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<document id="1FB03504223F5B1302391AD36EA3555F" ID-ISSN="1536-9307" ID-ZooBank="07C4C2D6-E904-41DF-9B26-B7C80C689CDC" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638264206251" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2019" docId="03DD87EBFFCBFFEF4685F9DA325FF9A8" docLanguage="en" docName="Euscorpius.294.1-16.pdf" docOrigin="Euscorpius 294" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Neobuthus solegladi Kovařík 2019, sp. n." docType="treatment" docUuid="846B77B6-C223-4C99-AECB-5A23A8EDF8D9" docUuidSource="ZooBank" docVersion="3" lastPageNumber="15" masterDocId="FFE4FF93FFC3FFFF455CFFD33374FFEA" masterDocTitle="Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXII. Two new species of Neobuthus from Somaliland (Buthidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1699436607717" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9F80D6ED479A5A077CD6279BC8B0ED8D">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXII. Two new species of Neobuthus from Somaliland (Buthidae)</mods:title>
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<mods:date id="6BED71DA5F79493238A290F067A391F7">2019</mods:date>
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<heading id="D0838191FFCBFFF74685F9DA366DF9CE" box="[985,1305,1545,1572]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="7" reason="2">
<taxonomicName id="4C744D7EFFCBFFF74685F9DA37A4F9CE" ID-CoL="6S8GK" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[985,1232,1545,1572]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">Neobuthus solegladi</taxonomicName>
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<emphasis id="B900EAEFFFCBFFF7418BF9D93787F9CE" bold="true" box="[1239,1267,1546,1572]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF7405EF9D93666F9CE" bold="true" box="[1282,1298,1546,1572]" pageId="8" pageNumber="7">n</emphasis>
.
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<subSubSection id="C36E6576FFCBFFF746AAF9F83633F969" pageId="8" pageNumber="7" type="description">
<paragraph id="8BCB36FDFFCBFFF746AAF9F836CCF989" blockId="8.[827,1464,1545,1667]" pageId="8" pageNumber="7">
(
<figureCitation id="134F2A78FFCBFFF746A1F9F837EFF9A9" box="[1021,1179,1579,1603]" captionStart-0="Figures 27–28" captionStart-1="Figures 29–32" captionStart-2="Figures 33–40" captionStart-3="Figures 41–59" captionStart-4="Figures 60–68" captionStart-5="Figure 69" captionStart-6="Figure 70" captionStartId-0="9.[113,190,2015,2037]" captionStartId-1="10.[114,191,1988,2010]" captionStartId-2="11.[113,190,1958,1980]" captionStartId-3="12.[115,192,1901,1923]" captionStartId-4="13.[113,188,1987,2009]" captionStartId-5="14.[115,183,1138,1160]" captionStartId-6="15.[112,180,1521,1543]" captionTargetBox-0="[115,1476,232,2003]" captionTargetBox-1="[121,1470,240,1960]" captionTargetBox-2="[136,1466,262,1924]" captionTargetBox-3="[133,1476,240,1874]" captionTargetBox-4="[114,1475,240,1951]" captionTargetBox-5="[113,1471,251,1126]" captionTargetBox-6="[117,1473,239,1511]" captionTargetId-0="figure-29@9.[115,1476,230,2006]" captionTargetId-1="figure-53@10.[113,1474,234,1965]" captionTargetId-2="figure-73@11.[113,1474,234,1930]" captionTargetId-3="figure-112@12.[115,1476,234,1880]" captionTargetId-4="figure-55@13.[114,1475,240,1956]" captionTargetId-5="figure-475@14.[113,1471,231,1126]" captionTargetId-6="figure-276@15.[113,1476,234,1515]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="Figures 27–28. Neobuthus solegladi sp. n., holotype male (27) and paratype female (28) in vivo habitus." captionText-1="Figures 29–32: Neobuthus solegladi sp. n. Figures 29–30. Male holotype, dorsal (29) and ventral (30) views.Figures 31–32. Female paratype, dorsal (31) and ventral (32) views. Scale bar: 10 mm." captionText-2="Figures 33–40: Neobuthus solegladi sp. n. Figures. 33, 35–37. Male holotype, telson lateral (33), metasoma and telson lateral (35), dorsal (36), and ventral (37) views. Figures. 34, 38–40. Female paratype, telson lateral (34), metasoma and telson lateral (38), dorsal (39), and ventral (40) views. Scale bars: 10 mm (35–37, 38–40)." captionText-3="Figures 41–59: Neobuthus solegladi sp. n., pedipalp. Figures 41–49: Male holotype, chela dorsal (41), external (42), and ventral (43) views, patella dorsal (44), external (45) and ventral (46) views, femur and trochanter internal (47), dorsal (48), and ventral (49) views. Figures 50–59. Female paratype, chela dorsal (50), external (51), and ventral (52) views, patella dorsal (53), external (54) and ventral (55) views, femur and trochanter ventral (56) and dorsal (57) views, movable (58) and fixed (59) finger dentate margin. The trichobothrial pattern is indicated in Figures 42–48 (white circles)." captionText-4="Figures 60–68: Neobuthus solegladi sp. n. Figures 60, 62: Male holotype, carapace and tergites (60), coxosternal area and sternites (62). Figures 61, 63–68: Female paratype, carapace and tergites (61), coxosternal area and sternites (63), and right legs I–IV, retrolateral aspect (64–68)." captionText-5="Figure 69. Neobuthus solegladi sp. n., type locality." captionText-6="Figure 70: Map showing distribution of Neobuthus spp." pageId="8" pageNumber="7">Figures 27–70</figureCitation>
,
<tableCitation id="C6F60346FFCBFFF741F9F9F83786F9A9" box="[1189,1266,1579,1603]" captionStart="Table 1" captionStartId="8.[113,169,953,974]" captionText="Table 1. Comparative measurements of adults of Neobuthus haeckeli sp. n. and N. solegladi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="8" pageNumber="7">Table 1</tableCitation>
)
<emphasis id="B900EAEFFFCBFFF74667F99836CCF989" bold="true" box="[827,1464,1611,1635]" pageId="8" pageNumber="7">http://zoobank.org/urn:lsid:zoobank.org:act:846B77B6-</emphasis>
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<emphasis id="B900EAEFFFCBFFF746F0F9B83633F969" bold="true" box="[940,1351,1643,1667]" pageId="8" pageNumber="7">C223-4C99-AECB-5A23A8EDF8D9</emphasis>
</paragraph>
</subSubSection>
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<paragraph id="8BCB36FDFFCBFFF7466CF97F3637F909" blockId="8.[816,1474,1707,1763]" pageId="8" pageNumber="7">
<materialsCitation id="3B1C3CA0FFCBFFF7466CF97F3764F92E" box="[816,1040,1707,1732]" collectionCode="LOCALITY, AND" pageId="8" pageNumber="7" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF885FFFCBFFF7466CF97F3012F929" box="[816,870,1708,1731]" pageId="8" pageNumber="7">TYPE</typeStatus>
<collectionCode id="ED65AE38FFCBFFF7462EF9783764F92E" box="[882,1040,1707,1732]" pageId="8" pageNumber="7">LOCALITY AND</collectionCode>
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<materialsCitation id="3B1C3CA0FFCBFFF74140F97F379BF909" collectionCode="REPOSITORY" elevation="982" latitude="9.884666" longLatPrecision="1" longitude="43.96136" pageId="8" pageNumber="7" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF885FFFCBFFF74140F97F373AF929" box="[1052,1102,1708,1731]" pageId="8" pageNumber="7">TYPE</typeStatus>
<collectionCode id="ED65AE38FFCBFFF74106F97837A2F929" box="[1114,1238,1707,1731]" pageId="8" pageNumber="7">REPOSITORY</collectionCode>
. Somaliland, Agabar,
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,
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a. s. l.
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;
<collectionCode id="ED65AE38FFCBFFF741A0F918364BF909" box="[1276,1343,1739,1763]" pageId="8" pageNumber="7">FKCP</collectionCode>
.
</paragraph>
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<materialsCitation id="3B1C3CA0FFCBFFF7466CF8DF36F5F889" collectingDate="2019-07-09" collectionCode="TYPE, MATERIAL" collectorName="Locality No. &amp; F. Kovarik" elevation="982" latitude="9.884666" longLatPrecision="1" longitude="43.96136" pageId="8" pageNumber="7" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="ED65AE38FFCBFFF7466CF8DF3084F8CE" box="[816,1008,1804,1828]" pageId="8" pageNumber="7">TYPE MATERIAL</collectionCode>
.
<emphasis id="B900EAEFFFCBFFF7414FF8D837E0F8C9" bold="true" box="[1043,1172,1803,1827]" pageId="8" pageNumber="7">Somaliland</emphasis>
, Agabar,
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,
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a. s. l.
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(
<collectorName id="2681532BFFCBFFF7413FF8F83798F8A9" box="[1123,1260,1835,1859]" pageId="8" pageNumber="7">Locality No.</collectorName>
<emphasis id="B900EAEFFFCBFFF741AEF8F83644F8A9" bold="true" box="[1266,1328,1835,1859]" pageId="8" pageNumber="7">19SO</emphasis>
),
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<collectingDate id="EF8EE9D5FFCBFFF7401AF8F836CAF8A9" box="[1350,1470,1835,1859]" pageId="8" pageNumber="7" value="2019-07-09">9.VII.2019</collectingDate>
</date>
,
<specimenCount id="9D72FD74FFCBFFF7466CF8983026F889" box="[816,850,1867,1891]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
(
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, 1717)
<specimenCount id="9D72FD74FFCBFFF7417CF8983735F889" box="[1056,1089,1867,1891]" pageId="8" pageNumber="7" type="female">2♀</specimenCount>
(
<typeStatus id="54CF885FFFCBFFF74109F89837B5F889" box="[1109,1217,1867,1891]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
), leg.
<collectorName id="2681532BFFCBFFF74057F89F36F5F889" box="[1291,1409,1867,1891]" pageId="8" pageNumber="7">F. Kovařík</collectorName>
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&amp;
<materialsCitation id="3B1C3CA0FFCBFFF740F1F89F30FDF869" collectionCode="T" collectorName="Mazuch" pageId="8" pageNumber="7" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED65AE38FFCBFFF740F1F89F36CBF889" box="[1453,1471,1868,1891]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="7">T</collectionCode>
.
<collectorName id="2681532BFFCBFFF7466CF8B830FDF869" box="[816,905,1899,1923]" pageId="8" pageNumber="7">Mazuch</collectorName>
</materialsCitation>
,
<collectionCode id="ED65AE38FFCBFFF746C8F8B830A3F869" box="[916,983,1899,1923]" pageId="8" pageNumber="7">FKCP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFCBFFF1466CF87832E0F8BC" lastPageId="14" lastPageNumber="13" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8BCB36FDFFCBFFF1466CF87832E0F8BC" blockId="8.[816,1474,1963,2019]" lastBlockId="14.[113,772,1182,1878]" lastPageId="14" lastPageNumber="13" pageId="8" pageNumber="7">
ETYMOLOGY. The specific epithet honors Michael E. Soleglad (
<collectingCountry id="F363766DFFCBFFF74664F8183004F809" box="[824,880,1995,2019]" name="United States of America" pageId="8" pageNumber="7">USA</collectingCountry>
) for his friendship and lifelong dedication to scorpions. DIAGNOSIS. Total length 17.7 mm (male), 22–25.7 mm (females); carapace with area between anterior median carinae yellow to orange brown; tergites with 3 dark stripes, median stripe flanked on either side by broad longitudinal yellow bands; pedipalp relatively slender, male with femur L/
<geoCoordinate id="EE40503AFFCDFFF147F2FACD3074FADC" box="[686,768,1310,1334]" degrees="2.36" direction="west" orientation="longitude" pageId="14" pageNumber="13" precision="555" value="-2.36">W 2.36</geoCoordinate>
, patella L/
<geoCoordinate id="EE40503AFFCDFFF14588FAED3250FABC" box="[212,292,1342,1366]" degrees="2.28" direction="west" orientation="longitude" pageId="14" pageNumber="13" precision="555" value="-2.28">W 2.28</geoCoordinate>
, chela L/
<geoCoordinate id="EE40503AFFCDFFF144DFFAED32A0FABC" box="[387,468,1342,1366]" degrees="4.27" direction="west" orientation="longitude" pageId="14" pageNumber="13" precision="555" value="-4.27">W 4.27</geoCoordinate>
; chela movable finger with 5 subrows of primary denticles, 3–4 external accessory denticles flanking proximal end of each subrow; trichobothria
<emphasis id="B900EAEFFFCDFFF147C0FAAD31DDFA7C" box="[668,681,1406,1430]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="17F034B8FFCDFFF147F5FA5D31C5FA76" attach="left" box="[681,689,1422,1436]" fontSize="6" pageId="14" pageNumber="13">2</subScript>
present on dorsal surface of femur and patella; dorsoexternal and ventroexternal carinae on pedipalp patella in female reduced; posterior margins of tergites usually with 3 pairs of macrosetae; pedipalps, legs, metasoma and telson with moderately short, not spiniform macrosetae in males, and long, fine setae in females; males with sternites III–VI with fine granulation, sternite VII finely granulated with 4, granulated carinae; females with sternites III–VI almost smooth, sternite VII finely granulated with 4 weak granulated carinae; metasoma I–III with median lateral and dorsal carinae present in both sexes; lateral surface of metasoma V granulated in both sexes, with granules separated; soles of telotarsi with relatively sparse setation, leg III of adults with 9–13 ventral macrosetae on telotarsus; pectine teeth: 18– 20 (male), 15–16 (females).
</paragraph>
</subSubSection>
<caption id="DF0B6675FFCAFFF6452DF80C3715F81F" box="[113,1121,2015,2037]" pageId="9" pageNumber="8" startId="9.[113,190,2015,2037]" targetBox="[115,1476,232,2003]" targetPageId="9">
<paragraph id="8BCB36FDFFCAFFF6452DF80C3715F81F" blockId="9.[113,1121,2015,2037]" box="[113,1121,2015,2037]" pageId="9" pageNumber="8">
<emphasis id="B900EAEFFFCAFFF6452DF80C3274F81E" bold="true" box="[113,256,2015,2037]" pageId="9" pageNumber="8">Figures 27–28</emphasis>
.
<taxonomicName id="4C744D7EFFCAFFF64450F80C32A4F81E" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[268,464,2015,2036]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCAFFF64450F80C32A4F81E" box="[268,464,2015,2036]" italics="true" pageId="9" pageNumber="8">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCAFFF6448AF80C317FF81E" box="[470,523,2015,2036]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B900EAEFFFCAFFF6448AF80C3299F81E" bold="true" box="[470,493,2015,2036]" pageId="9" pageNumber="8">sp</emphasis>
.
<emphasis id="B900EAEFFFCAFFF644A5F80C3172F81E" bold="true" box="[505,518,2015,2036]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, holotype male (27) and paratype female (28) in vivo habitus.
</paragraph>
</caption>
<caption id="DF0B6675FFC9FFF5452EF8173100F81D" pageId="10" pageNumber="9" startId="10.[114,191,1988,2010]" targetBox="[121,1470,240,1960]" targetPageId="10">
<paragraph id="8BCB36FDFFC9FFF5452EF8173100F81D" blockId="10.[114,1472,1988,2039]" pageId="10" pageNumber="9">
<emphasis id="B900EAEFFFC9FFF5452EF8173274F833" bold="true" box="[114,256,1988,2010]" pageId="10" pageNumber="9">Figures 29–32</emphasis>
:
<taxonomicName id="4C744D7EFFC9FFF54457F81732BAF833" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[267,462,1988,2009]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFC9FFF54457F81732BAF833" box="[267,462,1988,2009]" italics="true" pageId="10" pageNumber="9">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFC9FFF5448FF817317CF830" box="[467,520,1988,2010]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B900EAEFFFC9FFF5448FF817329EF833" bold="true" box="[467,490,1988,2009]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="B900EAEFFFC9FFF544A8F8173175F833" bold="true" box="[500,513,1988,2009]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B900EAEFFFC9FFF54750F81731EEF833" bold="true" box="[524,666,1988,2010]" pageId="10" pageNumber="9">Figures 29–30</emphasis>
. Male holotype, dorsal (29) and ventral (30) views.
<emphasis id="B900EAEFFFC9FFF541DFF8173799F833" bold="true" box="[1155,1261,1988,2010]" pageId="10" pageNumber="9">Figures 31</emphasis>
–3
<emphasis id="B900EAEFFFC9FFF54059F8173665F833" bold="true" box="[1285,1297,1988,2009]" pageId="10" pageNumber="9">2</emphasis>
. Female paratype, dorsal (31) and ventral (32) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="DF0B6675FFC8FFF4452DF8753142F81C" pageId="11" pageNumber="10" startId="11.[113,190,1958,1980]" targetBox="[136,1466,262,1924]" targetPageId="11">
<paragraph id="8BCB36FDFFC8FFF4452DF8753142F81C" blockId="11.[113,1471,1958,2038]" pageId="11" pageNumber="10">
<emphasis id="B900EAEFFFC8FFF4452DF8753275F851" bold="true" box="[113,257,1958,1980]" pageId="11" pageNumber="10">Figures 33–40</emphasis>
:
<taxonomicName id="4C744D7EFFC8FFF44453F87532A1F851" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[271,469,1958,1979]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFC8FFF44453F87532A1F851" box="[271,469,1958,1979]" italics="true" pageId="11" pageNumber="10">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFC8FFF44480F8753160F856" box="[476,532,1958,1980]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B900EAEFFFC8FFF44480F8753287F851" bold="true" box="[476,499,1958,1979]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B900EAEFFFC8FFF4475DF875317AF851" bold="true" box="[513,526,1958,1979]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B900EAEFFFC8FFF44747F875311CF856" bold="true" box="[539,616,1958,1980]" pageId="11" pageNumber="10">Figures</emphasis>
.
<emphasis id="B900EAEFFFC8FFF44729F87531F9F851" bold="true" box="[629,653,1958,1979]" pageId="11" pageNumber="10">33</emphasis>
,
<emphasis id="B900EAEFFFC8FFF447C7F87531A3F851" bold="true" box="[667,727,1958,1980]" pageId="11" pageNumber="10">35–37</emphasis>
. Male holotype, telson lateral (33), metasoma and telson lateral (35), dorsal (36), and ventral (37) views.
<emphasis id="B900EAEFFFC8FFF444DFF81032A4F833" bold="true" box="[387,464,1987,2009]" pageId="11" pageNumber="10">Figures</emphasis>
.
<emphasis id="B900EAEFFFC8FFF44487F8103287F832" bold="true" box="[475,499,1987,2008]" pageId="11" pageNumber="10">34</emphasis>
,
<emphasis id="B900EAEFFFC8FFF444A2F810314EF832" bold="true" box="[510,570,1987,2008]" pageId="11" pageNumber="10">38–40</emphasis>
. Female paratype, telson lateral (34), metasoma and telson lateral (38), dorsal (39), and ventral (40) views. Scale bars: 10 mm (35–37, 38–40).
</paragraph>
</caption>
<caption id="DF0B6675FFCFFFF3452FF8BE32E0F81D" pageId="12" pageNumber="11" startId="12.[115,192,1901,1923]" targetBox="[133,1476,240,1874]" targetPageId="12">
<paragraph id="8BCB36FDFFCFFFF3452FF8BE32E0F81D" blockId="12.[115,1473,1901,2039]" pageId="12" pageNumber="11">
<emphasis id="B900EAEFFFCFFFF3452FF8BE3276F868" bold="true" box="[115,258,1901,1923]" pageId="12" pageNumber="11">Figures 41–59</emphasis>
:
<taxonomicName id="4C744D7EFFCFFFF34452F8BE32A7F868" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[270,467,1901,1922]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCFFFF34452F8BE32A7F868" box="[270,467,1901,1922]" italics="true" pageId="12" pageNumber="11">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCFFFF34485F8BD317AF869" box="[473,526,1902,1923]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B900EAEFFFCFFFF34485F8BD3284F869" bold="true" box="[473,496,1902,1923]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B900EAEFFFCFFFF344A0F8BD317DF869" bold="true" box="[508,521,1902,1923]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, pedipalp.
<emphasis id="B900EAEFFFCFFFF34726F8BE307DF868" bold="true" box="[634,777,1901,1923]" pageId="12" pageNumber="11">Figures 41–49</emphasis>
: Male holotype, chela dorsal (41), external (42), and ventral (43) views, patella dorsal (44), external (45) and ventral (46) views, femur and trochanter internal (47), dorsal (48), and ventral (49) views.
<emphasis id="B900EAEFFFCFFFF34070F85936CEF875" bold="true" box="[1324,1466,1930,1952]" pageId="12" pageNumber="11">Figures 50–59</emphasis>
. Female paratype, chela dorsal (50), external (51), and ventral (52) views, patella dorsal (53), external (54) and ventral (55) views, femur and trochanter ventral (56) and dorsal (57) views, movable (58) and fixed (59) finger dentate margin. The trichobothrial pattern is indicated in Figures 42–48 (white circles).
</paragraph>
</caption>
<caption id="DF0B6675FFCEFFF2452DF81036F7F81C" pageId="13" pageNumber="12" startId="13.[113,188,1987,2009]" targetBox="[114,1475,240,1951]" targetPageId="13">
<paragraph id="8BCB36FDFFCEFFF2452DF81036F7F81C" blockId="13.[113,1471,1987,2038]" pageId="13" pageNumber="12">
<emphasis id="B900EAEFFFCEFFF2452DF8103389F833" bold="true" box="[113,253,1987,2009]" pageId="13" pageNumber="12">Figures 60–68</emphasis>
:
<taxonomicName id="4C744D7EFFCEFFF24454F81732BCF833" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[264,456,1988,2009]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCEFFF24454F81732BCF833" box="[264,456,1988,2009]" italics="true" pageId="13" pageNumber="12">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCEFFF24491F8173176F833" box="[461,514,1988,2009]" pageId="13" pageNumber="12" rank="species">
<emphasis id="B900EAEFFFCEFFF24491F8173297F833" bold="true" box="[461,483,1988,2009]" pageId="13" pageNumber="12">sp</emphasis>
.
<emphasis id="B900EAEFFFCEFFF244B2F817328FF833" bold="true" box="[494,507,1988,2009]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B900EAEFFFCEFFF2475AF81031E5F833" bold="true" box="[518,657,1987,2009]" pageId="13" pageNumber="12">Figures 60, 62</emphasis>
: Male holotype, carapace and tergites (60), coxosternal area and sternites (62).
<emphasis id="B900EAEFFFCEFFF2402FF81033BBF81F" bold="true" pageId="13" pageNumber="12">Figures 61, 63–68</emphasis>
: Female paratype, carapace and tergites (61), coxosternal area and sternites (63), and right legs I–IV, retrolateral aspect (64–68).
</paragraph>
</caption>
<caption id="DF0B6675FFCDFFF1452FFBA1311DFB62" box="[115,617,1138,1160]" pageId="14" pageNumber="13" startId="14.[115,183,1138,1160]" targetBox="[113,1471,251,1126]" targetPageId="14">
<paragraph id="8BCB36FDFFCDFFF1452FFBA1311DFB62" blockId="14.[115,617,1138,1160]" box="[115,617,1138,1160]" pageId="14" pageNumber="13">
<emphasis id="B900EAEFFFCDFFF1452FFBA133A0FB6D" bold="true" box="[115,212,1138,1160]" pageId="14" pageNumber="13">Figure 69</emphasis>
.
<taxonomicName id="4C744D7EFFCDFFF145BCFBA132D1FB6D" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[224,421,1138,1159]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCDFFF145BCFBA132D1FB6D" box="[224,421,1138,1159]" italics="true" pageId="14" pageNumber="13">Neobuthus solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCDFFF144F7FBA13294FB6D" box="[427,480,1138,1159]" pageId="14" pageNumber="13" rank="species">
<emphasis id="B900EAEFFFCDFFF144F7FBA132B6FB6D" bold="true" box="[427,450,1138,1159]" pageId="14" pageNumber="13">sp</emphasis>
.
<emphasis id="B900EAEFFFCDFFF14491FBA132AEFB6D" bold="true" box="[461,474,1138,1159]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, type locality.
</paragraph>
</caption>
<subSubSection id="C36E6576FFCDFFEF452DF8AD310EFA88" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13" type="description">
<paragraph id="8BCB36FDFFCDFFF1452DF8AD30C6F9DC" blockId="14.[113,771,1918,2038]" lastBlockId="14.[816,1475,1182,2038]" pageId="14" pageNumber="13">
DESCRIPTION. Total length of adult male 17.7 mm, of adult females 22–25.7 mm; measurements of carapace, telson, segments of metasoma and pedipalps given in
<tableCitation id="C6F60346FFCDFFF147FFF86D3188F83C" box="[675,764,1982,2006]" captionStart="Table 1" captionStartId="8.[113,169,953,974]" captionText="Table 1. Comparative measurements of adults of Neobuthus haeckeli sp. n. and N. solegladi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="14" pageNumber="13">Table 1</tableCitation>
; positions and distribution of trichobothria of pedipalps shown in
<figureCitation id="134F2A78FFCDFFF14612FB4D30A5FB5C" box="[846,977,1182,1206]" captionStart="Figures 41–59" captionStartId="12.[115,192,1901,1923]" captionTargetBox="[133,1476,240,1874]" captionTargetId="figure-112@12.[115,1476,234,1880]" captionTargetPageId="12" captionText="Figures 41–59: Neobuthus solegladi sp. n., pedipalp. Figures 41–49: Male holotype, chela dorsal (41), external (42), and ventral (43) views, patella dorsal (44), external (45) and ventral (46) views, femur and trochanter internal (47), dorsal (48), and ventral (49) views. Figures 50–59. Female paratype, chela dorsal (50), external (51), and ventral (52) views, patella dorsal (53), external (54) and ventral (55) views, femur and trochanter ventral (56) and dorsal (57) views, movable (58) and fixed (59) finger dentate margin. The trichobothrial pattern is indicated in Figures 42–48 (white circles)." pageId="14" pageNumber="13">Figs. 42–48</figureCitation>
; trichobothrium
<emphasis id="B900EAEFFFCDFFF141DBFB4D37E0FB5C" box="[1159,1172,1182,1206]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="17F034B8FFCDFFF141C8FB7D37E8FB56" attach="left" box="[1172,1180,1198,1212]" fontSize="6" pageId="14" pageNumber="13">2</subScript>
present on pedipalp femur and patella; base color pale yellow to light orange with variable fuscous pigmentation and extensive patterns of dark maculation on mesosoma, metasoma, partially on pedipalps and legs; chelicerae yellow with dark reticulation on anterior manus, dentition reddish.
<emphasis id="B900EAEFFFCDFFF1410FFAED3651FABC" box="[1107,1317,1342,1366]" italics="true" pageId="14" pageNumber="13">Sexual dimorphism</emphasis>
: strong, adult male substantially smaller, but without differences in shapes of pedipalps, metasoma and telson; pedipalp patella and femur granulate and matte in males, smooth and glossy in females; sternites smooth in females and granulated in male; macrosetae on pedipalps, legs, metasoma and telson much longer and finer in females than males; other sex differences cited below.
</paragraph>
<paragraph id="8BCB36FDFFCDFFF1466CF9ED3676F8FC" blockId="14.[816,1475,1182,2038]" pageId="14" pageNumber="13">
<emphasis id="B900EAEFFFCDFFF1466CF9ED30E1F9BC" bold="true" box="[816,917,1598,1622]" pageId="14" pageNumber="13">Pedipalp</emphasis>
(
<figureCitation id="134F2A78FFCDFFF146F6F9ED3745F9BC" box="[938,1073,1598,1622]" captionStart="Figures 41–59" captionStartId="12.[115,192,1901,1923]" captionTargetBox="[133,1476,240,1874]" captionTargetId="figure-112@12.[115,1476,234,1880]" captionTargetPageId="12" captionText="Figures 41–59: Neobuthus solegladi sp. n., pedipalp. Figures 41–49: Male holotype, chela dorsal (41), external (42), and ventral (43) views, patella dorsal (44), external (45) and ventral (46) views, femur and trochanter internal (47), dorsal (48), and ventral (49) views. Figures 50–59. Female paratype, chela dorsal (50), external (51), and ventral (52) views, patella dorsal (53), external (54) and ventral (55) views, femur and trochanter ventral (56) and dorsal (57) views, movable (58) and fixed (59) finger dentate margin. The trichobothrial pattern is indicated in Figures 42–48 (white circles)." pageId="14" pageNumber="13">Figs. 41–59</figureCitation>
). Pedipalp mostly sparsely hirsute; finely granulated in males and smooth in females; femur with five conspicuously granulose carinae, more strongly developed in male; patella with seven granulose carinae, well developed in male and reduced in females; dorsoexternal carinae on pedipalp patella in female reduced to absent; chela with carinae present but smooth in females;
</paragraph>
<paragraph id="8BCB36FDFFCDFFF1466CF8CD3719F89C" blockId="14.[816,1475,1182,2038]" pageId="14" pageNumber="13">chela movable finger with 5 and fixed finger with 4–5 subrows of primary denticles, 3–4 external accessory denticles flanking proximal end of each subrow.</paragraph>
<paragraph id="8BCB36FDFFCDFFF0466CF8AD3115F93A" blockId="14.[816,1475,1182,2038]" lastBlockId="15.[112,772,1521,2032]" lastPageId="15" lastPageNumber="14" pageId="14" pageNumber="13">
<emphasis id="B900EAEFFFCDFFF1466CF8AD30EBF87C" bold="true" box="[816,927,1918,1942]" pageId="14" pageNumber="13">Carapace</emphasis>
(
<figureCitation id="134F2A78FFCDFFF146EBF8AD3732F87C" box="[951,1094,1918,1942]" captionStart="Figures 60–68" captionStartId="13.[113,188,1987,2009]" captionTargetBox="[114,1475,240,1951]" captionTargetId="figure-55@13.[114,1475,240,1956]" captionTargetPageId="13" captionText="Figures 60–68: Neobuthus solegladi sp. n. Figures 60, 62: Male holotype, carapace and tergites (60), coxosternal area and sternites (62). Figures 61, 63–68: Female paratype, carapace and tergites (61), coxosternal area and sternites (63), and right legs I–IV, retrolateral aspect (64–68)." pageId="14" pageNumber="13">Figs. 60–61</figureCitation>
). Strongly trapezoidal (narrower anteriorly), wider than long (L/
<geoCoordinate id="EE40503AFFCDFFF141F3F84C3673F85C" box="[1199,1287,1950,1974]" degrees="0.85" direction="west" orientation="longitude" pageId="14" pageNumber="13" precision="555" value="-0.85">W 0.85</geoCoordinate>
–0.89); posterior median postocular area flat, anterior median preocular area gently sloped downwards towards anterior margin; lateral flanks steeply sloped; ocular tubercle broad, prominent, located slightly anterior to middle of carapace; anterior margin straight, finely microdenticulate, with coarser granules overlapping edge, bearing 8–10 macrosetae; anterior median carinae present, coarsely granular, other carinae indistinct; dense granulation covering most of carapace.
</paragraph>
<caption id="DF0B6675FFCCFFF0452CFA2231FFF9EC" box="[112,651,1521,1543]" pageId="15" pageNumber="14" startId="15.[112,180,1521,1543]" targetBox="[117,1473,239,1511]" targetPageId="15">
<paragraph id="8BCB36FDFFCCFFF0452CFA2231FFF9EC" blockId="15.[112,772,1521,2032]" box="[112,651,1521,1543]" pageId="15" pageNumber="14">
<emphasis id="B900EAEFFFCCFFF0452CFA2233A6F9EC" bold="true" box="[112,210,1521,1543]" pageId="15" pageNumber="14">Figure 70</emphasis>
: Map showing distribution of
<taxonomicName id="4C744D7EFFCCFFF044AAFA2231FFF9EC" box="[502,651,1521,1542]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAEFFFCCFFF044AAFA223129F9EC" box="[502,605,1521,1542]" italics="true" pageId="15" pageNumber="14">Neobuthus</emphasis>
spp.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BCB36FDFFCCFFF0452DF90B3207F83A" blockId="15.[112,772,1521,2032]" pageId="15" pageNumber="14">
<emphasis id="B900EAEFFFCCFFF0452DF90B33ABF91A" bold="true" box="[113,223,1752,1776]" pageId="15" pageNumber="14">Chelicera</emphasis>
. Fingers with typical buthid dentition (
<bibRefCitation id="EFE54B0CFFCCFFF047F7F90B33DDF8FA" author="VACHON" pageId="15" pageNumber="14" pagination="161 - 166" refId="ref6172" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
; Lowe &amp; Kovařík, 2016;
<bibRefCitation id="EFE54B0CFFCCFFF0448DF92B31C3F8FA" author="KOVARIK" box="[465,695,1784,1808]" pageId="15" pageNumber="14" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
); fixed finger with large distal denticle, 1 subdistal denticle and 2 basal denticles fused into bicusp, single denticle on ventral surface at level of bicusp; dorsal margin of movable finger with 5 denticles: 1 large distal denticle, medium-sized subdistal and medial, and 2 small, partially fused basal denticles; ventral margin with 2 denticles.
</paragraph>
<paragraph id="8BCB36FDFFCCFFEF452DF80B324AFDC8" blockId="15.[112,772,1521,2032]" lastBlockId="16.[113,772,235,1027]" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
<emphasis id="B900EAEFFFCCFFF0452DF80B339DF81A" bold="true" box="[113,233,2008,2032]" pageId="15" pageNumber="14">Mesosoma</emphasis>
(
<figureCitation id="134F2A78FFCCFFF045A1F80B32F1F81A" box="[253,389,2008,2032]" captionStart="Figures 60–68" captionStartId="13.[113,188,1987,2009]" captionTargetBox="[114,1475,240,1951]" captionTargetId="figure-55@13.[114,1475,240,1956]" captionTargetPageId="13" captionText="Figures 60–68: Neobuthus solegladi sp. n. Figures 60, 62: Male holotype, carapace and tergites (60), coxosternal area and sternites (62). Figures 61, 63–68: Female paratype, carapace and tergites (61), coxosternal area and sternites (63), and right legs I–IV, retrolateral aspect (64–68)." pageId="15" pageNumber="14">Figs. 60–63</figureCitation>
). Tergites I–VI bear three carinae of which the lateral pair may be less conspicuous mainly on tergites I–IV; tergite VII bears five well-defined carinae (median, submedians and laterals); tergites I–VI densely granular, with coarser granules on posterior lateral areas; tergite VII densely granular; sternites III–VI almost smooth in females, and finely granulated in male; sternite VII granulated in both sexes, more so in males, with four well-defined carinae; sternum
<typeStatus id="54CF885FFFCCFFF046AFF92A3755F8FB" box="[1011,1057,1785,1809]" pageId="15" pageNumber="14">type</typeStatus>
1, triangular in shape, smooth, with deep posteromedian invagination; genital opercula smooth; genital papillae present; pectines extending to around a half of sternite V in male and around a half of sternite IV in female; pectine teeth
<quantity id="4C8C9B18FFCCFFF046E3F8AB3769F87A" box="[959,1053,1912,1936]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="15" pageNumber="14" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
male, 15–16 (1x 15, 3 x16) in females; combs with 3 marginal lamellae and 7–8 middle lamellae; marginal lamellae, middle lamellae and fulcra with dense cover of short dark reddish macrosetae; fulcra with 2–4 setae.
<emphasis id="B900EAEFFFD3FFEF452DFF3833D2FEE9" bold="true" box="[113,166,235,259]" pageId="16" pageNumber="15">Legs</emphasis>
(
<figureCitation id="134F2A78FFD3FFEF45E9FF38324DFEE9" box="[181,313,235,259]" captionStart="Figures 60–68" captionStartId="13.[113,188,1987,2009]" captionTargetBox="[114,1475,240,1951]" captionTargetId="figure-55@13.[114,1475,240,1956]" captionTargetPageId="13" captionText="Figures 60–68: Neobuthus solegladi sp. n. Figures 60, 62: Male holotype, carapace and tergites (60), coxosternal area and sternites (62). Figures 61, 63–68: Female paratype, carapace and tergites (61), coxosternal area and sternites (63), and right legs I–IV, retrolateral aspect (64–68)." pageId="16" pageNumber="15">Figs. 64–68</figureCitation>
). Coxa, femora, patella and tibia of all legs bearing variable numbers of short to medium length, straight, dark-reddish macrosetae; tarsi with mix of short and longer, dark-reddish macrosetae; basitarsi I–III slightly compressed with flat retrolateral surfaces, with bristle combs consisting of retrosuperior series of longer macrosetae, plus retroinferior and proinferior series of shorter macrosetae; telotarsi with two rows of short macrosetae on ventral aspect, 9–13 macrosetae on telotarsus III; moderate tibial spurs present on leg IV and reduced on leg. III.
</paragraph>
<paragraph id="8BCB36FDFFD3FFEF452DFDF8324EFBE9" blockId="16.[113,772,235,1027]" pageId="16" pageNumber="15">
<emphasis id="B900EAEFFFD3FFEF452DFDF8321FFDA9" bold="true" box="[113,363,555,579]" pageId="16" pageNumber="15">Metasoma and telson</emphasis>
(
<figureCitation id="134F2A78FFD3FFEF4421FDF83177FDA9" box="[381,515,555,579]" captionStart="Figures 33–40" captionStartId="11.[113,190,1958,1980]" captionTargetBox="[136,1466,262,1924]" captionTargetId="figure-73@11.[113,1474,234,1930]" captionTargetPageId="11" captionText="Figures 33–40: Neobuthus solegladi sp. n. Figures. 33, 35–37. Male holotype, telson lateral (33), metasoma and telson lateral (35), dorsal (36), and ventral (37) views. Figures. 34, 38–40. Female paratype, telson lateral (34), metasoma and telson lateral (38), dorsal (39), and ventral (40) views. Scale bars: 10 mm (35–37, 38–40)." pageId="16" pageNumber="15">Figs. 33–40</figureCitation>
). Metasoma and telson sparsely hirsute, macrosetae moderately short in male and longer in female, straight and reddish to black; metasomal segments I–III with 10 carinae, IV with 8 carinae, V with 2 carinae; all carinae relatively well developed; segment IV with weakly indicated dorsolateral carinae; segment V with strong, granulate to dentate-lobate ventrolateral carinae; segments I– IV with dense granulation on all intercarinal surfaces except dorsal surfaces which are sparsely granulated, mainly in females; segment V densely granular on lateral and ventral surfaces, more coarsely so on ventral surface, granules not arranged along any traces of carinae; telson smooth, ventral surface sparsely, weakly granular; vesicle slightly elongated; aculeus stout, shorter than vesicle, tip of aculeus almost vertically directed.
</paragraph>
<paragraph id="8BCB36FDFFD3FFEF452DFBF8310EFA88" blockId="16.[113,773,1067,1379]" pageId="16" pageNumber="15">
AFFINITIES. The described features distinguish
<taxonomicName id="4C744D7EFFD3FFEF4721FBF83077FBA9" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[637,771,1067,1091]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFD3FFEF4721FBF83077FBA9" box="[637,771,1067,1091]" italics="true" pageId="16" pageNumber="15">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFD3FFEF452DFB9833C5FB88" box="[113,177,1099,1123]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B900EAEFFFD3FFEF452DFB9833FEFB89" bold="true" box="[113,138,1099,1123]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B900EAEFFFD3FFEF45C7FB9833DEFB89" bold="true" box="[155,170,1099,1123]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. According to the characters used in the key published in
<bibRefCitation id="EFE54B0CFFD3FFEF4747FBB8318BFB69" author="KOVARIK" box="[539,767,1131,1155]" pageId="16" pageNumber="15" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al. (2018)</bibRefCitation>
, the new species is most similar to
<taxonomicName id="4C744D7EFFD3FFEF44B2FB5833A5FB29" authority="Kovarik et al., 2018" authorityName="Kovarik" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="B900EAEFFFD3FFEF44B2FB5831F3FB49" box="[494,647,1163,1187]" italics="true" pageId="16" pageNumber="15">N. gubanensis</emphasis>
<bibRefCitation id="EFE54B0CFFD3FFEF47CEFB5833A5FB29" author="KOVARIK" pageId="16" pageNumber="15" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
</taxonomicName>
, which is also confirmed by DNA phylogeny (paper in preparation). Morphologically, it is difficult to distinguish these two species based on single specimens but we can see that female of
<taxonomicName id="4C744D7EFFD3FFEF4443FAD832D2FAC9" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[287,422,1291,1315]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFD3FFEF4443FAD832D2FAC9" box="[287,422,1291,1315]" italics="true" pageId="16" pageNumber="15">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFD3FFEF44EFFAD83282FAC8" box="[435,502,1291,1315]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B900EAEFFFD3FFEF44EFFAD832B8FAC9" bold="true" box="[435,460,1291,1315]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B900EAEFFFD3FFEF44BCFAD8329BFAC9" bold="true" box="[480,495,1291,1315]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
has smooth carinae on pedipalp chela (
<figureCitation id="134F2A78FFD3FFEF4446FAF8321DFAA8" box="[282,361,1322,1347]" captionStart="Figures 41–59" captionStartId="12.[115,192,1901,1923]" captionTargetBox="[133,1476,240,1874]" captionTargetId="figure-112@12.[115,1476,234,1880]" captionTargetPageId="12" captionText="Figures 41–59: Neobuthus solegladi sp. n., pedipalp. Figures 41–49: Male holotype, chela dorsal (41), external (42), and ventral (43) views, patella dorsal (44), external (45) and ventral (46) views, femur and trochanter internal (47), dorsal (48), and ventral (49) views. Figures 50–59. Female paratype, chela dorsal (50), external (51), and ventral (52) views, patella dorsal (53), external (54) and ventral (55) views, femur and trochanter ventral (56) and dorsal (57) views, movable (58) and fixed (59) finger dentate margin. The trichobothrial pattern is indicated in Figures 42–48 (white circles)." pageId="16" pageNumber="15">Fig. 50</figureCitation>
), which are absent in the females of
<taxonomicName id="4C744D7EFFD3FFEF47B7FAF8339DFA89" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="B900EAEFFFD3FFEF47B7FAF8339DFA89" italics="true" pageId="16" pageNumber="15">N. gubanensis</emphasis>
</taxonomicName>
(fig.
<quantity id="4C8C9B18FFD3FFEF447EFA983212FA89" box="[290,358,1355,1379]" metricMagnitude="0" metricUnit="m" metricValue="6.2484" pageId="16" pageNumber="15" unit="in" value="246.0">246 in</quantity>
<bibRefCitation id="EFE54B0CFFD3FFEF4431FA983119FA88" author="KOVARIK" box="[365,621,1354,1379]" pageId="16" pageNumber="15" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018: 50</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFD3FFEF452DFA58325FF9A8" pageId="16" pageNumber="15" type="discussion">
<paragraph id="8BCB36FDFFD3FFEF452DFA58325FF9A8" blockId="16.[113,772,1419,1602]" pageId="16" pageNumber="15">
COMMENTS ON LOCALITIES AND LIFE STRATEGY.
<taxonomicName id="4C744D7EFFD3FFEF4701FA5831A9FA49" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[605,733,1419,1443]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFD3FFEF4701FA5831A9FA49" box="[605,733,1419,1443]" italics="true" pageId="16" pageNumber="15">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFD3FFEF47BEFA5833F3FA28" pageId="16" pageNumber="15" rank="species">
<emphasis id="B900EAEFFFD3FFEF47BEFA58318FFA49" bold="true" box="[738,763,1419,1443]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B900EAEFFFD3FFEF452DFA7833F4FA29" bold="true" box="[113,128,1451,1475]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
inhabits rocky mountain area in central Somaliland. The
<typeStatus id="54CF885FFFD3FFEF452DFA1833DDFA09" box="[113,169,1483,1507]" pageId="16" pageNumber="15">types</typeStatus>
were collected at night in open terrain by UV detection together with
<taxonomicName id="4C744D7EFFD3FFEF4454FA383074F9E9" authority="Kovarik et al., 2019" authorityName="Kovarik" authorityYear="2019" box="[264,768,1515,1539]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B900EAEFFFD3FFEF4454FA38316FF9E9" box="[264,539,1515,1539]" italics="true" pageId="16" pageNumber="15">Parabuthus somalilandus</emphasis>
Kovařík et al., 2019
</taxonomicName>
,
<taxonomicName id="4C744D7EFFD3FFEF452DF9D8316DF9C9" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[113,537,1547,1571]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="polystictus">
<emphasis id="B900EAEFFFD3FFEF452DF9D83213F9C9" box="[113,359,1547,1571]" italics="true" pageId="16" pageNumber="15">Hottentotta polystictus</emphasis>
(Pocock, 1896)
</taxonomicName>
,
<taxonomicName id="4C744D7EFFD3FFEF477BF9D831BAF9C8" box="[551,718,1547,1571]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAEFFFD3FFEF477BF9D831D7F9C9" box="[551,675,1547,1571]" italics="true" pageId="16" pageNumber="15">Pandinurus</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="4C744D7EFFD3FFEF452DF9F8325FF9A8" box="[113,299,1579,1602]" class="Arachnida" family="Hemiscorpiidae" genus="Hemiscorpius" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAEFFFD3FFEF452DF9F83271F9A8" box="[113,261,1579,1602]" italics="true" pageId="16" pageNumber="15">Hemiscorpius</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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