodo.org/record/8027947/files/figure.png" pageId="3" pageNumber="29">Fig. 3</figureCitation>
|
|
), all of which live in shallow subterranean habitats (
|
|
<bibRefCitation id="EFE54B12070AFFFFFCFAFCDEEB6EFCD8" author="Culver, D. C. & Pipan, T." box="[806,1067,769,796]" pageId="3" pageNumber="29" refId="ref1585" refString="Culver, D. C. & Pipan, T. (2014) Shallow subterranean habitats: Ecology, evolution and conservation. Oxford University Press, 258 pp." type="book" year="2014">Culver & Pipan 2014</bibRefCitation>
|
|
) and, in accordance with their relict status, belong to monotypic genera.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |