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<document id="E743F37210916D2757EABB59E3B62B88" ID-CLB-Dataset="45246" ID-DOI="10.11646/zootaxa.3835.1.6" ID-GBIF-Dataset="725cafb5-06c8-4e80-af6b-259f0eb07575" ID-ISSN="1175-5326" ID-Zenodo-Dep="230695" ID-ZooBank="98E76FDF-C0D5-4FF4-BDB0-A3FECD5D2B63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460455512981" checkinUser="plazi" docAuthor="Almeida, Alexandre O., Anker, Arthur &amp; Mantelatto, Fernando L." docDate="2014" docId="03DD87F4DC1088766989FA0A1152FD31" docLanguage="en" docName="zt03835p120.pdf" docOrigin="Zootaxa 3835 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typton fapespae Almeida, Anker &amp; Mantelatto, 2014, sp. nov." docType="treatment" docUuid="274B3BF8-944B-4589-8C41-ABE9287E4369" docUuidSource="ZooBank" docVersion="9" lastPageNumber="119" masterDocId="FFE4FF8CDC11887F691EFFA21035FF99" masterDocTitle="A new snapping species of the shrimp genus Typton Costa, 1844 (Decapoda: Palaemonidae) from the coast of São Paulo, southeastern Brazil" masterLastPageNumber="120" masterPageNumber="110" pageNumber="111" updateTime="1698338559500" updateUser="ExternalLinkService">
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<mods:title id="AC22402D9832E8E25CD3B53973DE91AF">A new snapping species of the shrimp genus Typton Costa, 1844 (Decapoda: Palaemonidae) from the coast of São Paulo, southeastern Brazil</mods:title>
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<mods:namePart id="F2045A06ECD5D89369C6876C97D765A6">Almeida, Alexandre O.</mods:namePart>
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<mods:namePart id="CBBC85C6CC57EBA21A4D6FDE4B844B89">Anker, Arthur</mods:namePart>
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<mods:namePart id="D7B9C230607FCBD7DC32125443F2AA09">Mantelatto, Fernando L.</mods:namePart>
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<treatment id="03DD87F4DC1088766989FA0A1152FD31" ID-DOI="http://doi.org/10.5281/zenodo.6140847" ID-GBIF-Taxon="119575327" ID-Zenodo-Dep="6140847" LSID="urn:lsid:zoobank.org:act:274B3BF8-944B-4589-8C41-ABE9287E4369" httpUri="http://treatment.plazi.org/id/03DD87F4DC1088766989FA0A1152FD31" lastPageId="9" lastPageNumber="119" pageId="1" pageNumber="111">
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<heading id="D083818EDC10887E6989FA0A1186FA59" bold="true" box="[151,435,1447,1472]" fontSize="10" level="2" pageId="1" pageNumber="111" reason="2">
<emphasis id="B900EAF0DC10887E6989FA0A1186FA59" bold="true" box="[151,435,1447,1472]" pageId="1" pageNumber="111">
<taxonomicName id="4C744D61DC10887E6989FA0A1166FA59" ID-CoL="b113689a-95fa-424d-876b-71d7a8332170" box="[151,339,1447,1472]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="111" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC10887E6989FA0A1166FA59" bold="true" box="[151,339,1447,1472]" italics="true" pageId="1" pageNumber="111">Typton fapespae</emphasis>
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(
<figureCitation id="134F2A67DC10887E6981FA6E113DFA7C" box="[159,264,1484,1509]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1878,1900]" captionStartId-1="3.[151,250,1803,1825]" captionStartId-2="4.[151,250,1430,1452]" captionStartId-3="6.[151,250,1953,1975]" captionStartId-4="7.[151,250,1738,1760]" captionTargetBox-0="[218,1332,897,1829]" captionTargetBox-1="[166,1429,205,1756]" captionTargetBox-2="[169,1412,214,1386]" captionTargetBox-3="[183,1421,527,1916]" captionTargetBox-4="[167,1400,217,1694]" captionTargetId-0="figure@2.[208,1343,885,1858]" captionTargetId-1="figure@3.[157,1442,193,1780]" captionTargetId-2="figure@4.[163,1424,193,1408]" captionTargetId-3="figure@6.[183,1422,527,1918]" captionTargetId-4="figure@7.[167,1401,217,1695]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." captionText-1="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." captionText-2="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." captionText-3="FIGURE 4. Typton fapespae sp. nov., male holotype [MZUSP 31127] from São Sebastião, São Paulo, Brazil: A, habitus in life, dorsal view; B, same, lateral view. Photographs: A. Anker." captionText-4="FIGURE 5. Typton fapespae sp. nov., male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker." httpUri-0="https://zenodo.org/record/230696/files/figure.png" httpUri-1="https://zenodo.org/record/230697/files/figure.png" httpUri-2="https://zenodo.org/record/230698/files/figure.png" httpUri-3="https://zenodo.org/record/230699/files/figure.png" httpUri-4="https://zenodo.org/record/230700/files/figure.png" pageId="1" pageNumber="111">Figs. 15</figureCitation>
)
</paragraph>
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<taxonomicName id="4C744D61DC10887E6989F9B711ADF9B3" ID-CoL="965c8f67-54bd-405f-9086-db0b2799dab0" box="[151,408,1556,1578]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="111" phylum="Arthropoda" rank="species" species="gnathophylloides">
<emphasis id="B900EAF0DC10887E6989F9B711ADF9B3" box="[151,408,1556,1578]" italics="true" pageId="1" pageNumber="111">Typton gnathophylloides</emphasis>
</taxonomicName>
<treatmentCitation id="0AD510F3DC10887E68ADF9B612F5F9B3" author="Nalesso" box="[435,704,1556,1578]" page="96" pageId="1" pageNumber="111" year="1995">
<bibRefCitation id="EFE54B13DC10887E68ADF9B612A7F9B3" author="Nalesso" box="[435,658,1556,1578]" pageId="1" pageNumber="111" refString="Nalesso, R. C., Duarte, L. F. L., Pierozzi, I. Jr. &amp; Enumo, E. F. (1995) Tube epifauna of the polychaete Phyllochaetopterus socialis Claparede. Estuarine, Coastal and Shelf Science, 41, 91 - 100. http: // dx. doi. org / 10.1006 / ecss. 1995.0055" type="journal article" year="1995">
Nalesso
<emphasis id="B900EAF0DC10887E6B0FF9B71278F9B3" box="[529,589,1556,1578]" italics="true" pageId="1" pageNumber="111">et al.</emphasis>
1995
</bibRefCitation>
: 96
</treatmentCitation>
;
<treatmentCitation id="0AD510F3DC10887E6BCAF9B7142EF9B3" author="Duarte" box="[724,1051,1556,1578]" page="143" pageId="1" pageNumber="111" year="1996">
<bibRefCitation id="EFE54B13DC10887E6BCAF9B713D5F9B3" author="Duarte" box="[724,992,1556,1578]" pageId="1" pageNumber="111" refString="Duarte, L. F. L. &amp; Nalesso, R. C. (1996) The Sponge Zygomycale parishii (Bowerbank) and its endobiotic fauna. Estuarine, Coastal and Shelf Science, 42, 139 - 151. http: // dx. doi. org / 10.1006 / ecss. 1996.0011" type="journal article" year="1996">Duarte &amp; Nalesso 1996</bibRefCitation>
: 143
</treatmentCitation>
;
<treatmentCitation id="0AD510F3DC10887E6D30F9B61577F9B3" author="Amaral" box="[1070,1346,1556,1578]" page="249" pageId="1" pageNumber="111" year="2010">
<bibRefCitation id="EFE54B13DC10887E6D30F9B6153EF9B3" author="Amaral" box="[1070,1291,1556,1578]" pageId="1" pageNumber="111" refString="Amaral, A. C. Z., Migotto, A. E., Turra, A. &amp; Schaeffer-Novelli, Y. (2010) Araca: biodiversidade, impactos e ameacas. Biota Neotropica, 10, 219 - 264. http: // dx. doi. org / 10.1590 / s 1676 - 06032010000100022" type="journal article" year="2010">
Amaral
<emphasis id="B900EAF0DC10887E6D97F9B714F0F9B3" box="[1161,1221,1556,1578]" italics="true" pageId="1" pageNumber="111">et al.</emphasis>
2010
</bibRefCitation>
: 249
</treatmentCitation>
[not
<taxonomicName id="4C744D61DC10887E6C97F9B71221F9D1" ID-CoL="965c8f67-54bd-405f-9086-db0b2799dab0" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="111" phylum="Arthropoda" rank="species" species="gnathophylloides">
<emphasis id="B900EAF0DC10887E6C97F9B71140F9D1" italics="true" pageId="1" pageNumber="111">T. gnathophylloides</emphasis>
<bibRefCitation id="EFE54B13DC10887E6862F9901221F9D1" author="Holthuis" box="[380,532,1586,1608]" pageId="1" pageNumber="111" refString="Holthuis, L. B. (1951) A general revision of the Palaemonidae (Crustacea Decapoda Natantia) of the Americas. I. The subfamilies Euryrhynchidae and Pontoniinae. Allan Hancock Foundation Publications of the University of Southern California, Occasional Paper, 11, 1 - 332." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
</taxonomicName>
]
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material.
</emphasis>
<collectingCountry id="F3637672DC10887E684AF9D711AEF917" box="[340,411,1653,1678]" name="Brazil" pageId="1" pageNumber="111">Brazil</collectingCountry>
, coast of São Paulo.
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: male (pocl/cl 3.0/
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), São Sebastião, off Praia do Cabelo Gordo,
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, free diving, depth: less than
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, in sponge growing on large rock, coll. M. Tavares, J.B. Mendonça, A. Anker and P.P.G. Pachelle,
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[
<collectionCode id="ED65AE27DC10887E6D1BF91C1450F94F" box="[1029,1125,1726,1750]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="111">MZUSP</collectionCode>
31127].
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:
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(pocl/cl 2.6/
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, 2.8/
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), 2 ovigerous females (pocl/cl 2.3/
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, 3.4/4.0 mm), same collection data as for the
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[
<collectionCode id="ED65AE27DC10887E6840F8A4118BF887" box="[350,446,1798,1822]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="111">MZUSP</collectionCode>
3112831131];
<specimenCount id="9D72FD6BDC10887E6B64F8A412F2F887" box="[634,711,1797,1822]" pageId="1" pageNumber="111" type="male">1 male</specimenCount>
(pocl/cl 2.4/
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), Ubatuba, Praia do Cedro,
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, scuba diving, depth:
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, in
<taxonomicName id="4C744D61DC10887E6BBEF88B1437F8DB" box="[672,1026,1833,1858]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="1" pageNumber="111" phylum="Porifera" rank="species" species="angulosa" subGenus="Zygomycale">
<emphasis id="B900EAF0DC10887E6BBEF88B1437F8DB" box="[672,1026,1833,1858]" italics="true" pageId="1" pageNumber="111">Mycale (Zygomycale) angulosa</emphasis>
</taxonomicName>
, coll. F. Zara,
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[
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4486];
<specimenCount id="9D72FD6BDC10887E69F9F8EC117DF8FF" box="[231,328,1869,1894]" pageId="1" pageNumber="111" type="female">1 female</specimenCount>
(pocl/cl 1.9/
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), Ubatuba, Praia de Itaguá,
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, scuba diving, depth: less than
<quantity id="4C8C9B07DC10887E69CEF8D010CAF810" box="[208,255,1906,1930]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="111" unit="m" value="3.0">3 m</quantity>
, in
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<emphasis id="B900EAF0DC10887E6832F8D31226F810" box="[300,531,1905,1930]" italics="true" pageId="1" pageNumber="111">Schizoporella errata</emphasis>
(
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)
</taxonomicName>
(Gymnolaemata:
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), coll. I. Leone
<emphasis id="B900EAF0DC10887E6C14F8D01571F813" box="[1290,1348,1905,1930]" italics="true" pageId="1" pageNumber="111">et al.</emphasis>
,
<date id="FFCA1022DC10887E6C4AF8D310FBF837" pageId="1" pageNumber="111" value="2011-04-05">5 Apr. 2011</date>
[
<collectionCode id="ED65AE27DC10887E69C5F834111FF837" box="[219,298,1942,1966]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="1" pageNumber="111">CCDB</collectionCode>
3413].
</paragraph>
<paragraph id="8BCB36E2DC10887D69D9F81B1371FEFD" blockId="1.[151,1437,1653,2038]" lastBlockId="2.[151,1437,151,824]" lastPageId="2" lastPageNumber="112" pageId="1" pageNumber="111">
<emphasis id="B900EAF0DC10887E69D9F81B118FF84B" bold="true" box="[199,442,1977,2002]" pageId="1" pageNumber="111">Additional material.</emphasis>
<specimenCount id="9D72FD6BDC10887E68DFF8181222F84B" box="[449,535,1977,2002]" pageId="1" pageNumber="111" type="male">2 males</specimenCount>
(pocl/cl 4.2/
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),
<specimenCount id="9D72FD6BDC10887E6A0DF818134AF84B" box="[787,895,1977,2002]" pageId="1" pageNumber="111" type="female">2 females</specimenCount>
(pocl/cl 4.0/
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), Ubatuba, Praia Lamberto,
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<geoCoordinate id="EE405025DC10887E6853F87F11D6F86F" box="[333,483,2013,2038]" direction="west" orientation="longitude" pageId="1" pageNumber="111" precision="1" value="-45.11736">45°72.50”W</geoCoordinate>
,
<date id="FFCA1022DC10887E68EBF87F12BCF86F" box="[501,649,2013,2038]" pageId="1" pageNumber="111" value="1975-09-05">5 Sep. 1975</date>
[
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<collectionCode id="ED65AE27DC10887E6BE6F87C1307F86F" box="[760,818,2014,2038]" pageId="1" pageNumber="111">CRU</collectionCode>
14];
<specimenCount id="9D72FD6BDC10887E6A65F87C13EDF86F" box="[891,984,2013,2038]" pageId="1" pageNumber="111" type="male">2 males</specimenCount>
(pocl/cl 4.1/
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, 3.6/
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),
<specimenCount id="9D72FD6BDC10887D6C90F87C1139FF29" lastPageId="2" lastPageNumber="112" pageId="1" pageNumber="111" type="generic">3 specimens</specimenCount>
in poor condition, São Sebastião, Praia do Araçá,
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<geoCoordinate id="EE405025DC13887D6AE4FF3514AAFF29" box="[1018,1183,151,176]" direction="west" orientation="longitude" pageId="2" pageNumber="112" precision="1" value="-45.41303">45°2446.9”W</geoCoordinate>
,
<date id="FFCA1022DC13887D6DB3FF351575FF29" box="[1197,1344,151,176]" pageId="2" pageNumber="112" value="1975-01-26">26 Jan. 1975</date>
[
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<collectionCode id="ED65AE27DC13887D6989FF1E10E4FF4D" box="[151,209,188,212]" pageId="2" pageNumber="112">CRU</collectionCode>
20];
<specimenCount id="9D72FD6BDC13887D680FFF1E115FFF4C" box="[273,362,188,213]" pageId="2" pageNumber="112" type="male">9 males</specimenCount>
(pocl/cl
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/
<quantity id="4C8C9B07DC13887D6B78FF1E12E8FF4D" box="[614,733,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.4" metricValueMin="2.0" pageId="2" pageNumber="112" unit="mm" value="2.7" valueMax="3.4" valueMin="2.0">23.4 mm</quantity>
),
<specimenCount id="9D72FD6BDC13887D6BECFF1F1354FF4C" box="[754,865,188,213]" pageId="2" pageNumber="112" type="female">7 females</specimenCount>
(pocl/cl 2.84.2/3.64.7) São Sebastião, Praia do Araçá,
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<geoCoordinate id="EE405025DC13887D6897FF7D121BFF61" box="[393,558,223,248]" direction="west" orientation="longitude" pageId="2" pageNumber="112" precision="1" value="-45.41303">45°2446.9”W</geoCoordinate>
,
<date id="FFCA1022DC13887D6B24FF4212FBFF61" box="[570,718,224,248]" pageId="2" pageNumber="112" value="1974-09-14">14 Sep. 1974</date>
[
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<collectionCode id="ED65AE27DC13887D6A31FF42135CFF61" box="[815,873,224,248]" pageId="2" pageNumber="112">CRU</collectionCode>
21];
<specimenCount id="9D72FD6BDC13887D6AB8FF42143CFF61" box="[934,1033,223,248]" pageId="2" pageNumber="112" type="male">11 males</specimenCount>
(pocl/cl
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/
<quantity id="4C8C9B07DC13887D6C1BFF7D15A5FF61" box="[1285,1424,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.1" metricValueMin="3.4" pageId="2" pageNumber="112" unit="mm" value="4.25" valueMax="5.1" valueMin="3.4">3.45.1 mm</quantity>
),
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(pocl/cl
<quantity id="4C8C9B07DC13887D6875FEA611C7FE85" box="[363,498,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="2" pageNumber="112" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.14.9 mm</quantity>
/
<quantity id="4C8C9B07DC13887D6B1DFEA612A5FE85" box="[515,656,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="2" pageNumber="112" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.65.4 mm</quantity>
), São Sebastião, Praia do Araçá,
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<geoCoordinate id="EE405025DC13887D6DB3FEA61564FE84" box="[1197,1361,260,285]" direction="west" orientation="longitude" pageId="2" pageNumber="112" precision="1" value="-45.41303">45°2446.9”W</geoCoordinate>
,
<date id="FFCA1022DC13887D6C40FEA610FAFED9" pageId="2" pageNumber="112" value="1974-07-06">6 Jul. 1974</date>
[
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<collectionCode id="ED65AE27DC13887D6828FE8A1145FED9" box="[310,368,296,320]" pageId="2" pageNumber="112">CRU</collectionCode>
22];
<specimenCount id="9D72FD6BDC13887D68AFFE8A11CAFED9" box="[433,511,295,320]" pageId="2" pageNumber="112" type="male">1 male</specimenCount>
(pocl/cl 4.2/
<quantity id="4C8C9B07DC13887D6B8BFE8512C7FED9" box="[661,754,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="112" unit="mm" value="4.6">4.6 mm</quantity>
),
<specimenCount id="9D72FD6BDC13887D6A16FE8A1359FED9" box="[776,876,295,320]" pageId="2" pageNumber="112" type="female">1 female</specimenCount>
(pocl/cl 3.8/
<quantity id="4C8C9B07DC13887D6D1FFE8A146BFED9" box="[1025,1118,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="112" unit="mm" value="4.4">4.4 mm</quantity>
), Ubatuba, Praia Lamberto,
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<geoCoordinate id="EE405025DC13887D685BFEEE11EEFEFC" box="[325,475,332,357]" direction="west" orientation="longitude" pageId="2" pageNumber="112" precision="1" value="-45.11736">45°72.50”W</geoCoordinate>
,
<date id="FFCA1022DC13887D68F9FEEE1242FEFD" box="[487,631,332,356]" pageId="2" pageNumber="112" value="1974-06-10">10 Jun. 1974</date>
[
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<collectionCode id="ED65AE27DC13887D6BC9FEEE1324FEFD" box="[727,785,332,356]" pageId="2" pageNumber="112">CRU</collectionCode>
31].
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC13887769D9FECD1575FEFC" lastPageId="8" lastPageNumber="118" pageId="2" pageNumber="112" type="description">
<paragraph id="8BCB36E2DC13887D69D9FECD1222FD31" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="112">
<emphasis id="B900EAF0DC13887D69D9FECD116DFE11" bold="true" box="[199,344,367,392]" pageId="2" pageNumber="112">Description.</emphasis>
Carapace smooth, glabrous, slightly depressed, longer than deep. Rostrum arrow-shaped in dorsal view (
<figureCitation id="134F2A67DC13887D6837FE36115DFE35" box="[297,360,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
A), usually reaching base of cornea and sometimes extending beyond distal margin of eyes and half-length of dorsally visible part of first article of antennular peduncle, straight to slightly curving upwards, tip with single small seta (
<figureCitation id="134F2A67DC13887D68B9FE7E11DCFE6D" box="[423,489,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
AC). Paraorbital processes (
<figureCitation id="134F2A67DC13887D6A5FFE7E13B7FE6D" box="[833,898,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
AC) well developed, usually exceeding halflength of rostrum and half-length of cornea in lateral view, reaching level of most-anterior pterygostomial margin, non-divergent in dorsal view, pointing straight-forward or slightly upward in lateral view (
<figureCitation id="134F2A67DC13887D6D8DFD8614E6FDA5" box="[1171,1235,548,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
AC). Orbit (
<figureCitation id="134F2A67DC13887D6C71FD861097FDF9" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
B, C) feebly demarcated, without distinct infero-orbital angle or antennal spine; pterygostomial margin (
<figureCitation id="134F2A67DC13887D6C21FDE5154AFDF9" box="[1343,1407,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
B, C) slightly produced anteriorly, rounded to somewhat angular. Posteroventral margin of carapace rounded, with slight cardiac concavity (
<figureCitation id="134F2A67DC13887D68AEFD2D11C5FD31" box="[432,496,655,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
D).
</paragraph>
<paragraph id="8BCB36E2DC13887D69D9FD1614BBFCAE" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="112">
Abdomen (
<figureCitation id="134F2A67DC13887D685BFD1611B6FD55" box="[325,387,692,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
E) elongate, smooth, glabrous; first five somites (
<figureCitation id="134F2A67DC13887D6AAFFD1613DAFD55" box="[945,1007,692,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
E) with rounded pleura; posterodorsal margin of sixth somite (
<figureCitation id="134F2A67DC13887D68ABFD7511CDFD69" box="[437,504,727,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
F) with strong, sharp, subtriangular, median tooth; posterolateral angle of sixth somite (
<figureCitation id="134F2A67DC13887D69ECFD5E1105FC8D" box="[242,304,764,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112">Fig. 1</figureCitation>
EG) produced into strong acute tooth, posteroventral angle blunt, not posteriorly produced; first to third and sixth sternites unarmed, fourth and fifth each with low, rounded, median process.
</paragraph>
<caption id="DF0B666ADC13887D6989F8F41563F87E" httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="2" pageNumber="112" targetBox="[218,1332,897,1829]" targetPageId="2">
<paragraph id="8BCB36E2DC13887D6989F8F41563F87E" blockId="2.[151,1436,1878,2023]" pageId="2" pageNumber="112">
<emphasis id="B900EAF0DC13887D6989F8F41121F8F5" bold="true" box="[151,276,1878,1900]" pageId="2" pageNumber="112">FIGURE 1.</emphasis>
<taxonomicName id="4C744D61DC13887D6805F8FA11F5F8F5" box="[283,448,1878,1901]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="112" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC13887D6805F8FA11F5F8F5" box="[283,448,1878,1901]" italics="true" pageId="2" pageNumber="112">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC13887D68D6F8F5122CF8F5" bold="true" box="[456,537,1879,1900]" pageId="2" pageNumber="112">
<taxonomicNameLabel id="A233578BDC13887D68D6F8F5122CF8F5" box="[456,537,1879,1900]" pageId="2" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype [CCDB 4486] (A, B, DJ) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: AE = 0.5 mm, FH = 0.25 mm, I, J = 0.1 mm.
</paragraph>
</caption>
<caption id="DF0B666ADC12887C6989F8A910D8F840" httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="3" pageNumber="113" targetBox="[166,1429,205,1756]" targetPageId="3">
<paragraph id="8BCB36E2DC12887C6989F8A910D8F840" blockId="3.[151,1436,1803,2009]" pageId="3" pageNumber="113">
<emphasis id="B900EAF0DC12887C6989F8A91126F8B9" bold="true" box="[151,275,1803,1825]" pageId="3" pageNumber="113">FIGURE 2.</emphasis>
<taxonomicName id="4C744D61DC12887C6804F8AE118AF8B8" box="[282,447,1803,1825]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC12887C6804F8AE118AF8B8" box="[282,447,1803,1825]" italics="true" pageId="3" pageNumber="113">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC12887C68DBF8AE1223F8B8" bold="true" box="[453,534,1804,1825]" pageId="3" pageNumber="113">
<taxonomicNameLabel id="A233578BDC12887C68DBF8AE1223F8B8" box="[453,534,1804,1825]" pageId="3" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype [CCDB 3413] (A, B, E, KN) and male paratype [CCDB 4486] (C, D, FJ) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm.
</paragraph>
</caption>
<caption id="DF0B666ADC15887B6989FA3412B7F9FD" httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="4" pageNumber="114" targetBox="[169,1412,214,1386]" targetPageId="4">
<paragraph id="8BCB36E2DC15887B6989FA3412B7F9FD" blockId="4.[151,1437,1430,1636]" pageId="4" pageNumber="114">
<emphasis id="B900EAF0DC15887B6989FA341120FA32" bold="true" box="[151,277,1430,1452]" pageId="4" pageNumber="114">FIGURE 3.</emphasis>
<taxonomicName id="4C744D61DC15887B6800FA3511F3FA35" box="[286,454,1430,1452]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="114" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC15887B6800FA3511F3FA35" box="[286,454,1430,1452]" italics="true" pageId="4" pageNumber="114">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC15887B68D1FA341217FA32" bold="true" box="[463,546,1430,1451]" pageId="4" pageNumber="114">
<taxonomicNameLabel id="A233578BDC15887B68D1FA341217FA32" box="[463,546,1430,1451]" pageId="4" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype [CCDB 4486] (AJ) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: AC = 0.5 mm, DH = 0.25 mm, IL = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BCB36E2DC15887B69D9F9321414F8C4" blockId="4.[151,1436,1680,2029]" pageId="4" pageNumber="114">
Telson (
<figureCitation id="134F2A67DC15887B683CF9321157F930" box="[290,354,1680,1705]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="4" pageNumber="114">Fig. 1</figureCitation>
H) moderately slender, about 2.2 times as long as anterior width, posterior margin width about 0.35 times width of anterior margin; lateral margins slightly convex, tapering distally; dorsal surface with two pairs of well-developed spines, each about 0.1 of telson length, inserted at about 0.25 (anterior pair) and 0.6 (posterior pair) telson length; posterior margin (
<figureCitation id="134F2A67DC15887B6B5DF95E12B0F88C" box="[579,645,1788,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="4" pageNumber="114">
Fig.
<date id="FFCA1022DC15887B6B67F95F12B0F88C" box="[633,645,1789,1813]" pageId="4" pageNumber="114">1</date>
</figureCitation>
I, J) broadly rounded with minute median point; three pairs of spines present as following: small lateral spines and longer intermediate and submedian spines, intermediate being 2.02.8 times as long as lateral and 1.31.6 times as long as submedian (
<figureCitation id="134F2A67DC15887B6ADAF8E61430F8C4" box="[964,1029,1860,1885]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="4" pageNumber="114">
Fig.
<date id="FFCA1022DC15887B6AE7F8E71430F8C4" box="[1017,1029,1861,1885]" pageId="4" pageNumber="114">1</date>
</figureCitation>
I).
</paragraph>
<paragraph id="8BCB36E2DC15887A69D9F8CA1524FE84" blockId="4.[151,1436,1680,2029]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="115" pageId="4" pageNumber="114">
Antennular peduncle (
<figureCitation id="134F2A67DC15887B68DAF8CA123AF818" box="[452,527,1896,1921]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="4" pageNumber="114">Figs. 1</figureCitation>
A, 2A) slender, with proximal article subcylindrical, 3.8 times as long as distal width; statocyst well-developed, with statolith; stylocerite short, narrow; intermediate article shorter than distal article, their combined length equal to about 0.4 of proximal article length; lateral (= upper) flagellum (
<figureCitation id="134F2A67DC15887B6C2CF8121548F850" box="[1330,1405,1968,1993]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="4" pageNumber="114">Figs. 1</figureCitation>
A, 2A) short, first two articles of rami fused, stout, with 78 groups of aesthetascs, short ramus distally obsolete, longer ramus with 56 segments; mesial (= lower) flagellum long, with 910 segments. Antenna with antennal gland tubercle mesially; basicerite stout, unarmed laterally; scaphocerite (
<figureCitation id="134F2A67DC14887A6AC6FF1E1429FF4D" box="[984,1052,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C, D) vestigial, about 1.8 times as long as greatest width, terminally obliquely rounded, margins non-setose; carpocerite (
<figureCitation id="134F2A67DC14887A6DBCFF7D14D9FF61" box="[1186,1260,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
B) slender, not reaching end of antennular peduncle; flagellum (
<figureCitation id="134F2A67DC14887A6BA6FEA612CFFE85" box="[696,762,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
A) short, about 2.6 times as long as carpocerite.
</paragraph>
<paragraph id="8BCB36E2DC14887A69D9FE851578FE11" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
Mouthparts not dissected. Third maxilliped (
<figureCitation id="134F2A67DC14887A6BA1FE851337FED9" box="[703,770,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
E) with broad antepenultimate article, with both dorsal and ventral margins strongly convex; penultimate article about half as long as antepenultimate article, much more slender, with straight margins, ultimate article most slender, tapering towards apex; exopod well developed.
</paragraph>
<paragraph id="8BCB36E2DC14887A69D9FE361487FD81" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
First pereiopod (
<figureCitation id="134F2A67DC14887A689FFE3611F1FE35" box="[385,452,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
F) slender; ischium about three times as long as wide, unarmed; merus about five times as long as broad, slightly longer than carpus; carpus subcylindrical, broadening distally, about four times as long as broad and 1.5 times as long as chela; carpo-propodal brush lacking; chela with palm subcylindrical, slightly compressed; fingers robust, subspatulate, about 0.4 of chela length, with feebly bidentate tips.
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<paragraph id="8BCB36E2DC14887A69D9FD8613E8FC5E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
Major second pereiopod (
<figureCitation id="134F2A67DC14887A68F7FD86120DFDA5" box="[489,568,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Figs. 2</figureCitation>
GJ, 4, 5) greatly enlarged, much longer and more robust than minor second pereiopod; largest adult males with massive chela equaling combined length of cephalothorax and abdomen (
<figureCitation id="134F2A67DC14887A6C7AFDE51092FD1D" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[183,1421,527,1916]" captionTargetId="figure@6.[183,1422,527,1918]" captionTargetPageId="6" captionText="FIGURE 4. Typton fapespae sp. nov., male holotype [MZUSP 31127] from São Sebastião, São Paulo, Brazil: A, habitus in life, dorsal view; B, same, lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230699/files/figure.png" pageId="5" pageNumber="115">Figs. 4</figureCitation>
,
<figureCitation id="134F2A67DC14887A69ACFDCE10F5FD1C" box="[178,192,620,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1738,1760]" captionTargetBox="[167,1400,217,1694]" captionTargetId="figure@7.[167,1401,217,1695]" captionTargetPageId="7" captionText="FIGURE 5. Typton fapespae sp. nov., male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230700/files/figure.png" pageId="5" pageNumber="115">5</figureCitation>
A), females with much smaller chela, equaling or slightly exceeding length of cephalothorax (
<figureCitation id="134F2A67DC14887A6DC9FDCE152FFD1C" box="[1239,1306,620,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1738,1760]" captionTargetBox="[167,1400,217,1694]" captionTargetId="figure@7.[167,1401,217,1695]" captionTargetPageId="7" captionText="FIGURE 5. Typton fapespae sp. nov., male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230700/files/figure.png" pageId="5" pageNumber="115">Fig. 5</figureCitation>
B); ischium subequal to merus in length, unarmed; merus (
<figureCitation id="134F2A67DC14887A6BBFFD2D12D0FD31" box="[673,741,655,680]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">
Fig.
<date id="FFCA1022DC14887A6BC8FD3212D0FD31" box="[726,741,656,680]" pageId="5" pageNumber="115">2</date>
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I) about 1.3 times as long as broad, about 0.6 length of carpus and 0.2 length of palm, ventral surface with small, conical, distally more or less sharp teeth; carpus (
<figureCitation id="134F2A67DC14887A6C04FD161568FD55" box="[1306,1373,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
G, H) 0.30.4 length of palm, smooth, 1.6 times as long as maximal width, narrow proximally, distally widening, distoventral surface somewhat excavated; chela (
<figureCitation id="134F2A67DC14887A6BDEFD5E1337FC8D" box="[704,770,764,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
G, H, J) very large, with palm smooth, slightly compressed laterally, suboval in cross-section, flattened medially, slightly tapering distally, about twice as long as maximal height; dactylus (
<figureCitation id="134F2A67DC14887A6842FCE61195FCC5" box="[348,416,836,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
J) laterally compressed, about twice as long as maximal height, about 0.4 times length of palm; tip strongly curved, corneous; cutting edge conspicuously thickened, forming large bulge; pollex about as long as dactylus; tip curved, corneous; surface opposed to dactylus with relatively large and deep groove accommodating dactylar bulge, mesial and lateral margins entire, concave.
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<paragraph id="8BCB36E2DC14887A69D9FC761306FB71" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
Minor second pereiopod variously but always significantly smaller than major second pereiopod (
<figureCitation id="134F2A67DC14887A6C0FFC76156AFC75" box="[1297,1375,980,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Figs. 2</figureCitation>
KN, 4, 5); ischium (
<figureCitation id="134F2A67DC14887A6856FC5511B9FB89" box="[328,396,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
K) about 1.4 times length of merus, unarmed; merus (
<figureCitation id="134F2A67DC14887A6AE5FC551475FB89" box="[1019,1088,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
K) 1.5 times as long as broad, about 0.7 of carpal length and 0.4 of palm length, with minute conical teeth on ventral surface; carpus (
<figureCitation id="134F2A67DC14887A6C09FBBE156CFBAD" box="[1303,1369,1052,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
K, L) about 0.7 length of palm, smooth, about twice as long as maximal height, much wider distally; palm (
<figureCitation id="134F2A67DC14887A6C05FB9D156BFBC1" box="[1307,1374,1087,1112]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
K, L) smooth, non-setose, suboval in cross-section, about 1.8 times as long as maximal height; dactylus (
<figureCitation id="134F2A67DC14887A6C09FBC6156EFBE5" box="[1303,1371,1124,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
KN) about 0.6 length of palm, compressed, with curved tip; cutting edge sharp, with blunt proximal tooth laterally; pollex excavated on surface opposed to dactylus; lateral edge sharp, nearly entire; mesial edge projecting proximally as strong subtriangular tooth; finger tips simple.
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<paragraph id="8BCB36E2DC14887A69D9FB561220F925" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
Third pereiopod (
<figureCitation id="134F2A67DC14887A6890FB5611E4FA95" box="[398,465,1268,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
A, D) stout, with ischium about 0.8 times length of merus, unarmed; merus (
<figureCitation id="134F2A67DC14887A6C25FB56154BFA95" box="[1339,1406,1268,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
A) about twice as long as broad and 1.2 times length of carpus, unarmed; carpus (
<figureCitation id="134F2A67DC14887A6D57FAB514A5FAA9" box="[1097,1168,1303,1328]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
A) about 0.9 length of propodus, nearly three times as long as distal width, unarmed; propodus (
<figureCitation id="134F2A67DC14887A6AFCFA9E1410FACD" box="[994,1061,1340,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
A, D) about 2.8 times as long as wide, ventral margin with four or five irregularly spaced spines, distal pair of stouter spines fringing base of dactylus; dactylus (
<figureCitation id="134F2A67DC14887A6871FA26119AFA05" box="[367,431,1412,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
A, D) biunguiculate, compressed, about 0.25 of propodus length and 2.2 times longer than basal width, ventral margin of corpus slightly concave, crenulated, with acute, triangular, ventrally pointing secondary unguis; terminal unguis distinctly demarcated, about half as long as corpus, strongly curved, not crenulated proximally. Fourth pereiopod (
<figureCitation id="134F2A67DC14887A6B6FFA4D1283F991" box="[625,694,1519,1544]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
B, E) generally similar to third pereiopod; merus (
<figureCitation id="134F2A67DC14887A6DEBFA4D150DF991" box="[1269,1336,1519,1544]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
B) about 2.4 times as long as wide; carpus (
<figureCitation id="134F2A67DC14887A6B09F9B6126CF9B5" box="[535,601,1556,1581]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
B) slightly longer than propodus, about three times as long as distal width; propodus (
<figureCitation id="134F2A67DC14887A6811F9951164F9C9" box="[271,337,1591,1616]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
B, E) about three times as long as wide; ventral margin with three or four spines and distal pair of stouter spines; dactylus (
<figureCitation id="134F2A67DC14887A68AFF9FE11C0F9ED" box="[433,501,1628,1653]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="5" pageNumber="115">Fig. 3</figureCitation>
B, E) biunguiculate, nearly twice as long as basal width; ventral margin of corpus conspicuously crenulated, with triangular, ventrally pointing secondary unguis; terminal unguis equal or slightly shorter than half-length of corpus.
</paragraph>
<paragraph id="8BCB36E2DC14887A69D9F96513EAF844" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="115">
Fifth pereiopod (
<figureCitation id="134F2A67DC14887A6899F96511F8F979" box="[391,461,1735,1760]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C, F) noticeably more slender than third and fourth, with ischium about 0.6 length of merus, unarmed; merus (
<figureCitation id="134F2A67DC14887A68AEF94E11C6F89D" box="[432,499,1772,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C) about 3.5 times as long as wide and 1.4 length of carpus, unarmed; carpus (
<figureCitation id="134F2A67DC14887A6C71F94E1090F8B1" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C) 0.8 times length to propodus, about 3.3 times as long as distal width, unarmed; propodus (
<figureCitation id="134F2A67DC14887A6DADF8AD14C0F8B1" box="[1203,1269,1807,1832]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C, F) about 4.5 times as long as broad; ventral margin with one subdistal and one distal spines; distolateral surface with brush of grooming setae; dactylus (
<figureCitation id="134F2A67DC14887A68A2F8F511CAF8E9" box="[444,511,1879,1904]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="5" pageNumber="115">Fig. 2</figureCitation>
C, F) biunguiculate, compressed, about 0.2 of propodus length, 2.3 times as long as basal width; ventral margin of corpus slightly convex, distally conspicuously crenulated, with strong, subacute, ventrally pointing secondary unguis; terminal unguis well demarcated, more slender than that of third and fourth pereiopods, about half-length of corpus, curved, not crenulated proximally.
</paragraph>
<paragraph id="8BCB36E2DC17887969D9FF3513D5FED9" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="116">
First male pleopod (
<figureCitation id="134F2A67DC17887968B1FF3511C7FF29" box="[431,498,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="6" pageNumber="116">Fig. 3</figureCitation>
G) with endopod not reaching half-length of exopod, about three times as long as proximal width, elongate, with rounded tip, lateral margin with four longer plumose setae, apex and mesial margin with several shorter setae. Second male pleopod (
<figureCitation id="134F2A67DC1788796BA1FF7D12CAFF61" box="[703,767,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="6" pageNumber="116">Fig. 3</figureCitation>
H) with endopod about 0.8 length of exopod, with appendix interna at about 0.4 length of mesial margin; appendix masculina with corpus obsolete, being essentially represented by two stiff setulose setae protruding beyond appendix interna.
</paragraph>
<paragraph id="8BCB36E2DC17887969D9FEEE1478FE56" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="116">
Uropods (
<figureCitation id="134F2A67DC1788796825FEEE11BCFEFD" box="[315,393,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="6" pageNumber="116">Figs. 1</figureCitation>
E,
<date id="FFCA1022DC17887968AEFEEE11F0FEFD" box="[432,453,332,356]" pageId="6" pageNumber="116">3I</date>
L) with protopod unarmed; exopod slightly longer than telson, with lateral margin slightly convex, non-setose; distolateral margin (
<figureCitation id="134F2A67DC1788796BA1FECD1334FE11" box="[703,769,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="6" pageNumber="116">
Fig.
<date id="FFCA1022DC1788796BEBFED21334FE11" box="[757,769,368,392]" pageId="6" pageNumber="116">3</date>
</figureCitation>
IL) serrated, with 2-6 acute or subacute teeth; distolateral angle with acute tooth, reaching approximately half-length of stout spine; diaeresis inconspicuous; endopod slightly longer than exopod; proximal parts of lateral and mesial margins non-setose.
</paragraph>
<caption id="DF0B666ADC1788796989F803132DF84F" httpUri="https://zenodo.org/record/230699/files/figure.png" pageId="6" pageNumber="116" targetBox="[183,1421,527,1916]" targetPageId="6">
<paragraph id="8BCB36E2DC1788796989F803132DF84F" blockId="6.[151,1436,1953,2006]" pageId="6" pageNumber="116">
<emphasis id="B900EAF0DC1788796989F8031121F82F" bold="true" box="[151,276,1953,1975]" pageId="6" pageNumber="116">FIGURE 4.</emphasis>
<taxonomicName id="4C744D61DC1788796802F80011F6F82E" box="[284,451,1953,1975]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="116" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1788796802F80011F6F82E" box="[284,451,1953,1975]" italics="true" pageId="6" pageNumber="116">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC17887968D2F800122BF82E" bold="true" box="[460,542,1954,1975]" pageId="6" pageNumber="116">
<taxonomicNameLabel id="A233578BDC17887968D2F800122BF82E" box="[460,542,1954,1975]" pageId="6" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype [MZUSP 31127] from São Sebastião, São Paulo, Brazil: A, habitus in life, dorsal view; B, same, lateral view. Photographs: A. Anker.
</paragraph>
</caption>
<caption id="DF0B666ADC1688786989F96810CEF884" httpUri="https://zenodo.org/record/230700/files/figure.png" pageId="7" pageNumber="117" targetBox="[167,1400,217,1694]" targetPageId="7">
<paragraph id="8BCB36E2DC1688786989F96810CEF884" blockId="7.[151,1437,1738,1821]" pageId="7" pageNumber="117">
<emphasis id="B900EAF0DC1688786989F9681121F946" bold="true" box="[151,276,1738,1760]" pageId="7" pageNumber="117">FIGURE 5.</emphasis>
<taxonomicName id="4C744D61DC1688786802F96911F7F979" box="[284,450,1738,1760]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="117" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1688786802F96911F7F979" box="[284,450,1738,1760]" italics="true" pageId="7" pageNumber="117">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC16887868D7F968122EF946" bold="true" box="[457,539,1738,1759]" pageId="7" pageNumber="117">
<taxonomicNameLabel id="A233578BDC16887868D7F968122EF946" box="[457,539,1738,1759]" pageId="7" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker.
</paragraph>
</caption>
<paragraph id="8BCB36E2DC16887769D9F8E61153FE85" blockId="7.[151,1436,1860,2029]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="118" pageId="7" pageNumber="117">
<emphasis id="B900EAF0DC16887869D9F8E6110EF8C4" bold="true" box="[199,315,1860,1885]" pageId="7" pageNumber="117">Variation.</emphasis>
The most obvious variable character is the length of the rostrum, which can extend beyond the distal margin of the cornea (
<figureCitation id="134F2A67DC16887868C1F8CA1215F818" box="[479,544,1896,1921]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
A, B) or reach only to the middle of the cornea (
<figureCitation id="134F2A67DC1688786D43F8CA14AAF818" box="[1117,1183,1896,1921]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
C, 5). The paraorbital processes can be about as long as half-length of cornea (
<figureCitation id="134F2A67DC1688786A2FF82E1346F83C" box="[817,883,1932,1957]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
A, B) or significantly shorter (
<figureCitation id="134F2A67DC1688786DC3F82E1515F83C" box="[1245,1312,1932,1957]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
C, 5A). In addition, both the rostrum and the paraorbital processes are variable in their direction, pointing either straightforward or slightly upward (cf.
<figureCitation id="134F2A67DC16887868E7F876120DF874" box="[505,568,2004,2029]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
B and 1C). The pterygostomial region varies from broadly rounded (
<figureCitation id="134F2A67DC1688786C23F8761548F874" box="[1341,1405,2004,2029]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="7" pageNumber="117">Fig. 1</figureCitation>
B) to more angularly protruding (
<figureCitation id="134F2A67DC19887768F6FF351212FF29" box="[488,551,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="8" pageNumber="118">Fig. 1</figureCitation>
C). The number of spines on the ventral margin of the P3 and P4 propodi varies from four to five and three to four, respectively. There is also a significant variation in the number of teeth on the distolateral margin of the uropodal exopod, ranging from 26 (not including terminal tooth adjacent to strong spine) (
<figureCitation id="134F2A67DC19887769F5FEA61119FE85" box="[235,300,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="8" pageNumber="118">
Fig.
<date id="FFCA1022DC1988776801FEA61119FE85" box="[287,300,260,284]" pageId="8" pageNumber="118">3</date>
</figureCitation>
IL).
</paragraph>
<paragraph id="8BCB36E2DC19887769D9FE851575FEFC" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FE8511B0FED9" bold="true" box="[199,389,295,320]" pageId="8" pageNumber="118">Colour pattern.</emphasis>
Uniform translucent yellowish to pale orange, with some scattered reddish and yellowish chromatophores (
<figureCitation id="134F2A67DC1988776845FEEE119FFEFD" box="[347,426,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[183,1421,527,1916]" captionTargetId="figure@6.[183,1422,527,1918]" captionTargetPageId="6" captionText="FIGURE 4. Typton fapespae sp. nov., male holotype [MZUSP 31127] from São Sebastião, São Paulo, Brazil: A, habitus in life, dorsal view; B, same, lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230699/files/figure.png" pageId="8" pageNumber="118">Figs. 4</figureCitation>
,
<figureCitation id="134F2A67DC19887768A8FEEE11F3FEFC" box="[438,454,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1738,1760]" captionTargetBox="[167,1400,217,1694]" captionTargetId="figure@7.[167,1401,217,1695]" captionTargetPageId="7" captionText="FIGURE 5. Typton fapespae sp. nov., male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230700/files/figure.png" pageId="8" pageNumber="118">5</figureCitation>
); cornea silvery reddish; ovaries and freshly laid eggs yellow-orange (
<figureCitation id="134F2A67DC1988776DC4FEEE152BFEFC" box="[1242,1310,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1738,1760]" captionTargetBox="[167,1400,217,1694]" captionTargetId="figure@7.[167,1401,217,1695]" captionTargetPageId="7" captionText="FIGURE 5. Typton fapespae sp. nov., male paratype [MZUSP 31128] and ovigerous female paratype [MZUSP 31130] from São Sebastião, São Paulo, Brazil: A, habitus in life (male), lateral view; B, habitus in life (female), lateral view. Photographs: A. Anker." httpUri="https://zenodo.org/record/230700/files/figure.png" pageId="8" pageNumber="118">Fig. 5</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC19887769D9FECD13C9FE11" box="[199,1020,367,392]" pageId="8" pageNumber="118" type="materials_examined">
<paragraph id="8BCB36E2DC19887769D9FECD13C9FE11" blockId="8.[151,1437,151,2013]" box="[199,1020,367,392]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FECD1150FE11" bold="true" box="[199,357,367,392]" pageId="8" pageNumber="118">
<typeStatus id="54CF8840DC19887769D9FECD1137FE11" box="[199,258,367,392]" pageId="8" pageNumber="118">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3637672DC1988776872FECD1186FE11" box="[364,435,367,392]" name="Brazil" pageId="8" pageNumber="118">Brazil</collectingCountry>
, São Paulo, São Sebastião (Praia do Cabelo Gordo).
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC19887769D9FE3612CCFE6C" pageId="8" pageNumber="118" type="etymology">
<paragraph id="8BCB36E2DC19887769D9FE3612CCFE6C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FE36117BFE34" bold="true" box="[199,334,404,429]" pageId="8" pageNumber="118">Etymology.</emphasis>
The new species is named after Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP) for supporting studies on the origins, diversity and distribution of the flora and fauna of the State of São Paulo, especially through the Biota-FAPESP Program.
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC19887769D9FE5D1143FDA4" pageId="8" pageNumber="118" type="distribution">
<paragraph id="8BCB36E2DC19887769D9FE5D1143FDA4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FE5D1155FD81" bold="true" box="[199,352,511,536]" pageId="8" pageNumber="118">Distribution.</emphasis>
Presently known only from São Sebastião and Ubatuba, northern coast of the state of São Paulo, southeastern
<collectingCountry id="F3637672DC1988776834FD861147FDA4" box="[298,370,548,573]" name="Brazil" pageId="8" pageNumber="118">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC19887769D9FDE510DFFB5C" pageId="8" pageNumber="118" type="biology_ecology">
<paragraph id="8BCB36E2DC19887769D9FDE5123CFC19" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FDE51112FDF9" bold="true" box="[199,295,583,608]" pageId="8" pageNumber="118">Biology.</emphasis>
Like most if not all species of the genus,
<taxonomicName id="4C744D61DC1988776A19FDEB13BDFDF9" box="[775,904,584,608]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776A19FDEB13BDFDF9" box="[775,904,584,608]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776A8EFDE513DEFDF9" bold="true" box="[912,1003,583,608]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776A8EFDE513DEFDF9" box="[912,1003,583,608]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears to be an obligate associate of demosponges. All São Sebastião specimens collected by us were extracted from an unidentified sponge, whereas the male
<typeStatus id="54CF8840DC1988776818FD321153FD31" box="[262,358,656,680]" pageId="8" pageNumber="118" type="paratype">paratype</typeStatus>
from Ubatuba was found inside
<taxonomicName id="4C744D61DC1988776BF1FD2D146CFD31" box="[751,1113,655,680]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="118" phylum="Porifera" rank="species" species="angulosa" subGenus="Zygomycale">
<emphasis id="B900EAF0DC1988776BF1FD2D146CFD31" box="[751,1113,655,680]" italics="true" pageId="8" pageNumber="118">Mycale (Zygomycale) angulosa</emphasis>
</taxonomicName>
. The female
<typeStatus id="54CF8840DC1988776DE4FD32156EFD31" box="[1274,1371,656,680]" pageId="8" pageNumber="118" type="paratype">paratype</typeStatus>
from Ubatuba was extracted from a colony of the bryozoan
<taxonomicName id="4C744D61DC1988776A0CFD1613CEFD55" box="[786,1019,692,717]" class="Gymnolaemata" family="Schizoporellidae" genus="Schizoporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="118" phylum="Bryozoa" rank="species" species="errata">
<emphasis id="B900EAF0DC1988776A0CFD1613CEFD55" box="[786,1019,692,717]" italics="true" pageId="8" pageNumber="118">Schizoporella errata</emphasis>
</taxonomicName>
, most likely having escaped from a cryptic sponge growing inside or attached to the bryozoan colony. The specimens collected from the tangled matrix of tubes of the chaetopterid polychaete
<taxonomicName id="4C744D61DC1988776B51FD5E13B0FC8C" box="[591,901,764,789]" class="Polychaeta" family="Chaetopteridae" genus="Phyllochaetopterus" kingdom="Animalia" pageId="8" pageNumber="118" phylum="Annelida" rank="species" species="socialis">
<emphasis id="B900EAF0DC1988776B51FD5E13B0FC8C" box="[591,901,764,789]" italics="true" pageId="8" pageNumber="118">Phyllochaetopterus socialis</emphasis>
</taxonomicName>
and reported as
<taxonomicName id="4C744D61DC1988776D23FD5F152FFC8C" box="[1085,1306,764,789]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="gnathophylloides">
<emphasis id="B900EAF0DC1988776D23FD5F152FFC8C" box="[1085,1306,764,789]" italics="true" pageId="8" pageNumber="118">T. gnathophylloides</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE54B13DC1988776C5DFD5E1115FCA1" pageId="8" pageNumber="118" refString="Nalesso, R. C., Duarte, L. F. L., Pierozzi, I. Jr. &amp; Enumo, E. F. (1995) Tube epifauna of the polychaete Phyllochaetopterus socialis Claparede. Estuarine, Coastal and Shelf Science, 41, 91 - 100. http: // dx. doi. org / 10.1006 / ecss. 1995.0055" type="journal article">
Nalesso
<emphasis id="B900EAF0DC1988776989FC8310FAFCA1" box="[151,207,799,824]" italics="true" pageId="8" pageNumber="118">et al.</emphasis>
(1995)
</bibRefCitation>
were likely associated to cryptic sponges. Additional material reported by
<bibRefCitation id="EFE54B13DC1988776D7BFC82154BFCA1" author="Duarte" box="[1125,1406,799,824]" pageId="8" pageNumber="118" refString="Duarte, L. F. L. &amp; Nalesso, R. C. (1996) The Sponge Zygomycale parishii (Bowerbank) and its endobiotic fauna. Estuarine, Coastal and Shelf Science, 42, 139 - 151. http: // dx. doi. org / 10.1006 / ecss. 1996.0011" type="journal article" year="1996">Duarte &amp; Nalesso (1996)</bibRefCitation>
as
<taxonomicName id="4C744D61DC1988776989FCE71141FCC4" box="[151,372,836,861]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="gnathophylloides">
<emphasis id="B900EAF0DC1988776989FCE71141FCC4" box="[151,372,836,861]" italics="true" pageId="8" pageNumber="118">T. gnathophylloides</emphasis>
</taxonomicName>
was also found inside
<taxonomicName id="4C744D61DC1988776B69FCE613ECFCC4" box="[631,985,836,861]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="118" phylum="Porifera" rank="species" species="angulosa" subGenus="Zygomycale">
<emphasis id="B900EAF0DC1988776B69FCE613ECFCC4" box="[631,985,836,861]" italics="true" pageId="8" pageNumber="118">Mycale (Zygomycale) angulosa</emphasis>
</taxonomicName>
, suggesting that this sponge may be the main host of
<taxonomicName id="4C744D61DC1988776834FCCB119CFC19" box="[298,425,872,896]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776834FCCB119CFC19" box="[298,425,872,896]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC19887768AEFCC5123CFC19" bold="true" box="[432,521,871,896]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC19887768AEFCC5123CFC19" box="[432,521,871,896]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BCB36E2DC19887769D9FC2E10DFFB5C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
While photographing the São Sebastião specimens of
<taxonomicName id="4C744D61DC1988776A3EFC2F13AAFC3D" box="[800,927,908,932]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776A3EFC2F13AAFC3D" box="[800,927,908,932]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776ABBFC2E13CBFC3C" bold="true" box="[933,1022,908,933]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776ABBFC2E13CBFC3C" box="[933,1022,908,933]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the second author (AA) noticed that most shrimps in an irritated state produced a fairly strong snapping sound, presumably by rapidly closing the major second pereiopod, in a way similar to that of
<taxonomicName id="4C744D61DC1988776B86FC76143BFC74" authority="Borradaile, 1915" authorityName="Borradaile" authorityYear="1915" box="[664,1038,980,1005]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAF0DC1988776B86FC761373FC74" box="[664,838,980,1005]" italics="true" pageId="8" pageNumber="118">Periclimenaeus</emphasis>
<bibRefCitation id="EFE54B13DC1988776A50FC76143BFC74" author="Borradaile" box="[846,1038,980,1005]" pageId="8" pageNumber="118" refString="Borradaile, L. A. (1915) Notes on Carides. The Annals and Magazine of Natural History, Series 8, 15, 205 - 213." type="journal article" year="1915">Borradaile, 1915</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C744D61DC1988776D04FC7714DDFC74" box="[1050,1256,980,1005]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776D04FC7714DDFC74" box="[1050,1256,980,1005]" italics="true" pageId="8" pageNumber="118">Typton spongicola</emphasis>
</taxonomicName>
, a close eastern Atlantic relative of
<taxonomicName id="4C744D61DC198877686EFC5B11C5FB89" box="[368,496,1016,1040]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC198877686EFC5B11C5FB89" box="[368,496,1016,1040]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC19887768EBFC55127BFB89" bold="true" box="[501,590,1015,1040]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC19887768EBFC55127BFB89" box="[501,590,1015,1040]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see below), is also known to produce snapping sounds (
<bibRefCitation id="EFE54B13DC1988776DD4FC551547FB89" author="Johnston" box="[1226,1394,1015,1040]" pageId="8" pageNumber="118" refString="Johnston, T. F. (1946) Underwater noise caused by snapping shrimp. University of California, Division of War Research at the U. S. Navy Electronics Laboratory. UCDWR U 337, 1 - 76." type="journal article" year="1946">Johnston 1946</bibRefCitation>
). In both species, the pollex of the major chela of P2 is deeply depressed accommodating a large flat bulge of the dactylus (
<figureCitation id="134F2A67DC1988776818FB9D1179FBC1" box="[262,332,1087,1112]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="8" pageNumber="118">Fig. 2</figureCitation>
J; see also
<bibRefCitation id="EFE54B13DC19887768D3FBE21263FBC1" author="Bruce" box="[461,598,1088,1112]" pageId="8" pageNumber="118" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
: fig. 5A, B), a configuration that may be responsible for a mechanically induced snapping sound. Thorough studies determining the exact nature and physical properties of the snapping sound are needed for the snapping species of
<taxonomicName id="4C744D61DC1988776BB0FB2B12CFFB39" box="[686,762,1161,1184]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAF0DC1988776BB0FB2B12CFFB39" box="[686,762,1161,1184]" italics="true" pageId="8" pageNumber="118">Typton</emphasis>
</taxonomicName>
, but also for
<taxonomicName id="4C744D61DC1988776A85FB25147CFB39" box="[923,1097,1159,1184]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAF0DC1988776A85FB25147CFB39" box="[923,1097,1159,1184]" italics="true" pageId="8" pageNumber="118">Periclimenaeus</emphasis>
</taxonomicName>
and several other pontoniine genera.
</paragraph>
</subSubSection>
<subSubSection id="C36E6569DC19887669D9FB6D1152FD31" lastPageId="9" lastPageNumber="119" pageId="8" pageNumber="118" type="discussion">
<paragraph id="8BCB36E2DC19887769D9FB6D1321F9B4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="118">
<emphasis id="B900EAF0DC19887769D9FB6D110EFB71" bold="true" box="[199,315,1231,1256]" pageId="8" pageNumber="118">Remarks.</emphasis>
Only three other species of
<taxonomicName id="4C744D61DC1988776B9EFB7312F9FB71" box="[640,716,1233,1256]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAF0DC1988776B9EFB7312F9FB71" box="[640,716,1233,1256]" italics="true" pageId="8" pageNumber="118">Typton</emphasis>
</taxonomicName>
have a strong median tooth on the posterodorsal margin of the sixth abdominal somite, viz.
<taxonomicName id="4C744D61DC19887768F0FB5712B9FA94" box="[494,652,1268,1293]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B900EAF0DC19887768F0FB5712B9FA94" box="[494,652,1268,1293]" italics="true" pageId="8" pageNumber="118">T. hephaestus</emphasis>
</taxonomicName>
(eastern Pacific),
<taxonomicName id="4C744D61DC1988776A78FB5713DDFA94" box="[870,1000,1268,1293]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1988776A78FB5713DDFA94" box="[870,1000,1268,1293]" italics="true" pageId="8" pageNumber="118">T. holthuisi</emphasis>
</taxonomicName>
(central Atlantic), and
<taxonomicName id="4C744D61DC1988776C1FFB5715A9FA94" box="[1281,1436,1268,1293]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776C1FFB5715A9FA94" box="[1281,1436,1268,1293]" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
(eastern Atlantic).
<taxonomicName id="4C744D61DC198877686FFABB1218FAA9" box="[369,557,1304,1328]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC198877686FFABB1218FAA9" box="[369,557,1304,1328]" italics="true" pageId="8" pageNumber="118">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776B26FAB512A3FAA9" bold="true" box="[568,662,1303,1328]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776B26FAB512A3FAA9" box="[568,662,1303,1328]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the poorly described and illustrated
<taxonomicName id="4C744D61DC1988776C99FABB1121FACC" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B900EAF0DC1988776C99FABB1121FACC" italics="true" pageId="8" pageNumber="118">T. hephaestus</emphasis>
</taxonomicName>
by the presence of several (26) teeth on the distolateral margin of the uropodal exopod, above the terminal tooth adjacent to a stout spine [
<figureCitation id="134F2A67DC1988776B43FAFD1295FAE1" box="[605,672,1375,1400]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="8" pageNumber="118">
Fig.
<date id="FFCA1022DC1988776B8DFAC21295FAE1" box="[659,672,1376,1400]" pageId="8" pageNumber="118">3</date>
</figureCitation>
IL; this margin has no teeth in
<taxonomicName id="4C744D61DC1988776D12FAC31490FAE1" box="[1036,1189,1375,1400]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B900EAF0DC1988776D12FAC31490FAE1" box="[1036,1189,1375,1400]" italics="true" pageId="8" pageNumber="118">T. hephaestus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE54B13DC1988776DFCFAFD15BDFAE1" author="Holthuis" box="[1250,1416,1375,1400]" pageId="8" pageNumber="118" refString="Holthuis, L. B. (1951) A general revision of the Palaemonidae (Crustacea Decapoda Natantia) of the Americas. I. The subfamilies Euryrhynchidae and Pontoniinae. Allan Hancock Foundation Publications of the University of Southern California, Occasional Paper, 11, 1 - 332." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
)]. The hammer-shaped dactylus of the major second pereiopod in
<taxonomicName id="4C744D61DC1988776A42FA2713C6FA04" box="[860,1011,1412,1437]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B900EAF0DC1988776A42FA2713C6FA04" box="[860,1011,1412,1437]" italics="true" pageId="8" pageNumber="118">T. hephaestus</emphasis>
</taxonomicName>
provides another character separating the two species. However, both features (uropodal exopod and major P2) were indirectly inferred from the description of
<taxonomicName id="4C744D61DC1988776822FA6F118BFA7C" box="[316,446,1484,1509]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="vulcanus">
<emphasis id="B900EAF0DC1988776822FA6F118BFA7C" box="[316,446,1484,1509]" italics="true" pageId="8" pageNumber="118">T. vulcanus</emphasis>
</taxonomicName>
, which was described by
<bibRefCitation id="EFE54B13DC1988776BF6FA6E13AEFA7C" author="Holthuis" box="[744,923,1484,1509]" pageId="8" pageNumber="118" refString="Holthuis, L. B. (1951) A general revision of the Palaemonidae (Crustacea Decapoda Natantia) of the Americas. I. The subfamilies Euryrhynchidae and Pontoniinae. Allan Hancock Foundation Publications of the University of Southern California, Occasional Paper, 11, 1 - 332." type="journal article" year="1951">Holthuis (1951)</bibRefCitation>
as &quot;extremely close&quot;, with &quot;the only distinct difference between the two forms [being] the shape of the posterior margin of the sixth abdominal segment&quot;. A redescription of
<taxonomicName id="4C744D61DC1988776850F9B711D3F9B4" box="[334,486,1556,1581]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B900EAF0DC1988776850F9B711D3F9B4" box="[334,486,1556,1581]" italics="true" pageId="8" pageNumber="118">T. hephaestus</emphasis>
</taxonomicName>
is indeed highly desirable.
</paragraph>
<paragraph id="8BCB36E2DC19887669D9F9951229FD81" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,680]" lastPageId="9" lastPageNumber="119" pageId="8" pageNumber="118">
The serrated distolateral margin of the uropodal exopod characteristic of
<taxonomicName id="4C744D61DC1988776AE7F99B144CF9C9" box="[1017,1145,1592,1616]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776AE7F99B144CF9C9" box="[1017,1145,1592,1616]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776D61F99514ECF9C9" bold="true" box="[1151,1241,1591,1616]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776D61F99514ECF9C9" box="[1151,1241,1591,1616]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also present in
<taxonomicName id="4C744D61DC1988776989F9FF1105F9EC" box="[151,304,1628,1653]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776989F9FF1105F9EC" box="[151,304,1628,1653]" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D61DC1988776875F9FF11DEF9EC" box="[363,491,1628,1653]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1988776875F9FF11DEF9EC" box="[363,491,1628,1653]" italics="true" pageId="8" pageNumber="118">T. holthuisi</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C744D61DC1988776B6FF9FF12C6F9ED" box="[625,755,1628,1652]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776B6FF9FF12C6F9ED" box="[625,755,1628,1652]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776BE2F9FE136DF9EC" bold="true" box="[764,856,1628,1653]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776BE2F9FE136DF9EC" box="[764,856,1628,1653]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from these two species by the subquadrate merus of the major second pereiopod bearing a field of small but conspicuous, sharp denticles on the ventral surface (
<figureCitation id="134F2A67DC19887769E7F906110AF925" box="[249,319,1700,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="8" pageNumber="118">Fig. 2</figureCitation>
E, H); the merus is more rectangular and ventrally unarmed in
<taxonomicName id="4C744D61DC1988776D06F9071484F924" box="[1048,1201,1700,1725]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776D06F9071484F924" box="[1048,1201,1700,1725]" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D61DC1988776DF2F9071559F924" box="[1260,1388,1700,1725]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1988776DF2F9071559F924" box="[1260,1388,1700,1725]" italics="true" pageId="8" pageNumber="118">T. holthuisi</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFE54B13DC1988776989F96A1115F979" author="Bruce" box="[151,288,1736,1760]" pageId="8" pageNumber="118" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
;
<bibRefCitation id="EFE54B13DC198877682EF96A11DDF979" author="De" box="[304,488,1736,1760]" pageId="8" pageNumber="118" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC198877682EF96A1164F946" box="[304,337,1736,1759]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="118">De</collectingCountry>
Grave 2010
</bibRefCitation>
). Additional distinguishing features of
<taxonomicName id="4C744D61DC1988776AAFF96B1400F979" box="[945,1077,1736,1760]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776AAFF96B1400F979" box="[945,1077,1736,1760]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776D5EF96514ABF979" bold="true" box="[1088,1182,1735,1760]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776D5EF96514ABF979" box="[1088,1182,1735,1760]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are the more angular pterygostomial region (cf.
<figureCitation id="134F2A67DC19887768DEF94E11CBF89D" box="[448,510,1772,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="8" pageNumber="118">Fig. 1</figureCitation>
C and
<bibRefCitation id="EFE54B13DC1988776B54F94F12E5F89D" author="Bruce" box="[586,720,1772,1796]" pageId="8" pageNumber="118" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
: figs. 1, 2A and
<bibRefCitation id="EFE54B13DC1988776A9BF94F1401F89D" author="De" box="[901,1076,1772,1796]" pageId="8" pageNumber="118" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC1988776A9BF94F1393F89D" box="[901,934,1773,1796]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="118">De</collectingCountry>
Grave 2010
</bibRefCitation>
: fig. 1A); the stouter dactylus of the major second cheliped (about 2.1 times as long as maximal depth in
<taxonomicName id="4C744D61DC1988776AC3F8B3146AF8B1" box="[989,1119,1808,1832]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776AC3F8B3146AF8B1" box="[989,1119,1808,1832]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776D76F8AD14F6F8B1" bold="true" box="[1128,1219,1807,1832]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776D76F8AD14F6F8B1" box="[1128,1219,1807,1832]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. 2.6 times in
<taxonomicName id="4C744D61DC1988776C99F8B31127F8D4" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776C99F8B31127F8D4" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
and 2.4 times in
<taxonomicName id="4C744D61DC19887768FFF8971254F8D4" box="[481,609,1844,1869]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC19887768FFF8971254F8D4" box="[481,609,1844,1869]" italics="true" pageId="8" pageNumber="118">T. holthuisi</emphasis>
</taxonomicName>
); and the more robust third and fourth pereiopods (e.g., with P3 merus about twice as long as wide in
<taxonomicName id="4C744D61DC1988776B1FF8FB12B6F8E9" box="[513,643,1880,1904]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1988776B1FF8FB12B6F8E9" box="[513,643,1880,1904]" italics="true" pageId="8" pageNumber="118">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1988776B92F8F512D2F8E9" bold="true" box="[652,743,1879,1904]" pageId="8" pageNumber="118">
<taxonomicNameLabel id="A233578BDC1988776B92F8F512D2F8E9" box="[652,743,1879,1904]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. four times in
<taxonomicName id="4C744D61DC1988776AA6F8FB1464F8E9" box="[952,1105,1879,1904]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776AA6F8FB1464F8E9" box="[952,1105,1879,1904]" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
and almost three times in
<taxonomicName id="4C744D61DC1988776C99F8FB10CCF80C" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1988776C99F8FB10CCF80C" italics="true" pageId="8" pageNumber="118">T. holthuisi</emphasis>
</taxonomicName>
). The new species differs specifically from
<taxonomicName id="4C744D61DC1988776BEBF8DF13BBF80C" box="[757,910,1916,1941]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="118" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1988776BEBF8DF13BBF80C" box="[757,910,1916,1941]" italics="true" pageId="8" pageNumber="118">T. spongicola</emphasis>
</taxonomicName>
by the lower positioned paraorbital processes (cf.
<figureCitation id="134F2A67DC19887769DAF8021131F821" box="[196,260,1952,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="8" pageNumber="118">Fig. 1</figureCitation>
B, C and
<bibRefCitation id="EFE54B13DC198877686CF80311CCF821" author="Bruce" box="[370,505,1952,1976]" pageId="8" pageNumber="118" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
: fig. 2A); the shape of the antennal scaphocerite (cf.
<figureCitation id="134F2A67DC1988776D51F80214A4F821" box="[1103,1169,1952,1977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="8" pageNumber="118">Fig. 2</figureCitation>
D and
<bibRefCitation id="EFE54B13DC1988776DC3F803155EF821" author="Bruce" box="[1245,1387,1952,1976]" pageId="8" pageNumber="118" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce, 2009</bibRefCitation>
: fig. 3D); and the stronger median tooth on the posterodorsal margin of the sixth abdominal somite, reaching beyond lateral teeth in
<taxonomicName id="4C744D61DC188876685EFF3B11F5FF29" box="[320,448,152,176]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC188876685EFF3B11F5FF29" box="[320,448,152,176]" italics="true" pageId="9" pageNumber="119">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC18887668D9FF351214FF29" bold="true" box="[455,545,151,176]" pageId="9" pageNumber="119">
<taxonomicNameLabel id="A233578BDC18887668D9FF351214FF29" box="[455,545,151,176]" pageId="9" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. not reaching beyond them in
<taxonomicName id="4C744D61DC1888766A85FF3B1407FF29" box="[923,1074,151,176]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1888766A85FF3B1407FF29" box="[923,1074,151,176]" italics="true" pageId="9" pageNumber="119">T. spongicola</emphasis>
</taxonomicName>
(cf.
<figureCitation id="134F2A67DC1888766D76FF351493FF29" box="[1128,1190,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="9" pageNumber="119">Fig. 1</figureCitation>
F, G and
<bibRefCitation id="EFE54B13DC1888766C0CFF3A15ADFF29" author="Bruce" box="[1298,1432,152,176]" pageId="9" pageNumber="119" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
: figs. 1, 3J).
<taxonomicName id="4C744D61DC1888766837FF1F11D6FF4D" box="[297,483,188,212]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1888766837FF1F11D6FF4D" box="[297,483,188,212]" italics="true" pageId="9" pageNumber="119">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC18887668F1FF1E1278FF4C" bold="true" box="[495,589,188,213]" pageId="9" pageNumber="119">
<taxonomicNameLabel id="A233578BDC18887668F1FF1E1278FF4C" box="[495,589,188,213]" pageId="9" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs specifically from
<taxonomicName id="4C744D61DC1888766A65FF1F13C8FF4C" box="[891,1021,188,213]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766A65FF1F13C8FF4C" box="[891,1021,188,213]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
by the pollex of the minor second pereiopod ending in a simple tip (vs. with bifid tip in
<taxonomicName id="4C744D61DC1888766BE0FF43134BFF61" box="[766,894,223,248]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766BE0FF43134BFF61" box="[766,894,223,248]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
, cf.
<figureCitation id="134F2A67DC1888766AACFF7D13C3FF61" box="[946,1014,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1803,1825]" captionTargetBox="[166,1429,205,1756]" captionTargetId="figure@3.[157,1442,193,1780]" captionTargetPageId="3" captionText="FIGURE 2. Typton fapespae sp. nov., female paratype [CCDB 3413] (A, B, E, K N) and male paratype [CCDB 4486] (C, D, F J) from Ubatuba, São Paulo, Brazil: A, antennule and antenna, ventral view; B, same, proximal articles, ventral view; C, antenna, proximal portion, ventromesial view; D, same, scaphocerite, dorsal view; E, third maxilliped, lateral view; F, first pereiopod, lateral view; G, major second pereiopod, lateral view; H, same, mesial view; I, same, merus, lateral view; J, same, distal portion of chela, mesial view; K, minor second pereiopod, lateral view; L, same, carpus and chela, mesial view; M, same, fingers, mesial view; N, same, lateral view. Scale bars: A, F, J, K, L = 0.5 mm; B, C, E, I, M, N = 0.25 mm; D = 0.1 mm; G, H = 1 mm." httpUri="https://zenodo.org/record/230697/files/figure.png" pageId="9" pageNumber="119">Fig. 2</figureCitation>
J, L and
<bibRefCitation id="EFE54B13DC1888766D44FF421539FF61" author="De" box="[1114,1292,224,248]" pageId="9" pageNumber="119" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC1888766D44FF42144EFF6E" box="[1114,1147,224,247]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="119">De</collectingCountry>
Grave 2010
</bibRefCitation>
: fig. 2J); the dactyli of the ambulatory pereiopods (P35) furnished with much smaller denticles on the corpus and with an unarmed unguis (vs. with much larger denticles on the corpus and some denticles also on the unguis in
<taxonomicName id="4C744D61DC1888766C09FE8B15A0FED9" box="[1303,1429,295,320]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766C09FE8B15A0FED9" box="[1303,1429,295,320]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
, cf.
<figureCitation id="134F2A67DC18887669A4FEEE10CFFEFD" box="[186,250,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1430,1452]" captionTargetBox="[169,1412,214,1386]" captionTargetId="figure@4.[163,1424,193,1408]" captionTargetPageId="4" captionText="FIGURE 3. Typton fapespae sp. nov., male paratype [CCDB 4486] (A J) and female paratype [CCDB 3413] (K, L) from Ubatuba, São Paulo, Brazil: A, third pereiopod, lateral view; B, fourth pereiopod, lateral view; C, fifth pereiopod, lateral view; D, third pereiopod, dactylus and propodus, mesial view; E, fourth pereiopod, dactylus and propodus, lateral view; F, fifth pereiopod, dactylus and propodus, lateral view; G, first pleopod, ventral view; H, second pleopod, ventral view; I, left uropodal exopod (male), distolateral angle, dorsal view; J, right uropodal exopod (male), distolateral angle, dorsal view; K, left uropodal exopod (female), distolateral angle, dorsal view; L, right uropodal exopod (female), distolateral angle, dorsal view. Scale bars: A C = 0.5 mm, D H = 0.25 mm, I L = 0.1 mm." httpUri="https://zenodo.org/record/230698/files/figure.png" pageId="9" pageNumber="119">Fig. 3</figureCitation>
D, E and
<bibRefCitation id="EFE54B13DC188876687AFEEF1221FEFD" author="De" box="[356,532,332,356]" pageId="9" pageNumber="119" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC188876687AFEEF11B0FEFD" box="[356,389,333,356]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="119">De</collectingCountry>
Grave 2010
</bibRefCitation>
: fig. 3D, F); the shorter dorsolateral spines of the telson (about 0.10 telson length in
<taxonomicName id="4C744D61DC18887669AAFED31100FE11" box="[180,309,368,392]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC18887669AAFED31100FE11" box="[180,309,368,392]" italics="true" pageId="9" pageNumber="119">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1888766822FECD11A2FE11" bold="true" box="[316,407,367,392]" pageId="9" pageNumber="119">
<taxonomicNameLabel id="A233578BDC1888766822FECD11A2FE11" box="[316,407,367,392]" pageId="9" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs.
<quantity id="4C8C9B07DC18887668D9FED21223FE11" box="[455,534,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="119" unit="in" value="0.14">0.14 in</quantity>
<taxonomicName id="4C744D61DC1888766B03FED312A9FE11" box="[541,668,367,392]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766B03FED312A9FE11" box="[541,668,367,392]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
, cf.
<figureCitation id="134F2A67DC1888766BD1FECD133BFE11" box="[719,782,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="9" pageNumber="119">Fig. 1</figureCitation>
H and
<bibRefCitation id="EFE54B13DC1888766A40FED21425FE11" author="De" box="[862,1040,368,392]" pageId="9" pageNumber="119" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC1888766A40FED2134AFE1E" box="[862,895,368,391]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="119">De</collectingCountry>
Grave 2010
</bibRefCitation>
: fig.
<date id="FFCA1022DC1888766D57FED21454FE11" box="[1097,1121,368,392]" pageId="9" pageNumber="119">1I</date>
); and the presence (in most specimens) of a minute median tooth on the posterior margin of the telson (absent in
<taxonomicName id="4C744D61DC1888766D55FE3714FDFE34" box="[1099,1224,404,429]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766D55FE3714FDFE34" box="[1099,1224,404,429]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
, cf.
<figureCitation id="134F2A67DC1888766DE7FE361502FE35" box="[1273,1335,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[218,1332,897,1829]" captionTargetId="figure@2.[208,1343,885,1858]" captionTargetPageId="2" captionText="FIGURE 1. Typton fapespae sp. nov., male paratype [CCDB 4486] (A, B, D J) and female paratype [CCDB 3413] (C) from Ubatuba, São Paulo, Brazil: A, frontal region and cephalic appendages, dorsal view; B, frontal region, lateral view; C, frontal region and proximal part of cephalic appendages, lateral view; D, posterior margin of carapace, lateral view; E, abdomen, lateral view; F, sixth abdominal somite, dorsal view; G, same, lateral view; H, telson, dorsal view; I, same, posterior margin, dorsal view; J, same, spines and setae ommited, dorsal view. Scale bars: A E = 0.5 mm, F H = 0.25 mm, I, J = 0.1 mm." httpUri="https://zenodo.org/record/230696/files/figure.png" pageId="9" pageNumber="119">Fig. 1</figureCitation>
J and
<bibRefCitation id="EFE54B13DC1888766C65FE371116FE49" author="De" pageId="9" pageNumber="119" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC1888766C65FE3715A9FE35" box="[1403,1436,405,428]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="119">De</collectingCountry>
Grave 2010
</bibRefCitation>
: fig. 1J). Regarding its size,
<taxonomicName id="4C744D61DC1888766B63FE1B1334FE49" box="[637,769,440,464]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC1888766B63FE1B1334FE49" box="[637,769,440,464]" italics="true" pageId="9" pageNumber="119">T. fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC1888766A12FE15135FFE49" bold="true" box="[780,874,439,464]" pageId="9" pageNumber="119">
<taxonomicNameLabel id="A233578BDC1888766A12FE15135FFE49" box="[780,874,439,464]" pageId="9" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(pocl
<quantity id="4C8C9B07DC1888766AA5FE1A147DFE49" box="[955,1096,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.4" metricValueMin="1.9" pageId="9" pageNumber="119" unit="mm" value="2.65" valueMax="3.4" valueMin="1.9">1.93.4 mm</quantity>
) is intermediate between the somewhat smaller
<taxonomicName id="4C744D61DC1888766876FE7F11D0FE6C" box="[360,485,476,501]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B900EAF0DC1888766876FE7F11D0FE6C" box="[360,485,476,501]" italics="true" pageId="9" pageNumber="119">T. holthuisi</emphasis>
</taxonomicName>
(pocl
<quantity id="4C8C9B07DC1888766B35FE7E1286FE6D" box="[555,691,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="9" pageNumber="119" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
,
<bibRefCitation id="EFE54B13DC1888766BA0FE7F135FFE6D" author="De" box="[702,874,476,500]" pageId="9" pageNumber="119" refString="De Grave, S. (2010) A new species of the genus Typton Costa (Decapoda, Palaemonidae, Pontoniinae) from Ascension Island. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 209 - 218. [Brill, Leiden]" type="journal article" year="2010">
<collectingCountry id="F3637672DC1888766BA0FE7F12EAFE6D" box="[702,735,477,500]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="119">De</collectingCountry>
Grave 2010
</bibRefCitation>
) and much larger
<taxonomicName id="4C744D61DC1888766D2DFE7F14FCFE6C" box="[1075,1225,476,501]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="spongicola">
<emphasis id="B900EAF0DC1888766D2DFE7F14FCFE6C" box="[1075,1225,476,501]" italics="true" pageId="9" pageNumber="119">T. spongicola</emphasis>
</taxonomicName>
(pocl of ovigerous females
<quantity id="4C8C9B07DC18887669EBFE5D114BFD8E" box="[245,382,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="9" pageNumber="119" unit="mm" value="7.0" valueMax="7.5" valueMin="6.5">6.57.5 mm</quantity>
,
<bibRefCitation id="EFE54B13DC1888766897FDA21225FD81" author="Bruce" box="[393,528,512,536]" pageId="9" pageNumber="119" refString="Bruce, A. J. (2009) A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine, 50, 382 - 394." type="journal article" year="2009">Bruce 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8BCB36E2DC18887669D9FD871152FD31" blockId="9.[151,1436,151,680]" pageId="9" pageNumber="119">
<taxonomicName id="4C744D61DC18887669D9FD8711B6FDA5" box="[199,387,548,572]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="fapespae" status="sp. nov.">
<emphasis id="B900EAF0DC18887669D9FD8711B6FDA5" box="[199,387,548,572]" italics="true" pageId="9" pageNumber="119">Typton fapespae</emphasis>
</taxonomicName>
<emphasis id="B900EAF0DC188876688EFD8611C5FDA4" bold="true" box="[400,496,548,573]" pageId="9" pageNumber="119">
<taxonomicNameLabel id="A233578BDC188876688EFD8611C5FDA4" box="[400,496,548,573]" pageId="9" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from all other western Atlantic species of
<taxonomicName id="4C744D61DC1888766C57FD8715A6FDA5" box="[1353,1427,549,572]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAF0DC1888766C57FD8715A6FDA5" box="[1353,1427,549,572]" italics="true" pageId="9" pageNumber="119">Typton</emphasis>
</taxonomicName>
, including all species previously reported from Brazilian waters (
<taxonomicName id="4C744D61DC1888766A67FDEB13C5FDF9" box="[889,1008,585,608]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="carneus">
<emphasis id="B900EAF0DC1888766A67FDEB13C5FDF9" box="[889,1008,585,608]" italics="true" pageId="9" pageNumber="119">T. carneus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D61DC1888766D1EFDEB14BDFDF9" box="[1024,1160,583,608]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B900EAF0DC1888766D1EFDEB14BDFDF9" box="[1024,1160,583,608]" italics="true" pageId="9" pageNumber="119">T. distinctus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D61DC1888766D87FDEB1542FDF9" box="[1177,1399,583,608]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="gnathophylloides">
<emphasis id="B900EAF0DC1888766D87FDEB1542FDF9" box="[1177,1399,583,608]" italics="true" pageId="9" pageNumber="119">T. gnathophylloides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D61DC1888766C99FDEB1130FD1D" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="prionurus">
<emphasis id="B900EAF0DC1888766C99FDEB1130FD1D" italics="true" pageId="9" pageNumber="119">T. prionurus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D61DC1888766804FDCF11AFFD1D" box="[282,410,621,644]" class="Malacostraca" family="Palaemonidae" genus="Typton" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B900EAF0DC1888766804FDCF11AFFD1D" box="[282,410,621,644]" italics="true" pageId="9" pageNumber="119">T. tortugae</emphasis>
</taxonomicName>
), by the presence of a strong median tooth on the posterodorsal margin of the sixth abdominal somite.
</paragraph>
</subSubSection>
</treatment>
</document>