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<document id="D9324388E06D25B10C23B4197550619C" ID-DOI="10.5281/zenodo.5348617" ID-ISSN="2345-7600" ID-Zenodo-Dep="5348617" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630204003608" checkinUser="tatiana" docAuthor="Dow, R. A. &amp; Orr, A. G." docDate="2012" docId="03DD87E5FFCF6170FC34FB56FB1A9F95" docLanguage="en" docName="60rbz361-397.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Telosticta janeus Dow &amp; Orr 2012, new species" docType="treatment" docVersion="3" lastPageNumber="395" masterDocId="FFE4FF9DFFEE6152FFF2FFABFFAF9A63" masterDocTitle="Telosticta, A New Damselfly Genus From Borneo And Palawan (Odonata: Zygoptera: Platystictidae)" masterLastPageNumber="397" masterPageNumber="361" pageNumber="394" updateTime="1699184126446" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BADA7DE4B90A41D7DAEB0CA3A42DEFB3">Telosticta, A New Damselfly Genus From Borneo And Palawan (Odonata: Zygoptera: Platystictidae)</mods:title>
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<mods:namePart id="9D5694E98E6DC0C996B08DA771A3D554">Dow, R. A.</mods:namePart>
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<mods:namePart id="60E2CF74596D59608F2220CEB0CD7A33">Orr, A. G.</mods:namePart>
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<mods:typeOfResource id="408447C226FF6CA889718DD22FF1B31D">text</mods:typeOfResource>
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<mods:title id="ECFA80444CAC41B1057CA765426BCD0E">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="AE4C4B82103714FB62A625BD7784D7DA">2012</mods:date>
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<mods:number id="D9DBD1162C90572F5582B3E8D90FC447">2012-08-31</mods:number>
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<treatment id="03DD87E5FFCF6170FC34FB56FB1A9F95" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95" httpUri="http://treatment.plazi.org/id/03DD87E5FFCF6170FC34FB56FB1A9F95" lastPageId="34" lastPageNumber="395" pageId="33" pageNumber="394">
<subSubSection id="C36E6578FFCF6173FC34FB56FABD9F75" box="[966,1298,1277,1302]" pageId="33" pageNumber="394" type="nomenclature">
<paragraph id="8BCB36F3FFCF6173FC34FB56FABD9F75" blockId="33.[926,1337,1277,1334]" box="[966,1298,1277,1302]" pageId="33" pageNumber="394">
<heading id="D083819FFFCF6173FC34FB56FABD9F75" bold="true" box="[966,1298,1277,1302]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="394" reason="2">
<emphasis id="B900EAE1FFCF6173FC34FB56FABD9F75" bold="true" box="[966,1298,1277,1302]" pageId="33" pageNumber="394">
<taxonomicName id="4C744D70FFCF6173FC34FB56FBD09F75" ID-CoL="9X4GV" authority="Dow &amp; Orr, 2012" authorityName="Dow &amp; Orr" authorityYear="2012" box="[966,1151,1277,1302]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="394" phylum="Arthropoda" rank="species" species="janeus" status="sp. nov.">
<emphasis id="B900EAE1FFCF6173FC34FB56FBD09F75" bold="true" box="[966,1151,1277,1302]" italics="true" pageId="33" pageNumber="394">Telosticta janeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A233579AFFCF6173FB7FFB55FABD9F75" box="[1165,1298,1278,1302]" pageId="33" pageNumber="394" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCF6173FC6CFAB5FA969F55" box="[926,1337,1310,1334]" pageId="33" pageNumber="394" type="description">
<paragraph id="8BCB36F3FFCF6173FC6CFAB5FA969F55" blockId="33.[926,1337,1277,1334]" box="[926,1337,1310,1334]" pageId="33" pageNumber="394">
(
<figureCitation id="134F2A76FFCF6173FC54FAB5FBE09F55" box="[934,1103,1310,1334]" captionStart="Fig" captionStartId="19.[114,149,2035,2057]" captionTargetBox="[114,1453,401,1976]" captionTargetId="figure-98@19.[114,1453,401,1976]" captionTargetPageId="19" captionText="Fig. 16. Markings of T. belalongensis and T. janeus. T. belalongensis holotype: A, antehumeral markings; C, terminal abdominal segments, dorsal view; E, terminal abdominal segments, lateral view. T. janeus holotype: B, antehumeral markings; D, terminal abdominal segments, dorsal view; F, terminal abdominal segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5348649" httpUri="https://zenodo.org/record/5348649/files/figure.png" pageId="33" pageNumber="394">Figs. 16B, D, F</figureCitation>
,
<figureCitation id="134F2A76FFCF6173FBAEFAB5FB039F55" box="[1116,1196,1310,1334]" captionStart="Fig" captionStartId="20.[136,171,2062,2084]" captionTargetBox="[140,1471,1495,2006]" captionTargetId="figure-576@20.[123,1491,1485,2021]" captionTargetPageId="20" captionText="Fig. 17. T. belalongensis and T. janeus. T. belalongensis holotype: A, posterior pronotal lobe, lateral; C, right inferior anal appendage, ventral view. T. janeus holotype: B, posterior pronotal lobe, lateral; D, right inferior anal appendage, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5348651" httpUri="https://zenodo.org/record/5348651/files/figure.png" pageId="33" pageNumber="394">17B, D</figureCitation>
,
<figureCitation id="134F2A76FFCF6173FB4BFAB5FAA69F55" box="[1209,1289,1310,1334]" captionStart="Fig" captionStartId="21.[112,147,2063,2085]" captionTargetBox="[113,1452,1051,2011]" captionTargetId="figure-431@21.[101,1469,1019,2020]" captionTargetPageId="21" captionText="Fig. 18. T. belalongensis and T. janeus. T. belalongensis holotype:A, anal appendages dorsal; C, anal appendages lateral. T. janeus holotype: B, anal appendages dorsal; D, anal appendages lateral." figureDoi="http://doi.org/10.5281/zenodo.5348653" httpUri="https://zenodo.org/record/5348653/files/figure.png" pageId="33" pageNumber="394">18B, D</figureCitation>
,
<figureCitation id="134F2A76FFCF6173FAE4FAB5FA9E9F55" box="[1302,1329,1310,1334]" captionStart="Fig" captionStartId="22.[835,870,2063,2085]" captionTargetBox="[835,1473,1434,2023]" captionTargetId="figure-749@22.[835,1472,1434,2023]" captionTargetPageId="22" captionText="Fig. 19. Distribution of T. bidayuh (cross), T. janeus (open square), T. feronia (black square), and T. longigaster (circle)." figureDoi="http://doi.org/10.5281/zenodo.5348655" httpUri="https://zenodo.org/record/5348655/files/figure.png" pageId="33" pageNumber="394">19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCF6173FCDEFAF7FBFB9C3B" pageId="33" pageNumber="394" type="materials_examined">
<paragraph id="8BCB36F3FFCF6173FCDEFAF7FA279FEC" blockId="33.[812,1451,1372,1424]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEFAF7FC5B9F11" bold="true" box="[812,1012,1372,1394]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEFAF7FC429F11" bold="true" box="[812,1005,1372,1394]" italics="true" pageId="33" pageNumber="394">Material examined</emphasis>
.
</emphasis>
<materialsCitation id="3B1C3CAEFFCF6173FBECFAF6FA279FEC" collectingDate="1987-03-19" collectionCode="RMNH" collectorName="J. v. Tol" country="Malaysia" location="Danum Valley" pageId="33" pageNumber="394" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54CF8851FFCF6173FBECFAF6FBD29F10" box="[1054,1149,1373,1395]" pageId="33" pageNumber="394" type="holotype">Holotype</typeStatus>
: male (JvT 19977),
<collectingCountry id="F3637663FFCF6173FABEFAF6FA079F10" box="[1356,1448,1373,1395]" name="Malaysia" pageId="33" pageNumber="394">Malaysia</collectingCountry>
,
<collectingRegion id="49B0F811FFCF6173FCDEFAD1FCC69FF3" box="[812,873,1402,1424]" country="Malaysia" name="Sabah" pageId="33" pageNumber="394">Sabah</collectingRegion>
,
<location id="8EAB6028FFCF6173FC86FAD1FBAF9FF3" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCF6173FC86FAD1FBAF9FF3" box="[884,1024,1402,1424]" country="Malaysia" name="Danum Valley" pageId="33" pageNumber="394" stateProvince="Sabah">Danum Valley</location>
,
<date id="FFCA1033FFCF6173FBF9FAD1FB269FEC" box="[1035,1161,1402,1423]" pageId="33" pageNumber="394" value="1987-03-19">
<collectingDate id="EF8EE9DBFFCF6173FBF9FAD1FB269FEC" box="[1035,1161,1402,1423]" pageId="33" pageNumber="394" value="1987-03-19">19 Mar.1987</collectingDate>
</date>
, coll.
<collectorName id="26815325FFCF6173FB37FAD1FABA9FF3" box="[1221,1301,1402,1424]" pageId="33" pageNumber="394">J. v. Tol</collectorName>
, in
<collectionCode id="ED65AE36FFCF6173FACBFAD1FA2B9FEC" box="[1337,1412,1402,1423]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="33" pageNumber="394" type="Museum">RMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCB36F3FFCF6173FCDEFA1FFBFB9C3B" blockId="33.[812,1451,1459,1625]" pageId="33" pageNumber="394">
<materialsCitation id="3B1C3CAEFFCF6173FCDEFA1FFC5B9F86" collectingDate="1987-03-19" collectionCode="RMNH" collectorName="J. v. Tol" country="Malaysia" location="Danum Valley" pageId="33" pageNumber="394" specimenCount="4" specimenCount-male="4" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54CF8851FFCF6173FCDEFA1FFC269FAA" box="[812,905,1460,1481]" pageId="33" pageNumber="394" type="paratype">Paratypes</typeStatus>
(all from
<collectingCountry id="F3637663FFCF6173FC1BFA18FBEC9FAA" box="[1001,1091,1459,1481]" name="Malaysia" pageId="33" pageNumber="394">Malaysia</collectingCountry>
,
<collectingRegion id="49B0F811FFCF6173FBBEFA18FB249FAA" box="[1100,1163,1459,1481]" country="Malaysia" name="Sabah" pageId="33" pageNumber="394">Sabah</collectingRegion>
):
<specimenCount id="9D72FD7AFFCF6173FB69FA18FB4A9FAA" box="[1179,1253,1459,1481]" pageId="33" pageNumber="394" type="male">4 males</specimenCount>
(JvT 199759), data as holotype,
<collectionCode id="ED65AE36FFCF6173FC5BFA7BFC5B9F86" box="[937,1012,1488,1509]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="33" pageNumber="394" type="Museum">RMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B1C3CAEFFCF6173FC0DFA7BFAFF9C61" collectingDate="1987-03-20" collectionCode="RMNH" collectorName="J. v. Tol" country="Malaysia" elevation="160" pageId="33" pageNumber="394" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D72FD7AFFCF6173FC0DFA7BFBE59F85" box="[1023,1098,1488,1510]" pageId="33" pageNumber="394" type="male">3 males</specimenCount>
(JvT 199802), same area, brooklet at “West 5”,
<quantity id="4C8C9B16FFCF6173FC5BFA46FC479C61" box="[937,1000,1517,1539]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="33" pageNumber="394" unit="m" value="160.0">
<elevation id="0059D1C0FFCF6173FC5BFA46FC479C61" box="[937,1000,1517,1539]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="33" pageNumber="394" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<date id="FFCA1033FFCF6173FC01FA46FBDE9C61" box="[1011,1137,1517,1538]" pageId="33" pageNumber="394" value="1987-03-20">
<collectingDate id="EF8EE9DBFFCF6173FC01FA46FBDE9C61" box="[1011,1137,1517,1538]" pageId="33" pageNumber="394" value="1987-03-20">20 Mar.1987</collectingDate>
</date>
, coll.
<collectorName id="26815325FFCF6173FB5EFA46FB559C60" box="[1196,1274,1517,1539]" pageId="33" pageNumber="394">J. v. Tol</collectorName>
,
<collectionCode id="ED65AE36FFCF6173FAF7FA46FAFF9C61" box="[1285,1360,1517,1538]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="33" pageNumber="394" type="Museum">RMNH</collectionCode>
</materialsCitation>
;
<specimenCount id="9D72FD7AFFCF6173FAA8FA46FA089C60" box="[1370,1447,1517,1539]" pageId="33" pageNumber="394" type="male">4 males</specimenCount>
, Imbak Canyon, Ridge Trail, sub camp B,
<date id="FFCA1033FFCF6173FB37F9A1FA989C7C" box="[1221,1335,1546,1567]" pageId="33" pageNumber="394" value="2010-12-02">2 Dec.2010</date>
, coll. CYC, in collection CYC; male, same area, “small river”,
<date id="FFCA1033FFCF6173FAC0F98CFA089C5F" box="[1330,1447,1575,1596]" pageId="33" pageNumber="394" value="2010-12-03">3 Dec.2010</date>
, coll. CYC, in collection CYC.
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCF6173FCDEF92AFAD09D1A" pageId="33" pageNumber="394" type="diagnosis">
<paragraph id="8BCB36F3FFCF6173FCDEF92AFAD09D1A" blockId="33.[812,1452,1665,1913]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF92AFC0E9CF9" bold="true" box="[812,929,1665,1690]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF92AFC359CFA" bold="true" box="[812,922,1665,1689]" italics="true" pageId="33" pageNumber="394">Diagnosis</emphasis>
.
</emphasis>
— Readily distinguished from all other known
<taxonomicName id="4C744D70FFCF6173FCDEF908FC499CDA" box="[812,998,1697,1722]" class="Insecta" family="Platystictidae" genus="Telosticta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="394" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAE1FFCF6173FCDEF908FC3C9CD9" box="[812,915,1699,1722]" italics="true" pageId="33" pageNumber="394">Telosticta</emphasis>
species
</taxonomicName>
except
<taxonomicName id="4C744D70FFCF6173FBCBF908FB499CD9" authorityName="Dow &amp; Orr" authorityYear="2012" box="[1081,1254,1699,1722]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="394" phylum="Arthropoda" rank="species" species="belalongensis">
<emphasis id="B900EAE1FFCF6173FBCBF908FB499CD9" box="[1081,1254,1699,1722]" italics="true" pageId="33" pageNumber="394">T. belalongensis</emphasis>
</taxonomicName>
by the form of the superior appendages, with the terminal part flattened and folded over on its self. Separated from
<taxonomicName id="4C744D70FFCF6173FB2BF948FA279C99" authorityName="Dow &amp; Orr" authorityYear="2012" box="[1241,1416,1763,1786]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="394" phylum="Arthropoda" rank="species" species="belalongensis">
<emphasis id="B900EAE1FFCF6173FB2BF948FA279C99" box="[1241,1416,1763,1786]" italics="true" pageId="33" pageNumber="394">T. belalongensis</emphasis>
</taxonomicName>
by its shorter lateral processes of the posterior pronotal lobe; superior anal appendages without a well-developed dorsal projection, but with a dorsal crest near the end, and the shape of the spine and scoop of the inferior anal appendages.
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCF6173FCDEF80BFBD89DBA" pageId="33" pageNumber="394" type="etymology">
<paragraph id="8BCB36F3FFCF6173FCDEF80BFBD89DBA" blockId="33.[812,1451,1952,2009]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF80BFC079DD9" bold="true" box="[812,936,1952,1978]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF80BFC0E9DDA" bold="true" box="[812,929,1952,1977]" italics="true" pageId="33" pageNumber="394">Etymology</emphasis>
.
</emphasis>
<taxonomicName id="4C744D70FFCF6173FC3FF808FBBD9DD9" authorityName="Dow &amp; Orr" authorityYear="2012" box="[973,1042,1955,1978]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="394" phylum="Arthropoda" rank="species" species="janeus">
<emphasis id="B900EAE1FFCF6173FC3FF808FBBD9DD9" box="[973,1042,1955,1978]" italics="true" pageId="33" pageNumber="394">janeus</emphasis>
</taxonomicName>
, a noun in apposition; a Latinised form of Jan, named for Jan van Tol.
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCF6170FCDEF7AAFC3F9FF5" lastPageId="34" lastPageNumber="395" pageId="33" pageNumber="394" type="description">
<paragraph id="8BCB36F3FFCF6173FCDEF7AAFA04925A" blockId="33.[812,1451,2048,2105]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF7AAFBC09279" bold="true" box="[812,1135,2048,2074]" pageId="33" pageNumber="394">
<emphasis id="B900EAE1FFCF6173FCDEF7AAFBC7927A" bold="true" box="[812,1128,2048,2073]" italics="true" pageId="33" pageNumber="394">
Description of
<typeStatus id="54CF8851FFCF6173FC3DF7ABFB83927A" box="[975,1068,2048,2073]" pageId="33" pageNumber="394" type="holotype">holotype</typeStatus>
male
</emphasis>
.
</emphasis>
— Head: Labium pale. Basal 4/5 of labrum blue, black along free margin. Anteclypeus
</paragraph>
<paragraph id="8BCB36F3FFCC6170FF7AFF15FDD29BD5" blockId="34.[136,775,190,438]" pageId="34" pageNumber="395">blue, postclypeus shining black. Mandible bases blue in corner by clypeus, black below, genae black. Vertex and frons bronzy black, occiput shining black. Ratio of width of compound eye to width of vertex measured at level of lateral ocelli ca 9/10. Transverse occipital carina with lateral extremities prominent. Ocelli whitish. Antenna with scape and top ca 1/4 of pedicel brown basal 3/4 of pedicel greyish pale, flagellum dark brown (missing on right).</paragraph>
<paragraph id="8BCB36F3FFCC6170FF7AFE75FE7A9F15" blockId="34.[136,776,478,1398]" pageId="34" pageNumber="395">
Thorax: Prothorax, largely creamy white with blue patches dorsally on middle pronotal lobe, except to rear of propleuron and the rear part of middle pronotal lobe where black and dark brown with paler areas, and whole posterior pronotal lobe which is black, becoming brown towards tips of lateral pronotal processes, which are moderately long. (
<figureCitation id="134F2A76FFCC6170FD65FDD5FD5598F5" box="[663,762,638,662]" captionStart="Fig" captionStartId="20.[136,171,2062,2084]" captionTargetBox="[140,1471,1495,2006]" captionTargetId="figure-576@20.[123,1491,1485,2021]" captionTargetPageId="20" captionText="Fig. 17. T. belalongensis and T. janeus. T. belalongensis holotype: A, posterior pronotal lobe, lateral; C, right inferior anal appendage, ventral view. T. janeus holotype: B, posterior pronotal lobe, lateral; D, right inferior anal appendage, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5348651" httpUri="https://zenodo.org/record/5348651/files/figure.png" pageId="34" pageNumber="395">Fig. 17B</figureCitation>
). Synthorax: Mesepisternum bronzy black, with a pair of long pale blue antehumeral markings, tapering to rear and occupying ca 4/5 of the length (
<figureCitation id="134F2A76FFCC6170FE25FD75FD979895" box="[471,568,734,758]" captionStart="Fig" captionStartId="19.[114,149,2035,2057]" captionTargetBox="[114,1453,401,1976]" captionTargetId="figure-98@19.[114,1453,401,1976]" captionTargetPageId="19" captionText="Fig. 16. Markings of T. belalongensis and T. janeus. T. belalongensis holotype: A, antehumeral markings; C, terminal abdominal segments, dorsal view; E, terminal abdominal segments, lateral view. T. janeus holotype: B, antehumeral markings; D, terminal abdominal segments, dorsal view; F, terminal abdominal segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5348649" httpUri="https://zenodo.org/record/5348649/files/figure.png" pageId="34" pageNumber="395">Fig. 16B</figureCitation>
). Antealar triangles mostly black, except at rear where pale. Mesepimeron mostly bronzy black. Metepisternum largely occupied by a pale band, spilling over onto the lower part of the mesepimeron, becoming blue towards wing bases, with a bronzy black wedge below, this extending from the wing bases to the metinfraepisternum. Metepimeron mostly pale, with a dark brown area below the metepleural suture centrally. Venter of synthorax pale. Legs: each with coxa and trochanter cream, femur dirty cream with a narrow dark stripe long the extensor surface, and obscure grey and dark markings elsewhere, black immediately above joint. Tibia similar but more extensive dark stripe on extensor surface and another on flexor surface. Tarsus dirty cream, black around joints, claws pale brown. Wings: 13 Px in Fw, 12 Px in Hw. Vein ab absent. R
<subScript id="17F034B6FFCC6170FEFFFB61FEBA9EBA" attach="left" box="[269,277,1226,1241]" fontSize="6" pageId="34" pageNumber="395">4</subScript>
arising distal to subnodus in Fw, very slightly distal to subnodus in Hw, IR
<subScript id="17F034B6FFCC6170FE36FB41FE639E9A" attach="left" box="[452,460,1258,1273]" fontSize="6" pageId="34" pageNumber="395">3</subScript>
joined to it by a short stalk. Pterostigma approximately trapezoidal with costal side shorter than anal side, and proximal side shorter than distal side, dark brown with slightly paler border, covering slightly more than one underlying cell.
</paragraph>
<paragraph id="8BCB36F3FFCC6170FF7AFA35FAD69A95" blockId="34.[136,775,1438,2070]" lastBlockId="34.[835,1474,190,246]" pageId="34" pageNumber="395">
Abdomen: Largely brown, darkening to rear. S1 brown above and to rear laterally, white otherwise. S2 brown with a pale basal annulus (possibly pale blue in life), narrowly interrupted dorsally, laterally extending to the posterior carina as a pale wedge. S37 brown with a narrow pale basal annulus interrupted dorsally and extended laterally near lower margin of tergite for a distance, each segment becoming darker apically. S810 black with a pair of pale blue dorsal-lateral markings on S8, very narrowly separated dorsally, and a basal dorsal marking on S9 (
<figureCitation id="134F2A76FFCC6170FE8AF915FE5A9CB5" box="[376,501,1726,1750]" captionStart="Fig" captionStartId="19.[114,149,2035,2057]" captionTargetBox="[114,1453,401,1976]" captionTargetId="figure-98@19.[114,1453,401,1976]" captionTargetPageId="19" captionText="Fig. 16. Markings of T. belalongensis and T. janeus. T. belalongensis holotype: A, antehumeral markings; C, terminal abdominal segments, dorsal view; E, terminal abdominal segments, lateral view. T. janeus holotype: B, antehumeral markings; D, terminal abdominal segments, dorsal view; F, terminal abdominal segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5348649" httpUri="https://zenodo.org/record/5348649/files/figure.png" pageId="34" pageNumber="395">Fig. 16D, F</figureCitation>
). Appendages (
<figureCitation id="134F2A76FFCC6170FD65F915FCAD9CB5" box="[663,770,1726,1750]" captionStart="Fig" captionStartId="20.[136,171,2062,2084]" captionTargetBox="[140,1471,1495,2006]" captionTargetId="figure-576@20.[123,1491,1485,2021]" captionTargetPageId="20" captionText="Fig. 17. T. belalongensis and T. janeus. T. belalongensis holotype: A, posterior pronotal lobe, lateral; C, right inferior anal appendage, ventral view. T. janeus holotype: B, posterior pronotal lobe, lateral; D, right inferior anal appendage, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5348651" httpUri="https://zenodo.org/record/5348651/files/figure.png" pageId="34" pageNumber="395">Figs. 17D</figureCitation>
,
<figureCitation id="134F2A76FFCC6170FF7AF975FF7C9C95" box="[136,211,1758,1782]" captionStart="Fig" captionStartId="21.[112,147,2063,2085]" captionTargetBox="[113,1452,1051,2011]" captionTargetId="figure-431@21.[101,1469,1019,2020]" captionTargetPageId="21" captionText="Fig. 18. T. belalongensis and T. janeus. T. belalongensis holotype:A, anal appendages dorsal; C, anal appendages lateral. T. janeus holotype: B, anal appendages dorsal; D, anal appendages lateral." figureDoi="http://doi.org/10.5281/zenodo.5348653" httpUri="https://zenodo.org/record/5348653/files/figure.png" pageId="34" pageNumber="395">18B, D</figureCitation>
) largely dark brown with obscure paler areas interiorly and dorsally. Superior appendage ca 2.5 times the length of S10, with robust but blunt interior projection at ca 1/3 length directed inwards. Dorsal projection a small rounded swelling only narrowly separated from the basal projection, but there is a thin subapical ca semicircular crest, prominent in lateral view. Expanded in distal 1/2 but folded under itself to nearly form an open-mouthed tube with the outer corner produced in a distinctive manner (
<figureCitation id="134F2A76FFCC6170FE71F875FE4D9D95" box="[387,482,2014,2038]" captionStart="Fig" captionStartId="21.[112,147,2063,2085]" captionTargetBox="[113,1452,1051,2011]" captionTargetId="figure-431@21.[101,1469,1019,2020]" captionTargetPageId="21" captionText="Fig. 18. T. belalongensis and T. janeus. T. belalongensis holotype:A, anal appendages dorsal; C, anal appendages lateral. T. janeus holotype: B, anal appendages dorsal; D, anal appendages lateral." figureDoi="http://doi.org/10.5281/zenodo.5348653" httpUri="https://zenodo.org/record/5348653/files/figure.png" pageId="34" pageNumber="395">Fig. 18B</figureCitation>
). Inferior appendage almost as long as superior, stem robust and scoop strongly expanded then tapering to tip, up- and out-turned almost immediately, spine directed inwards, slightly upwards and to rear.
</paragraph>
<paragraph id="8BCB36F3FFCC6170FCB1FEB4FB7E9B35" blockId="34.[835,1474,286,342]" pageId="34" pageNumber="395">Measurements (mm): Abdomen without anal appendages 33, superior appendage ca 1.2, Hw 20.5.</paragraph>
<paragraph id="8BCB36F3FFCC6170FCB1FED6FB9F9BF5" blockId="34.[835,1072,381,406]" box="[835,1072,381,406]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FED6FC329BF5" bold="true" box="[835,925,381,406]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FED6FC399BF5" bold="true" box="[835,918,381,406]" italics="true" pageId="34" pageNumber="395">Female</emphasis>
.
</emphasis>
— Unknown
</paragraph>
<paragraph id="8BCB36F3FFCC6170FCB1FE15FB029EF5" blockId="34.[835,1474,446,1174]" pageId="34" pageNumber="395">
Variation in
<materialsCitation id="3B1C3CAEFFCC6170FC2AFE14FA869B95" location="There" pageId="34" pageNumber="395" specimenCount="1" typeStatus="paratype">
<emphasis id="B900EAE1FFCC6170FC2AFE14FBE49BB5" bold="true" box="[984,1099,446,470]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FC2AFE14FBEB9BB5" bold="true" box="[984,1092,447,470]" italics="true" pageId="34" pageNumber="395">
<typeStatus id="54CF8851FFCC6170FC2AFE14FBEB9BB5" box="[984,1092,447,470]" pageId="34" pageNumber="395" type="paratype">paratypes</typeStatus>
</emphasis>
.
</emphasis>
<location id="8EAB6028FFCC6170FB88FE15FB159BB5" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCC6170FB88FE15FB159BB5" box="[1146,1210,446,470]" name="There" pageId="34" pageNumber="395">There</location>
are small variations in the markings of the head and thorax in the
</materialsCitation>
<typeStatus id="54CF8851FFCC6170FAC1FE74FA319B94" box="[1331,1438,479,503]" pageId="34" pageNumber="395" type="paratype">paratypes</typeStatus>
; in particular the extent of the intrusion of the pale stripe on the metepisternum onto the mesepimeron is greater than in the
<materialsCitation id="3B1C3CAEFFCC6170FC9CFD95FAF59895" county="There" location="In" municipality="In" pageId="34" pageNumber="395" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF8851FFCC6170FC9CFD95FC7F9835" box="[878,976,574,598]" pageId="34" pageNumber="395" type="holotype">holotype</typeStatus>
.
<location id="8EAB6028FFCC6170FC2FFD94FC5C9835" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCC6170FC2FFD94FC5C9835" box="[989,1011,575,598]" county="There" municipality="In" name="In" pageId="34" pageNumber="395">In</location>
one of the
<location id="8EAB6028FFCC6170FB88FD95FAB29835" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCC6170FB88FD95FAB29835" box="[1146,1309,574,598]" county="There" municipality="In" name="Imbak Canyon" pageId="34" pageNumber="395">Imbak Canyon</location>
specimens the antehumeral stripes are shorter, ca 2/3 of the length of the metepisternum.
<collectingMunicipality id="6BAFAC89FFCC6170FC1CFDD4FBAB98F5" box="[1006,1028,639,662]" pageId="34" pageNumber="395">In</collectingMunicipality>
<specimenCount id="9D72FD7AFFCC6170FBF8FDD4FB3C98F4" box="[1034,1171,639,663]" pageId="34" pageNumber="395" type="generic" typeStatus="paratype">one paratype</specimenCount>
the lateral extremities of the transverse carina are much less prominent.
<collectingCounty id="62AA4E7FFFCC6170FAD7FD35FACC98D5" box="[1317,1379,670,694]" pageId="34" pageNumber="395">There</collectingCounty>
is some variation in the length of the lateral processes of the posterior pronotal lobe, in one case they are longer than in the
</materialsCitation>
<materialsCitation id="3B1C3CAEFFCC6170FAADFD75FB8999B5" collectorName="The S" location="The" pageId="34" pageNumber="395" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF8851FFCC6170FAADFD75FA119895" box="[1375,1470,734,758]" pageId="34" pageNumber="395" type="holotype">holotype</typeStatus>
, in another they are shorter and poorly developed.
<location id="8EAB6028FFCC6170FABCFD55FAD89975" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCC6170FABCFD55FAD89975" box="[1358,1399,766,790]" name="The" pageId="34" pageNumber="395">The</location>
colour of the pterostigma varies from deep chestnut brown to almost black and the paler border is often only partially present and varies from just paler than the central part to considerably paler; in
<specimenCount id="9D72FD7AFFCC6170FC56FCD4FB8099F4" box="[932,1071,895,919]" pageId="34" pageNumber="395" type="generic" typeStatus="paratype">one paratype</specimenCount>
no border is apparent.
<collectorName id="26815325FFCC6170FAD9FCD5FAC499F5" box="[1323,1387,894,918]" pageId="34" pageNumber="395">The S</collectorName>
8 dorsal markings are sometimes fused and sometimes more widely separated than in the
</materialsCitation>
<materialsCitation id="3B1C3CAEFFCC6170FBDCFC15FB069EF5" county="Danum Valley" location="The" municipality="The Imbak Canyon" pageId="34" pageNumber="395" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF8851FFCC6170FBDCFC15FB2199B5" box="[1070,1166,958,982]" pageId="34" pageNumber="395" type="holotype">holotype</typeStatus>
, and the dorsal part appears to fade somewhat with increasing maturity.
<location id="8EAB6028FFCC6170FAFDFC75FA979995" LSID="urn:lsid:plazi:treatment:03DD87E5FFCF6170FC34FB56FB1A9F95:8EAB6028FFCC6170FAFDFC75FA979995" box="[1295,1336,990,1014]" county="Danum Valley" municipality="The Imbak Canyon" name="The" pageId="34" pageNumber="395">The</location>
extent of the S9 marking varies considerably, and it is sometimes divided into two.
<collectingMunicipality id="6BAFAC89FFCC6170FC5FFBB5FB2F9E55" box="[941,1152,1054,1078]" pageId="34" pageNumber="395">The Imbak Canyon</collectingMunicipality>
specimens are slightly larger on average than those from the
<collectingCounty id="62AA4E7FFFCC6170FB6AFB94FA9D9E35" box="[1176,1330,1086,1110]" pageId="34" pageNumber="395">Danum Valley</collectingCounty>
, and have S1 paler dorsally and slightly more extensive pale colouration along the lower margins of S37
</materialsCitation>
.
</paragraph>
<paragraph id="8BCB36F3FFCC6170FCB1FB14FA149E95" blockId="34.[835,1473,1214,1270]" pageId="34" pageNumber="395">
Measurements (mm): Abdomen without appendages 3135, Hw 19.522.5. Wings with 1113 Px in Fw,
<quantity id="4C8C9B16FFCC6170FADAFB75FA279E95" box="[1320,1416,1246,1270]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="34" pageNumber="395" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
Hw.
</paragraph>
<paragraph id="8BCB36F3FFCC6170FCB1FAB6FC3F9FF5" blockId="34.[835,1474,1309,1430]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FAB6FC549F55" bold="true" box="[835,1019,1309,1334]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FAB6FC5B9F55" bold="true" box="[835,1012,1309,1334]" italics="true" pageId="34" pageNumber="395">Biological notes</emphasis>
.
</emphasis>
— Little has been recorded on the biology of this species, but all specimens have been collected at small streams in mixed dipterocarp forest in hilly or mountainous terrain.
</paragraph>
</subSubSection>
<subSubSection id="C36E6578FFCC6170FCB1FA16FB1A9F95" pageId="34" pageNumber="395" type="distribution">
<paragraph id="8BCB36F3FFCC6170FCB1FA16FB1A9F95" blockId="34.[835,1474,1469,1526]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FA16FC609FB5" bold="true" box="[835,975,1469,1494]" pageId="34" pageNumber="395">
<emphasis id="B900EAE1FFCC6170FCB1FA16FC679FB5" bold="true" box="[835,968,1469,1494]" italics="true" pageId="34" pageNumber="395">Distribution</emphasis>
.
</emphasis>
— Known only from the Danum Valley and Imbak Canyon in
<collectingRegion id="49B0F811FFCC6170FBF5FA75FBE79F95" box="[1031,1096,1502,1526]" country="Malaysia" name="Sabah" pageId="34" pageNumber="395">Sabah</collectingRegion>
(
<figureCitation id="134F2A76FFCC6170FBAAFA75FB079F95" box="[1112,1192,1502,1526]" captionStart="Fig" captionStartId="22.[835,870,2063,2085]" captionTargetBox="[835,1473,1434,2023]" captionTargetId="figure-749@22.[835,1472,1434,2023]" captionTargetPageId="22" captionText="Fig. 19. Distribution of T. bidayuh (cross), T. janeus (open square), T. feronia (black square), and T. longigaster (circle)." figureDoi="http://doi.org/10.5281/zenodo.5348655" httpUri="https://zenodo.org/record/5348655/files/figure.png" pageId="34" pageNumber="395">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>