treatments-xml/data/03/DD/87/03DD87E12E17C47EA69EFDF0FA31676D.xml

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<mods:title id="7467439A6444874285011FCE244283FF">Four new species of the subgenus Pachylaelaps (Longipachylaelaps) from Italy, with a new identification key to the known species (Acari: Pachylaelapidae)</mods:title>
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<paragraph id="8BCB36F72E17C47DA69EFDF0FB206267" blockId="17.[151,1162,548,574]" box="[151,1162,548,574]" pageId="17" pageNumber="112">
<heading id="D083819B2E17C47DA69EFDF0FB206267" bold="true" box="[151,1162,548,574]" fontSize="11" level="1" pageId="17" pageNumber="112" reason="1">
<emphasis id="B900EAE52E17C47DA69EFDF0FB206267" bold="true" box="[151,1162,548,574]" pageId="17" pageNumber="112">
Key to known species of the subgenus 
<taxonomicName id="4C744D742E17C47DA47AFDF0FBBB6267" ID-CoL="8W4W4" authority="Longipachylaelaps" authorityName="Longipachylaelaps" box="[627,1041,548,574]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAE52E17C47DA47AFDF0FCB96267" bold="true" box="[627,787,548,574]" italics="true" pageId="17" pageNumber="112">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E17C47DA52DFDF0FBA26267" bold="true" box="[804,1032,548,574]" italics="true" pageId="17" pageNumber="112">Longipachylaelaps</emphasis>
)
</taxonomicName>
(females)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EFDBEFC826302" blockId="17.[151,1437,618,1219]" pageId="17" pageNumber="112">
<materialsCitation id="3B1C3CAA2E17C47DA69EFDBEFC8E6302" ID-GBIF-Occurrence="1434656193" collectingDate="2008-05-10" collectorName="Serra da Labruja Mts &amp; San Bento da Porta Aberta &amp; P. Fenda" country="Turkey" county="Partial" determinerName="P. Masan" elevation="260" location="Karg" municipality="European" pageId="17" pageNumber="112" specimenCount="1" stateProvince="Viana do Castelo">
<collectingCounty id="62AA4E7B2E17C47DA69EFDBEFF4A62DA" box="[151,224,618,643]" pageId="17" pageNumber="112">Partial</collectingCounty>
keys to 
<collectingMunicipality id="6BAFAC8D2E17C47DA74FFDBFFE1862DB" box="[326,434,619,642]" pageId="17" pageNumber="112">European</collectingMunicipality>
species of 
<taxonomicName id="4C744D742E17C47DA43FFDBFFD6762DA" box="[566,717,619,643]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAE52E17C47DA43FFDBFFD6762DA" box="[566,717,619,643]" italics="true" pageId="17" pageNumber="112">Pachylaelaps</emphasis>
</taxonomicName>
may be found in 
<bibRefCitation id="EFE54B062E17C47DA596FDBFFB8E62DA" author="Karg" box="[927,1060,619,643]" pageId="17" pageNumber="112" refString="Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1 - 475." type="journal article" year="1971">
<location id="8EAB602C2E17C47DA596FDBFFC7D62DA" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA596FDBFFC7D62DA" box="[927,983,619,643]" country="Turkey" county="Partial" municipality="European" name="Karg" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Karg</location>
(1971
</bibRefCitation>
, 
<bibRefCitation id="EFE54B062E17C47DA23BFDBFFBC762DA" author="Karg" box="[1074,1133,619,643]" pageId="17" pageNumber="112" refString="Karg, W. (1993) Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach. Raubmilben. 2. Uberarbeitete Auflage. Die Tierwelt Deutschlands, 59, 1 - 523." type="journal article" year="1993">1993</bibRefCitation>
), 
<bibRefCitation id="EFE54B062E17C47DA28DFDBEFAE962DA" author="Koroleva" box="[1156,1347,618,643]" pageId="17" pageNumber="112" refString="Koroleva, E. V. (1977) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites, Mesostigmata. Nauka Press, Leningrad, pp. 411 - 483. [in Russian]" type="book chapter" year="1977">
<location id="8EAB602C2E17C47DA28DFDBEFB4662DA" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA28DFDBEFB4662DA" box="[1156,1260,618,643]" country="Turkey" county="Partial" municipality="European" name="Koroleva" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Koroleva</location>
(1977)
</bibRefCitation>
, 
<bibRefCitation id="EFE54B062E17C47DA358FDBFFF5A62FE" author="Masan" pageId="17" pageNumber="112" refString="Masan, P. (2007 a) A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea). NOI Press, Bratislava, 247 pp." type="book" year="2007" yearSuffix="a">
<location id="8EAB602C2E17C47DA358FDBFFA3662DA" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA358FDBFFA3662DA" box="[1361,1436,619,643]" country="Turkey" county="Partial" municipality="European" name="Masan" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Mašán</location>
(2007a)
</bibRefCitation>
, and 
<location id="8EAB602C2E17C47DA739FD5BFED162FE" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA739FD5BFED162FE" box="[304,379,655,679]" country="Turkey" county="Partial" municipality="European" name="Masan" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Mašán</location>
<emphasis id="B900EAE52E17C47DA78DFD5BFE1D62FF" box="[388,439,654,678]" italics="true" pageId="17" pageNumber="112">et al</emphasis>
. (2016). A new key to the subgenus 
<emphasis id="B900EAE52E17C47DA56EFD5AFB9662FF" box="[871,1084,654,678]" italics="true" pageId="17" pageNumber="112">
<location id="8EAB602C2E17C47DA56EFD5AFB9662FF" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA56EFD5AFB9662FF" box="[871,1084,654,678]" country="Turkey" county="Partial" municipality="European" name="Longipachylaelaps" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Longipachylaelaps</location>
</emphasis>
is needed to include a species described recently from 
<collectingCountry id="F36376672E17C47DA7A3FD66FE506292" box="[426,506,690,715]" name="Turkey" pageId="17" pageNumber="112">Turkey</collectingCountry>
(
<bibRefCitation id="EFE54B062E17C47DA403FD66FD356292" author="Ozbek" box="[522,671,690,715]" pageId="17" pageNumber="112" refString="Ozbek, H. H. (2015) An unusual new species of Pachylaelaps (Acari: Pachylaelapidae) from Turkey. International Journal of Acarology, 41, 67 - 70." type="journal article" year="2015">
<location id="8EAB602C2E17C47DA403FD66FDFD6292" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA403FD66FDFD6292" box="[522,599,690,715]" country="Turkey" county="Partial" municipality="European" name="Ozbek" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Özbek</location>
, 2015
</bibRefCitation>
), the four new species described in this study, and a revised species of 
<location id="8EAB602C2E17C47DA6B1FD02FEEB62B6" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA6B1FD02FEEB62B6" box="[184,321,726,751]" country="Turkey" county="Partial" municipality="European" name="Hirschmann" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Hirschmann</location>
&amp; 
<location id="8EAB602C2E17C47DA760FD03FE1D62B7" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA760FD03FE1D62B7" box="[361,439,727,750]" country="Turkey" county="Partial" municipality="European" name="Krauss" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Krauss</location>
(1965), namely 
<taxonomicName id="4C744D742E17C47DA471FD03FC9062B7" box="[632,826,726,751]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="granulifer" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA471FD03FD2C62B7" box="[632,646,727,750]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA4A9FD03FD0562B7" box="[672,687,727,750]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA4C0FD02FC9062B7" box="[713,826,726,750]" italics="true" pageId="17" pageNumber="112">granulifer</emphasis>
</taxonomicName>
, based on numerous specimens collected in 
<collectingCountry id="F36376672E17C47DA334FD02FA3662B6" box="[1341,1436,726,751]" name="Portugal" pageId="17" pageNumber="112">Portugal</collectingCountry>
(
<location id="8EAB602C2E17C47DA697FD2EFEE1634A" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA697FD2EFEE1634A" box="[158,331,762,787]" country="Turkey" county="Partial" municipality="European" name="North Portugal" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">North Portugal</location>
, 
<location id="8EAB602C2E17C47DA75FFD2FFDE5634B" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA75FFD2FFDE5634B" box="[342,591,762,787]" country="Turkey" county="Partial" municipality="European" name="Serra da Labruja Mts." pageId="17" pageNumber="112" stateProvince="Viana do Castelo">
<collectorName id="268153212E17C47DA75FFD2FFDE0634B" box="[342,586,762,787]" pageId="17" pageNumber="112">Serra da Labruja Mts</collectorName>
.
</location>
, 
<location id="8EAB602C2E17C47DA453FD2FFC48634A" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA453FD2FFC48634A" box="[602,994,762,787]" country="Turkey" county="Partial" municipality="European" name="San Bento da Porta Aberta Village" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">
<collectorName id="268153212E17C47DA453FD2FFC2D634A" box="[602,903,762,787]" pageId="17" pageNumber="112">San Bento da Porta Aberta</collectorName>
Village
</location>
, 
<collectingRegion id="49B0F8152E17C47DA5E7FD2EFB07634A" box="[1006,1197,762,787]" country="Portugal" name="Viana do Castelo" pageId="17" pageNumber="112">Viana do Castelo</collectingRegion>
Cadaster, non-native eucalyptus forest (
<emphasis id="B900EAE52E17C47DA77EFCCAFDCE636F" box="[375,612,798,822]" italics="true" pageId="17" pageNumber="112">
<location id="8EAB602C2E17C47DA77EFCCAFE5E636F" LSID="urn:lsid:plazi:treatment:03DD87E12E17C47EA69EFDF0FA31676D:8EAB602C2E17C47DA77EFCCAFE5E636F" box="[375,500,798,822]" country="Turkey" county="Partial" municipality="European" name="Eucalyptus" pageId="17" pageNumber="112" stateProvince="Viana do Castelo">Eucalyptus</location>
globulus
</emphasis>
), humid leaf litter and soil detritus, altitude 
<quantity id="4C8C9B122E17C47DA29FFCCAFB42636F" box="[1174,1256,798,823]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="112" unit="m" value="260.0">
<elevation id="0059D1C42E17C47DA29FFCCAFB42636F" box="[1174,1256,798,823]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="112" unit="m" value="260.0">260 m</elevation>
</quantity>
, 41°56'02,3&quot;N, 08°37'49,9&quot;W, 
<date id="FFCA10372E17C47DA748FC97FE736302" box="[321,473,835,859]" pageId="17" pageNumber="112" value="2008-05-10">
<collectingDate id="EF8EE9DF2E17C47DA748FC97FE736302" box="[321,473,835,859]" pageId="17" pageNumber="112" value="2008-05-10">10 May 2008</collectingDate>
</date>
, lgt. 
<collectorName id="268153212E17C47DA419FC97FDDD6302" box="[528,631,834,859]" pageId="17" pageNumber="112">P. Fenďa</collectorName>
, det. 
<determinerName id="7E17DA332E17C47DA4B8FC97FCB66302" box="[689,796,835,859]" pageId="17" pageNumber="112">P. Mašán</determinerName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCB36F72E17C47DA6CEFCB2FBF5649A" blockId="17.[151,1437,618,1219]" pageId="17" pageNumber="112">
The identification of 
<taxonomicName id="4C744D742E17C47DA7C9FCB2FDFD6327" box="[448,599,870,894]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAE52E17C47DA7C9FCB2FDFD6327" box="[448,599,870,894]" italics="true" pageId="17" pageNumber="112">Pachylaelaps</emphasis>
</taxonomicName>
species is complicated by the incomplete descriptions of some species. Therefore some species are not included in the keys presented in this paper, namely 
<taxonomicName id="4C744D742E17C47DA30CFC5FFCB3639E" authority="Hirschmann &amp; Krauss, 1965" authorityName="Hirschmann &amp; Krauss" authorityYear="1965" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="dubius" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA30CFC5FFA3663FA" box="[1285,1436,907,931]" italics="true" pageId="17" pageNumber="112">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E17C47DA6A9FC7AFEDF639F" box="[160,373,942,966]" italics="true" pageId="17" pageNumber="112">Longipachylaelaps</emphasis>
) 
<emphasis id="B900EAE52E17C47DA78FFC7AFE7B639F" box="[390,465,942,966]" italics="true" pageId="17" pageNumber="112">dubius</emphasis>
Hirschmann &amp; Krauss, 1965
</taxonomicName>
and 
<taxonomicName id="4C744D742E17C47DA559FC7BFB63639E" authority="Schmolzer, 1992" authorityName="Schmolzer" authorityYear="1992" box="[848,1225,942,967]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="obirensis" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA559FC7BFCF4639F" box="[848,862,943,966]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA57CFC7BFC2E639F" box="[885,900,943,966]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA595FC7AFBAB639F" box="[924,1025,942,966]" italics="true" pageId="17" pageNumber="112">obirensis</emphasis>
Schmölzer, 1992
</taxonomicName>
, including also the two dubious species 
<taxonomicName id="4C744D742E17C47DA774FC07FD6B63B2" authority="Berlese, 1920" authorityName="Berlese" authorityYear="1920" box="[381,705,978,1003]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="gallicus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA774FC07FE2163B3" box="[381,395,979,1002]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA7ABFC07FE1B63B3" box="[418,433,979,1002]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA7C1FC07FD8B63B2" box="[456,545,979,1003]" italics="true" pageId="17" pageNumber="112">gallicus</emphasis>
Berlese, 1920
</taxonomicName>
and 
<taxonomicName id="4C744D742E17C47DA4FFFC07FBD863B2" authority="Duges, 1834" authorityName="Duges" authorityYear="1834" box="[758,1138,979,1003]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="tetragonoides" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA4FFFC07FCAE63B3" box="[758,772,979,1002]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA512FC07FC8063B3" box="[795,810,979,1002]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA548FC07FC7163B2" box="[833,987,979,1003]" italics="true" pageId="17" pageNumber="112">tetragonoides</emphasis>
Dugès, 1834
</taxonomicName>
. The male described under the name 
<taxonomicName id="4C744D742E17C47DA70FFC23FDCF6456" authority="Halbert, 1915" authorityName="Halbert" authorityYear="1915" box="[262,613,1014,1039]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="longisetis" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA70FFC23FEBE6457" box="[262,276,1015,1038]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA724FC23FE966457" box="[301,316,1015,1038]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA75FFC22FE6B6457" box="[342,449,1014,1038]" italics="true" pageId="17" pageNumber="112">longisetis</emphasis>
Halbert, 1915
</taxonomicName>
by 
<bibRefCitation id="EFE54B062E17C47DA49BFC23FC246456" author="Evans" box="[658,910,1014,1039]" pageId="17" pageNumber="112" refString="Evans, G. O. &amp; Hyatt, K. H. (1956) British mites of the genus Pachylaelaps Berlese (Gamasina - Pachylaelaptidae). Entomologist's Monthly Magazine, 92, 118 - 129." type="journal article" year="1956">Evans &amp; Hyatt (1956)</bibRefCitation>
is considered here as a misidentification of 
<taxonomicName id="4C744D742E17C47DA38EFC23FD91646A" authority="Evans &amp; Hyatt, 1956" authorityName="Evans &amp; Hyatt" authorityYear="1956" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="undulatus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA38EFC23FA3F6457" box="[1415,1429,1015,1038]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA6A9FBCFFF05646B" box="[160,175,1051,1074]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA6C3FBCFFE92646A" box="[202,312,1051,1075]" italics="true" pageId="17" pageNumber="112">undulatus</emphasis>
<bibRefCitation id="EFE54B062E17C47DA74AFBCFFD91646A" author="Evans" box="[323,571,1050,1075]" pageId="17" pageNumber="112" refString="Evans, G. O. &amp; Hyatt, K. H. (1956) British mites of the genus Pachylaelaps Berlese (Gamasina - Pachylaelaptidae). Entomologist's Monthly Magazine, 92, 118 - 129." type="journal article" year="1956">Evans &amp; Hyatt, 1956</bibRefCitation>
</taxonomicName>
, but this suggestion needs to be verified by further study. The keys below contain 26 species of which 15 species are based on both adult stages, six species exclusively on males (
<taxonomicName id="4C744D742E17C47DA34EFBEBFA3F640E" box="[1351,1429,1087,1111]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="conifer">
<emphasis id="B900EAE52E17C47DA34EFBEBFA3F640E" box="[1351,1429,1087,1111]" italics="true" pageId="17" pageNumber="112">conifer</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA69EFBB7FEAB6422" box="[151,257,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B900EAE52E17C47DA69EFBB7FEAB6422" box="[151,257,1123,1147]" italics="true" pageId="17" pageNumber="112">decipiens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA706FBB7FED96422" box="[271,371,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B900EAE52E17C47DA706FBB7FED96422" box="[271,371,1123,1147]" italics="true" pageId="17" pageNumber="112">gibbosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA788FBB7FE406422" box="[385,490,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="hestulifer">
<emphasis id="B900EAE52E17C47DA788FBB7FE406422" box="[385,490,1123,1147]" italics="true" pageId="17" pageNumber="112">hestulifer</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA7F1FBB7FD2D6422" box="[504,647,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="longicrinitus">
<emphasis id="B900EAE52E17C47DA7F1FBB7FD2D6422" box="[504,647,1123,1147]" italics="true" pageId="17" pageNumber="112">longicrinitus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C744D742E17C47DA4CCFBB7FCA46423" box="[709,782,1123,1146]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="virago">
<emphasis id="B900EAE52E17C47DA4CCFBB7FCA46423" box="[709,782,1123,1146]" italics="true" pageId="17" pageNumber="112">virago</emphasis>
</taxonomicName>
), and five species only on females (
<taxonomicName id="4C744D742E17C47DA2ADFBB7FAB26422" box="[1188,1304,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B900EAE52E17C47DA2ADFBB7FAB26422" box="[1188,1304,1123,1147]" italics="true" pageId="17" pageNumber="112">anatolicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA32EFBB7FA3F6422" box="[1319,1429,1123,1147]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="bifurciger">
<emphasis id="B900EAE52E17C47DA32EFBB7FA3F6422" box="[1319,1429,1123,1147]" italics="true" pageId="17" pageNumber="112">bifurciger</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA69EFB52FEA864C7" box="[151,258,1158,1182]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="longisetis">
<emphasis id="B900EAE52E17C47DA69EFB52FEA864C7" box="[151,258,1158,1182]" italics="true" pageId="17" pageNumber="112">longisetis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C744D742E17C47DA718FB52FED564C7" box="[273,383,1158,1182]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B900EAE52E17C47DA718FB52FED564C7" box="[273,383,1158,1182]" italics="true" pageId="17" pageNumber="112">undulatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C744D742E17C47DA7B7FB53FDB264C7" box="[446,536,1159,1182]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B900EAE52E17C47DA7B7FB53FDB264C7" box="[446,536,1159,1182]" italics="true" pageId="17" pageNumber="112">vicarius</emphasis>
</taxonomicName>
). The identification key for male stages is newly developed for the purposes of this paper. The key to identification of females is derived from that by 
<bibRefCitation id="EFE54B062E17C47DA5A6FB7FFBF0649A" author="Masan" box="[943,1114,1195,1219]" pageId="17" pageNumber="112" refString="Masan, P. (2007 a) A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea). NOI Press, Bratislava, 247 pp." type="book" year="2007" yearSuffix="a">Mašán (2007a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E17C47EA69EFB26FA31676D" lastPageId="18" lastPageNumber="113" pageId="17" pageNumber="112" type="key">
<keyStep id="30802E522E17C47DA69EFB26FA316506" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EFB26FA36657D" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EFB26FA36657D" pageId="17" pageNumber="112">
1. Cheliceral digits polydentate, each with at least ten denticles; central projection of epistome apically pointed, lanceolate; length of dorsal shield 720–810 µm............................................... 
<taxonomicName id="4C744D742E17C47DA27FFAC4FA36657D" ID-CoL="8Y7KZ" authority="Ozbek, 2015" authorityName="Ozbek" authorityYear="2015" box="[1142,1436,1295,1317]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="anatolicus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA27FFAC4FB28657C" box="[1142,1154,1296,1317]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA29FFAC4FB09657C" box="[1174,1187,1296,1317]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA2BEFADBFAB1657C" box="[1207,1307,1295,1317]" italics="true" pageId="17" pageNumber="112">anatolicus</emphasis>
<bibRefCitation id="EFE54B062E17C47DA328FADBFA36657D" author="Ozbek" box="[1313,1436,1295,1316]" pageId="17" pageNumber="112" refString="Ozbek, H. H. (2015) An unusual new species of Pachylaelaps (Acari: Pachylaelapidae) from Turkey. International Journal of Acarology, 41, 67 - 70." type="journal article" year="2015">Özbek, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EFAF9FA316506" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EFAF9FA316506" pageId="17" pageNumber="112">- Cheliceral digits oligodentate, each with at most five denticles; central projection of epistome apically widened, with anterior margin densely crenellated............................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E17C47DA69EFAB3FA31658D" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EFAB3FA3165C3" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EFAB3FA3165C3" pageId="17" pageNumber="112">
2. Structures of sperm induction system between coxae IV well sclerotised (at least some basal or distal components), usually well discernible and striking in their lines (
<figureCitation id="134F2A722E17C47DA472FA51FD4065C3" box="[635,746,1413,1434]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,263,956,979]" captionTargetBox="[151,1436,193,935]" captionTargetId="figure@4.[151,1436,193,935]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Pachylaelaps (Longipachylaelaps) abludens sp. nov., sperm induction structures, female, variant forms. 10 ‒ 11. Normal forms; 12 ‒ 13. Aberrant forms in strongly pressed specimens. Not scaled." httpUri="https://zenodo.org/record/322029/files/figure.png" pageId="17" pageNumber="112">Figs 10–13</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA4FDFA51FC9865C3" box="[756,818,1413,1434]" captionStart="FIGURES 23 – 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 – 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="17" pageNumber="112">23–26</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA535FA51FCD065C3" box="[828,890,1413,1434]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="17" pageNumber="112">49–52</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA58DFA51FC6B65C3" box="[900,961,1413,1434]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">57–61</figureCitation>
).................................... 3
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EFA76FA31658D" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EFA76FA31658D" pageId="17" pageNumber="112">
- Structures of sperm induction system weakly sclerotised, hyaline and transparent, scarcely detectable, often poorly developed or fully reduced (
<figureCitation id="134F2A722E17C47DA78EFA6BFE67658D" box="[391,461,1471,1492]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">Fig. 62</figureCitation>
)............................................................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E17C47DA69EFA09FA31663E" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EFA09FA316656" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EFA09FA316656" pageId="17" pageNumber="112">3. Sperm induction system tubular: tubiform structures simple, short or elongated, straight or curved, sometimes variously convoluted or broadened distally............................................................................. 4</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EF9C3FA31663E" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF9C3FA31663E" pageId="17" pageNumber="112">
- Sperm induction system sacculate or otherwise modified: tubiform structures absent and transformed into saccules with short tubiform opening only slightly protuberant above enlarged base (
<figureCitation id="134F2A722E17C47DA559F9E1FC3D6613" box="[848,919,1589,1610]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">Fig. 61</figureCitation>
), or into specific sickle-shaped structures (
<figureCitation id="134F2A722E17C47DA338F9E1FADD6613" box="[1329,1399,1589,1610]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">Fig. 60</figureCitation>
).................................................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E17C47DA69EF9BBFA316685" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EF9BBFA3666FB" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF9BBFA3666FB" pageId="17" pageNumber="112">
4. Movable digit of chelicera tridentate; tubiform structures of sperm induction system short and narrow, straight or slightly curved, regularly sclerotised (
<figureCitation id="134F2A722E17C47DA7F7F959FDEC66FB" box="[510,582,1677,1698]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">Fig. 57</figureCitation>
); length of dorsal shield 750–800 µm............... 
<taxonomicName id="4C744D742E17C47DA27CF959FA3666FB" ID-CoL="8Y7KK" authority="Halbert, 1915" authorityName="Halbert" authorityYear="1915" box="[1141,1436,1676,1698]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="longisetis" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA27CF959FB2B66FB" box="[1141,1153,1677,1698]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA29DF959FB0B66FB" box="[1172,1185,1677,1698]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA2BFF958FAB866FB" box="[1206,1298,1676,1698]" italics="true" pageId="17" pageNumber="112">longisetis</emphasis>
Halbert, 1915
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EF97EFA316685" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF97EFA316685" pageId="17" pageNumber="112">
- Movable digit of chelicera bidentate; tubiform structures longer, often irregularly sclerotised, and otherwise formed as depicted in 
<figureCitation id="134F2A722E17C47DA758F913FA316685" box="[337,1435,1735,1756]" captionStart-0="FIGURES 57 – 62" captionStart-1="FIGURES 3 – 9" captionStartId-0="20.[151,263,1370,1393]" captionStartId-1="3.[151,263,1834,1857]" captionTargetBox-0="[151,1436,193,1349]" captionTargetBox-1="[160,1424,207,1798]" captionTargetId-0="figure@20.[151,1436,193,1349]" captionTargetId-1="figure@3.[151,1436,193,1813]" captionTargetPageId-0="20" captionTargetPageId-1="3" captionText-0="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." captionText-1="FIGURES 3 – 9. Pachylaelaps (Longipachylaelaps) abludens sp. nov. 3. Sperm induction structures, female; 4. Epistome, female; 5. Chelicera, male; 6. Chelicera, female; 7. Ventral spurs on leg II (femur, genu and tibia), male; 8. Palptibial projections, male; 9. Tarsus II, female, mediodistal part. Not scaled." httpUri-0="https://zenodo.org/record/322040/files/figure.png" httpUri-1="https://zenodo.org/record/322028/files/figure.png" pageId="17" pageNumber="112">Fig. 57................................................................................... 5</figureCitation>
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<keyLead id="308595C22E17C47DA69EF931FA3666A3" box="[151,1436,1765,1786]" pageId="17" pageNumber="112">
5. Tubiform structures of sperm induction system shorter, with their apices separated (
<figureCitation id="134F2A722E17C47DA20BF931FBDA66A3" box="[1026,1136,1765,1786]" captionStart="FIGURES 23 – 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 – 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="17" pageNumber="112">Figs 23–26</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA273F931FB1266A3" box="[1146,1208,1765,1786]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="17" pageNumber="112">51, 52</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA2CBF931FB5566A3" box="[1218,1279,1765,1786]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">58, 59</figureCitation>
).......... 6
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EF8D6FA36674E" blockId="17.[151,1436,1266,2020]" box="[151,1436,1794,1815]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF8D6FA36674E" box="[151,1436,1794,1815]" pageId="17" pageNumber="112">
- Tubiform structures longer, with their apices or distal sections adjacent, sometimes with bends (
<figureCitation id="134F2A722E17C47DA27FF8D6FB4E674E" box="[1142,1252,1794,1815]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,263,956,979]" captionTargetBox="[151,1436,193,935]" captionTargetId="figure@4.[151,1436,193,935]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Pachylaelaps (Longipachylaelaps) abludens sp. nov., sperm induction structures, female, variant forms. 10 ‒ 11. Normal forms; 12 ‒ 13. Aberrant forms in strongly pressed specimens. Not scaled." httpUri="https://zenodo.org/record/322029/files/figure.png" pageId="17" pageNumber="112">Figs 10–13</figureCitation>
, 
<figureCitation id="134F2A722E17C47DA2E7F8D6FA81674E" box="[1262,1323,1794,1815]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="17" pageNumber="112">49, 50</figureCitation>
)..... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E17C47DA69EF8CBFA36679E" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EF8CBFA3667D5" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF8CBFA3667D5" pageId="17" pageNumber="112">
6. Dorsal setae short: setae J1 (= 25–30 µm) with tips reaching between insertions of setae J1 and J2; setae J4 slightly shorter than setae J5 (J4/J5 0.87–0.96); pilus dentilis conspicuously enlarged, spine-like; tubiform structures of sperm induction system with narrow base and club-like apice (
<figureCitation id="134F2A722E17C47DA446F88EFD3D6736" box="[591,663,1882,1903]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="17" pageNumber="112">Fig. 58</figureCitation>
); smaller species, length of dorsal shield 510–565 µm................................................................................... 
<taxonomicName id="4C744D742E17C47DA5D1F8ACFA3667D5" ID-CoL="8Y7K6" authority="Masan, Ozbek &amp; Fenda, 2016" authorityName="Masan, Ozbek &amp; Fenda" authorityYear="2016" box="[984,1436,1911,1933]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="112" phylum="Arthropoda" rank="species" species="brevipilis" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E17C47DA5D1F8ACFC4E67D4" box="[984,996,1912,1933]" italics="true" pageId="17" pageNumber="112">P</emphasis>
. (
<emphasis id="B900EAE52E17C47DA5F1F8ACFBAF67D4" box="[1016,1029,1912,1933]" italics="true" pageId="17" pageNumber="112">L</emphasis>
.) 
<emphasis id="B900EAE52E17C47DA210F8A3FBD967D4" box="[1049,1139,1911,1933]" italics="true" pageId="17" pageNumber="112">brevipilis</emphasis>
<bibRefCitation id="EFE54B062E17C47DA270F8A3FA3667D5" author="Masan" box="[1145,1436,1911,1932]" pageId="17" pageNumber="112" refString="Masan, P., Ozbek, H. H. &amp; Fenda, P. (2016) Two new species of Pachylaelaps Berlese, 1888 from the Iberian Peninsula, with a key to European species (Acari, Gamasida, Pachylaelapidae). ZooKeys, 603, 71 - 95." type="journal article" year="2016">Mašán, Özbek &amp; Fenďa, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E17C47DA69EF841FA36679E" blockId="17.[151,1436,1266,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF841FA36679E" pageId="17" pageNumber="112">- Dorsal setae long: setae J1 (= 60–95 µm) with tips reaching to or beyond insertions of setae J2; setae J4 at least 1.5 as long as setae J5; pilus dentilis not hypertrophied, slender; larger species, dorsal shield at least 680 µm in length................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E17C47EA69EF81BFA31615D" lastPageId="18" lastPageNumber="113" pageId="17" pageNumber="112">
<paragraph id="8BCB36F72E17C47DA69EF81BFA3667BD" blockId="17.[151,1436,1266,2020]" box="[151,1436,1999,2020]" pageId="17" pageNumber="112">
<keyLead id="308595C22E17C47DA69EF81BFA3667BD" box="[151,1436,1999,2020]" pageId="17" pageNumber="112">7. Dorsal setae J5 markedly shortened, about 5–6 times shorter than setae J4; tubiform structures os sperm induction system reg-</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA6E8FF43FA31615D" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
ularly sclerotised; pilus dentilis minute, with upright position; length of dorsal shield 770–890 µm.............................................................................................. 
<taxonomicName id="4C744D742E14C47EA24BFF61FA366093" ID-CoL="8Y7KT" authority="Koroleva, 1977" authorityName="Koroleva" authorityYear="1977" box="[1090,1436,180,202]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="sublongisetis" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA24BFF61FBE46093" box="[1090,1102,181,202]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA26BFF61FBC56093" box="[1122,1135,181,202]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA28DFF60FAAB6093" box="[1156,1281,180,202]" italics="true" pageId="18" pageNumber="113">sublongisetis</emphasis>
<bibRefCitation id="EFE54B062E14C47EA30EFF61FA366093" author="Koroleva" box="[1287,1436,181,202]" pageId="18" pageNumber="113" refString="Koroleva, E. V. (1977) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites, Mesostigmata. Nauka Press, Leningrad, pp. 411 - 483. [in Russian]" type="book chapter" year="1977">Koroleva, 1977</bibRefCitation>
</taxonomicName>
<keyLead id="308595C22E14C47EA69EFF06FA31615D" pageId="18" pageNumber="113">- Setae J5 less shortened, about 1.4–2.2 times shorter than setae J4; basal or medial portion of tubiform structures unsclerotised, hyaline; pilus dentilis larger, curved and directed backward................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFED9FA366105" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFED9FA366166" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFED9FA366166" pageId="18" pageNumber="113">
8. Tubiform structures of sperm induction system with less sclerotised medial portion and well sclerotised narrow base (
<figureCitation id="134F2A722E14C47EA341FED9FA27617B" box="[1352,1421,269,290]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="18" pageNumber="113">Fig. 59</figureCitation>
); length of dorsal shield 680–805 µm.............................................. 
<taxonomicName id="4C744D742E14C47EA26EFEFEFA366166" ID-CoL="8Y7KL" authority="Willmann, 1938" authorityName="Willmann" authorityYear="1938" box="[1127,1436,297,319]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="longulus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA26EFEFEFBD96166" box="[1127,1139,298,319]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA28EFEFEFB3E6166" box="[1159,1172,298,319]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2A1FEFDFB516166" box="[1192,1275,297,319]" italics="true" pageId="18" pageNumber="113">longulus</emphasis>
Willmann, 1938
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<paragraph id="8BCB36F72E14C47EA69EFE93FA366105" blockId="18.[151,1437,151,1845]" box="[151,1436,327,348]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFE93FA366105" box="[151,1436,327,348]" pageId="18" pageNumber="113">
- Tubiform structures with less sclerotised and slightly expanded basal portion (
<figureCitation id="134F2A722E14C47EA5C4FE93FB966105" box="[973,1084,327,348]" captionStart="FIGURES 23 – 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 – 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="18" pageNumber="113">Figs 23–26</figureCitation>
, 
<figureCitation id="134F2A722E14C47EA24FFE93FB296105" box="[1094,1155,327,348]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="18" pageNumber="113">51, 52</figureCitation>
).................... 9
</keyLead>
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<paragraph id="8BCB36F72E14C47EA69EFEB1FA3661EC" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFEB1FA3661EC" pageId="18" pageNumber="113">
9. Tubiform structures of sperm induction system larger, as in 
<figureCitation id="134F2A722E14C47EA531FEB1FC036123" box="[824,937,357,378]" captionStart="FIGURES 23 – 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 – 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="18" pageNumber="113">Figs 23–26</figureCitation>
; genitiventral shield subequal in length and width (289–318 µm long, 289–310 µm wide, L/ 
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–1.05); anterior sections of peritremes longer, with tips reaching close to dorsal setae z1; length of dorsal shield 780–925 µm....................................... 
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<emphasis id="B900EAE52E14C47EA286FE74FB3161EC" box="[1167,1179,416,437]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA2A6FE74FB1661EC" box="[1199,1212,416,437]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2D8FE4BFAE361EC" box="[1233,1353,415,437]" italics="true" pageId="18" pageNumber="113">bergomensis</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E14C47EA346FE4BFA3661EC" bold="true" box="[1359,1436,415,437]" pageId="18" pageNumber="113">
<taxonomicNameLabel id="A233579E2E14C47EA346FE4BFA3661EC" box="[1359,1436,415,437]" pageId="18" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFE69FA366254" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFE69FA366254" pageId="18" pageNumber="113">
- Tubiform structures smaller, as in 
<figureCitation id="134F2A722E14C47EA438FE69FD09618B" box="[561,675,445,466]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="18" pageNumber="113">Figs 51, 52</figureCitation>
; genitiventral shield oblong (246–284 µm long, 203–225 µm wide, L/ 
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– 1.28); anterior sections of peritremes shorter, with tips reaching between setae z1 and z2; length of dorsal shield 665–800 µm.................................................................................. 
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<emphasis id="B900EAE52E14C47EA2B4FE2CFB636254" box="[1213,1225,504,525]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA2D4FE2CFB406254" box="[1245,1258,504,525]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2F7FE2CFAE36254" box="[1278,1353,504,525]" italics="true" pageId="18" pageNumber="113">pantinii</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E14C47EA346FE23FA366254" bold="true" box="[1359,1436,503,525]" pageId="18" pageNumber="113">
<taxonomicNameLabel id="A233579E2E14C47EA346FE23FA366254" box="[1359,1436,503,525]" pageId="18" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
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<keyLead id="308595C22E14C47EA69EFDC1FA316273" box="[151,1435,533,554]" pageId="18" pageNumber="113">
10. Tubiform structures of sperm induction system conspicuously elongated, with 1–3 bends (
<figureCitation id="134F2A722E14C47EA249FDC1FB076273" box="[1088,1197,533,554]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,263,956,979]" captionTargetBox="[151,1436,193,935]" captionTargetId="figure@4.[151,1436,193,935]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Pachylaelaps (Longipachylaelaps) abludens sp. nov., sperm induction structures, female, variant forms. 10 ‒ 11. Normal forms; 12 ‒ 13. Aberrant forms in strongly pressed specimens. Not scaled." httpUri="https://zenodo.org/record/322029/files/figure.png" pageId="18" pageNumber="113">Figs 10–13</figureCitation>
)............... 11
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFDE6FA36621E" blockId="18.[151,1437,151,1845]" box="[151,1436,562,583]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFDE6FA36621E" box="[151,1436,562,583]" pageId="18" pageNumber="113">
- Tubiform structures shorter, straight or regularly curved (
<figureCitation id="134F2A722E14C47EA50CFDE6FCD8621E" box="[773,882,562,583]" captionStart="FIGURES 49 – 52" captionStartId="15.[151,263,982,1005]" captionTargetBox="[151,1436,193,961]" captionTargetId="figure@15.[151,1436,193,961]" captionTargetPageId="15" captionText="FIGURES 49 – 52. Sperm induction structures, females, variant forms. 49 ‒ 50. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. (49. Normal form; 50. Aberrant form in strongly pressed specimen); 51 ‒ 52. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. (51. Normal form; 52. Aberrant form in strongly pressed specimen). Not scaled." httpUri="https://zenodo.org/record/322038/files/figure.png" pageId="18" pageNumber="113">Figs 49, 50</figureCitation>
).......................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFD9BFA3762AE" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFD9BFA3662C6" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFD9BFA3662C6" pageId="18" pageNumber="113">
11. Tubiform structures of sperm induction system helically convoluted, with 2–3 bends in medial and distal portions; dorsal setae J4 and J5 subequal in length; soft integument with 14 pairs of setae (six pairs of r-R setae and eight pairs of Jv-Zv setae); length of dorsal shield 836 µm............................................... 
<taxonomicName id="4C744D742E14C47EA24DFD5EFA3662C6" ID-CoL="8XTZK" authority="Moraza &amp; Pena, 2005" authorityName="Moraza &amp; Pena" authorityYear="2005" box="[1092,1436,649,671]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="silviae" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA24DFD5EFBFA62C6" box="[1092,1104,650,671]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA26DFD5EFBDB62C6" box="[1124,1137,650,671]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA28CFD5DFB6962C6" box="[1157,1219,649,671]" italics="true" pageId="18" pageNumber="113">silviae</emphasis>
Moraza &amp; Peña, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFD73FA3762AE" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFD73FA3762AE" pageId="18" pageNumber="113">
- Tubiform structures with one subterminal bend, as in 
<figureCitation id="134F2A722E14C47EA4E5FD73FCF662E5" box="[748,860,679,700]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,263,956,979]" captionTargetBox="[151,1436,193,935]" captionTargetId="figure@4.[151,1436,193,935]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Pachylaelaps (Longipachylaelaps) abludens sp. nov., sperm induction structures, female, variant forms. 10 ‒ 11. Normal forms; 12 ‒ 13. Aberrant forms in strongly pressed specimens. Not scaled." httpUri="https://zenodo.org/record/322029/files/figure.png" pageId="18" pageNumber="113">Figs 10–13</figureCitation>
; setae J4 as long as setae J5 (J4/J5 1.43–1.98); soft integument with 16 pairs of setae (seven pairs of r-R setae and nine pairs of Jv-Zv setae); length of dorsal shield 785–845 µm...................................................................................... 
<taxonomicName id="4C744D742E14C47EA2BDFD36FAE162AE" ID-CoL="8Y7JZ" box="[1204,1355,737,759]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="abludens" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA2BDFD36FB6A62AE" box="[1204,1216,738,759]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA2DDFD36FB4B62AE" box="[1236,1249,738,759]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2FCFD35FAE162AE" box="[1269,1355,737,759]" italics="true" pageId="18" pageNumber="113">abludens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233579E2E14C47EA35BFD36FA3762AE" box="[1362,1437,738,759]" pageId="18" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFD2BFA316335" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFD2BFA36636B" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFD2BFA36636B" pageId="18" pageNumber="113">
12. Dorsal setae J5 shortened (17.5 µm long), more than three times shorter than setae J4; tubiform structures of sperm induction system terminally truncate; length of dorsal shield 870 µm............................ 
<taxonomicName id="4C744D742E14C47EA278FCC9FA36636B" ID-CoL="8Y7KS" authority="Berlese, 1920" authorityName="Berlese" authorityYear="1920" box="[1137,1436,796,818]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="squamifer" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA278FCC9FBD7636B" box="[1137,1149,797,818]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA298FCC9FB34636B" box="[1169,1182,797,818]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2BBFCC8FAB8636B" box="[1202,1298,796,818]" italics="true" pageId="18" pageNumber="113">squamifer</emphasis>
Berlese, 1920
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFCEEFA316335" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFCEEFA316335" pageId="18" pageNumber="113">- Setae J5 normal in length (± 40 µm long), never more than two times shorter than setae J4; tubiform structures terminally rounded........................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFCA1FA3163BB" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFCA1FA3663F1" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFCA1FA3663F1" pageId="18" pageNumber="113">
13. Dorsal setae shorter: setae J1 with tips not reaching insertions of following setae J2; setae J5 shorter than setae J4 (J4/J5 1.45– 1.8); length of dorsal shield 830–865 µm............................................. 
<taxonomicName id="4C744D742E14C47EA29FFC46FAE363FE" ID-CoL="8Y7KM" box="[1174,1353,913,935]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="marcovallei" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA29FFC46FB0863FE" box="[1174,1186,914,935]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA2BFFC46FB6963FE" box="[1206,1219,914,935]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2DEFC45FAE363FE" box="[1239,1353,913,935]" italics="true" pageId="18" pageNumber="113">marcovallei</emphasis>
</taxonomicName>
<emphasis id="B900EAE52E14C47EA346FC46FA3663F1" bold="true" box="[1359,1436,914,936]" pageId="18" pageNumber="113">
<taxonomicNameLabel id="A233579E2E14C47EA346FC46FA3663F1" box="[1359,1436,914,936]" pageId="18" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFC7BFA3163BB" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFC7BFA3163BB" pageId="18" pageNumber="113">
- Dorsal setae longer: setae J1 with tips reaching beyond insertions of setae J2; setae J4 and J5 subequal in length (J5 less than 1.4 times shorter than setae J5); length of dorsal shield unknown............. 
<taxonomicName id="4C744D742E14C47EA5FBFC19FA3163BB" ID-CoL="8Y7KR" authority="Hirschmann &amp; Krauss, 1965" authorityName="Hirschmann &amp; Krauss" authorityYear="1965" box="[1010,1435,972,994]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="pulsator" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA5FBFC19FC5463BB" box="[1010,1022,973,994]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA21BFC19FBB563BB" box="[1042,1055,973,994]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA23DFC18FB2E63BB" box="[1076,1156,972,994]" italics="true" pageId="18" pageNumber="113">pulsator</emphasis>
Hirschmann &amp; Krauss, 1965
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFC3EFA3164CB" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFC3EFA316463" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFC3EFA316463" pageId="18" pageNumber="113">
14. Sperm induction system with specific sickle-shaped structures (
<figureCitation id="134F2A722E14C47EA549FC3EFC2263A6" box="[832,904,1002,1023]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="18" pageNumber="113">Fig. 60</figureCitation>
); pilus dentilis relatively robust, spine-like and directed backward; movable cheliceral digit with simple distal hook; genitiventral shield relatively narrower (length/width 1.14–1.27); length of dorsal shield 745–840 µm............................................... 
<taxonomicName id="4C744D742E14C47EA274FBF1FA316463" ID-CoL="8Y7KB" authority="Masan, 2007" authorityName="Masan" authorityYear="2007" box="[1149,1435,1060,1082]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="distinctus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA274FBF1FB236463" box="[1149,1161,1061,1082]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA295FBF1FB036463" box="[1180,1193,1061,1082]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2B7FBF0FAB06463" box="[1214,1306,1060,1082]" italics="true" pageId="18" pageNumber="113">distinctus</emphasis>
Mašán, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFB96FA3164CB" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFB96FA3164CB" pageId="18" pageNumber="113">- Sperm induction system sacculate: saccules with short tubiform opening slightly protuberant above enlarged base; pilus dentilis minute, with upright position; movable cheliceral digit with bifid distal hook; genitiventral shield relatively wider (length/ width 0.92–1.12)................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFB4EFA31651B" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFB4EFA3164B3" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFB4EFA3164B3" pageId="18" pageNumber="113">
15. Sperm ductus inside saccules relatively shorter, straight and directed to anterior margin of coxa IV; base of saccules abutting the coxa IV (
<figureCitation id="134F2A722E14C47EA76BFB63FE016495" box="[354,427,1207,1228]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="18" pageNumber="113">Fig. 61</figureCitation>
); dorsal setae J 
<date id="FFCA10372E14C47EA448FB63FD256495" box="[577,655,1207,1228]" pageId="18" pageNumber="113" value="1939-05-30">5 30–39</date>
µm long, about 2–3.5 times shorter than setae J4; genitiventral shield relatively narrower (length/width 1.03–1.12); length of dorsal shield 745–885 µm.................... 
<taxonomicName id="4C744D742E14C47EA282FB01FA3164B3" ID-CoL="8Y7KX" authority="Masan, 2007" authorityName="Masan" authorityYear="2007" box="[1163,1435,1237,1258]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="vicarius" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA282FB01FB3D64B3" box="[1163,1175,1237,1258]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA2A2FB01FB1264B3" box="[1195,1208,1237,1258]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2C5FB01FAB364B3" box="[1228,1305,1237,1258]" italics="true" pageId="18" pageNumber="113">vicarius</emphasis>
Mašán, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFB26FA31651B" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFB26FA31651B" pageId="18" pageNumber="113">
- Sperm ductus inside saccules relatively longer, slightly curved and directed between coxae III and IV; base of saccules slightly widened, abutting the coxae III and IV; setae J 
<date id="FFCA10372E14C47EA497FADBFD41657D" box="[670,747,1295,1316]" pageId="18" pageNumber="113" value="1920-05-25">5 20–25</date>
µm long, about 5–7 times shorter than setae J4; genitiventral shield relatively wider (length/width 0.9–0.95); length of dorsal shield 940–1,050 µm........... 
<taxonomicName id="4C744D742E14C47EA246FAF9FA31651B" ID-CoL="8Y7K5" authority="Koroleva, 1978" authorityName="Koroleva" authorityYear="1978" box="[1103,1435,1324,1346]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="bocharovae" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA246FAF9FBF1651B" box="[1103,1115,1325,1346]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA266FAF9FBD6651B" box="[1135,1148,1325,1346]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA299FAF8FAAA651B" box="[1168,1280,1324,1346]" italics="true" pageId="18" pageNumber="113">bocharovae</emphasis>
Koroleva, 1978
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFA9EFA366525" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFA9EFA366506" blockId="18.[151,1437,151,1845]" box="[151,1436,1354,1375]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFA9EFA366506" box="[151,1436,1354,1375]" pageId="18" pageNumber="113">16. Tubiform structures of sperm induction system elongated (with more or less adjacent tips), straight or slightly curved.... 17</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFAB3FA366525" blockId="18.[151,1437,151,1845]" box="[151,1436,1383,1404]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFAB3FA366525" box="[151,1436,1383,1404]" pageId="18" pageNumber="113">- Tubiform structures shortened (with well separated tips) or not detectable...................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EFA51FA366675" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EFA51FA31658D" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFA51FA31658D" pageId="18" pageNumber="113">
17. Tubiform structures thin and long, worm-like; opposite margins of genitiventral and anal shield straight and markedly separated; dorsal setae J4 and J5 short and subequal in length; epistome with distal projection narrow and bifurcate apically (often with small denticle between lateral cusps); length of dorsal shield 910 µm............... 
<taxonomicName id="4C744D742E14C47EA27BFA14FA31658D" ID-CoL="8Y7K4" authority="Berlese, 1920" authorityName="Berlese" authorityYear="1920" box="[1138,1435,1471,1493]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="bifurciger" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA27BFA14FBD4658C" box="[1138,1150,1472,1493]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA29BFA14FB35658C" box="[1170,1183,1472,1493]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2BAFA6BFAB8658C" box="[1203,1298,1471,1493]" italics="true" pageId="18" pageNumber="113">bifurciger</emphasis>
Berlese, 1920
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EFA09FA366675" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EFA09FA366675" pageId="18" pageNumber="113">
- Tubiform structures broadened, with slightly club-like tip; opposite margins of genitiventral and anal shield undulate and closely abutting each other; setae J 
<date id="FFCA10372E14C47EA425FA2EFDD36656" box="[556,633,1530,1551]" pageId="18" pageNumber="113" value="1933-05-30">5 30–33</date>
µm long, about 4–4.5 times shorter than setae J4; distal projection of epistome wide and densely crenulated anteriorly; length of dorsal shield 780–840 µm............ 
<taxonomicName id="4C744D742E14C47EA225F9CCFA366675" ID-CoL="8Y7KW" authority="Evans &amp; Hyatt, 1956" authorityName="Evans &amp; Hyatt" authorityYear="1956" box="[1068,1436,1559,1581]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="undulatus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA225F9CCFB926674" box="[1068,1080,1560,1581]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA245F9CCFBF36674" box="[1100,1113,1560,1581]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA264F9C3FB616674" box="[1133,1227,1559,1581]" italics="true" pageId="18" pageNumber="113">undulatus</emphasis>
<bibRefCitation id="EFE54B062E14C47EA2D8F9C3FA366675" author="Evans" box="[1233,1436,1559,1580]" pageId="18" pageNumber="113" refString="Evans, G. O. &amp; Hyatt, K. H. (1956) British mites of the genus Pachylaelaps Berlese (Gamasina - Pachylaelaptidae). Entomologist's Monthly Magazine, 92, 118 - 129." type="journal article" year="1956">Evans &amp; Hyatt, 1956</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EF9E1FA3166DD" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EF9E1FA31663E" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EF9E1FA31663E" pageId="18" pageNumber="113">
18. Tubiform structures short, broad, conical and delicately striated transversally (
<figureCitation id="134F2A722E14C47EA5C7F9E1FBBC6613" box="[974,1046,1589,1610]" captionStart="FIGURES 57 – 62" captionStartId="20.[151,263,1370,1393]" captionTargetBox="[151,1436,193,1349]" captionTargetId="figure@20.[151,1436,193,1349]" captionTargetPageId="20" captionText="FIGURES 57 – 62. Sperm induction structures, females. 57. Pachylaelaps (Longipachylaelaps) longisetis; 58. Pachylaelaps (Longipachylaelaps) brevipilis; 59. Pachylaelaps (Longipachylaelaps) longulus; 60. Pachylaelaps (Longipachylaelaps) distinctus; 61. Pachylaelaps (Longipachylaelaps) vicarius; 62. Pachylaelaps (Longipachylaelaps) carpathicus. Not scaled." httpUri="https://zenodo.org/record/322040/files/figure.png" pageId="18" pageNumber="113">Fig. 62</figureCitation>
); length of dorsal shield 685–835 µm............................................................................. 
<taxonomicName id="4C744D742E14C47EA26EF986FA31663E" ID-CoL="8Y7K7" authority="Masan, 2007" authorityName="Masan" authorityYear="2007" box="[1127,1435,1617,1639]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="carpathicus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA26EF986FBD9663E" box="[1127,1139,1618,1639]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA28EF986FB3E663E" box="[1159,1172,1618,1639]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2A0F985FAB0663E" box="[1193,1306,1617,1639]" italics="true" pageId="18" pageNumber="113">carpathicus</emphasis>
Mašán, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EF9BBFA3166DD" blockId="18.[151,1437,151,1845]" box="[151,1435,1647,1668]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EF9BBFA3166DD" box="[151,1435,1647,1668]" pageId="18" pageNumber="113">- Tubiform structures not detectable, absent................................................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30802E522E14C47EA69EF959FA31676D" pageId="18" pageNumber="113">
<paragraph id="8BCB36F72E14C47EA69EF959FA316685" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EF959FA316685" pageId="18" pageNumber="113">
19. Dorsal setae J5 shortened (15–25 µm long), more than three times shorter than setae J4 (82–105 µm long); genitiventral shield 240–300 µm wide, relatively narrower (L/W 0.97–1.11); soft integument with 16 pairs of setae (seven pairs of r-R setae and nine pairs of Jv-Zv setae); smaller species, length of dorsal shield 740–930 µm............ 
<taxonomicName id="4C744D742E14C47EA27DF91CFA316685" ID-CoL="8Y7KQ" authority="Masan, 2007" authorityName="Masan" authorityYear="2007" box="[1140,1435,1735,1757]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="perlucidus" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA27DF91CFB2A6684" box="[1140,1152,1736,1757]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA29AF91CFB0A6684" box="[1171,1184,1736,1757]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA2BCF913FAB06684" box="[1205,1306,1735,1757]" italics="true" pageId="18" pageNumber="113">perlucidus</emphasis>
Mašán, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCB36F72E14C47EA69EF931FA31676D" blockId="18.[151,1437,151,1845]" pageId="18" pageNumber="113">
<keyLead id="308595C22E14C47EA69EF931FA31676D" pageId="18" pageNumber="113">
- Dorsal setae J5 normal (50–66 µm long), less than 1.5 times shorter than setae J4 (60–78 µm long); genitiventral shield 375– 450 µm wide, relatively wider (L/W 0.86–0.97); soft integument with 14 pairs of setae (six pairs of r-R setae and eight pairs of Jv-Zv setae); larger species, length of dorsal shield 990–1,085 µm......... 
<taxonomicName id="4C744D742E14C47EA5E8F8F4FA31676D" ID-CoL="8Y7KG" authority="Hirschmann &amp; Krauss, 1965" authorityName="Hirschmann &amp; Krauss" authorityYear="1965" box="[993,1435,1823,1845]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="granulifer" subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E14C47EA5E8F8F4FC47676C" box="[993,1005,1824,1845]" italics="true" pageId="18" pageNumber="113">P</emphasis>
. (
<emphasis id="B900EAE52E14C47EA208F8F4FBA4676C" box="[1025,1038,1824,1845]" italics="true" pageId="18" pageNumber="113">L</emphasis>
.) 
<emphasis id="B900EAE52E14C47EA22BF8CBFB29676C" box="[1058,1155,1823,1845]" italics="true" pageId="18" pageNumber="113">granulifer</emphasis>
Hirschmann &amp; Krauss, 1965
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>