treatments-xml/data/03/DD/87/03DD87E12E03C46AA69EFEF3FE1A62F1.xml
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<document id="907DCC094875AAE42BBEF49C01CBE97E" ID="10.11646/zootaxa.4236.1.5SLASH10255" ID-CLB-Dataset="36476" ID-DOI="10.11646/zootaxa.4236.1.5" ID-GBIF-Dataset="9b07325e-60db-405c-97a2-831f68682f6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="322026" ID-ZooBank="04D3145C-1C8A-4AE7-968F-7F307E1E732A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487754882402" checkinUser="plazi" docAuthor="Mašán, Peter" docDate="2017" docId="03DD87E12E03C46AA69EFEF3FE1A62F1" docLanguage="en" docName="zootaxa.4236.1.5SLASH10255.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachylaelaps (Longipachylaelaps) bergomensis Mašán, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="101" masterDocId="FFE4FF992E06C46CA609FFD4FFAA6059" masterDocTitle="Four new species of the subgenus Pachylaelaps (Longipachylaelaps) from Italy, with a new identification key to the known species (Acari: Pachylaelapidae)" masterLastPageNumber="117" masterPageNumber="95" pageNumber="100" updateTime="1698437172724" updateUser="ExternalLinkService">
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<mods:title id="F04E8832B6B05882AC9696670FEA0B43">Four new species of the subgenus Pachylaelaps (Longipachylaelaps) from Italy, with a new identification key to the known species (Acari: Pachylaelapidae)</mods:title>
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<mods:namePart id="A67882C3AC497F82522D5D49CC16019C">Mašán, Peter</mods:namePart>
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<mods:date id="D1956A8F56AF362C7B55D857E61FBA2A">2017</mods:date>
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<treatment id="03DD87E12E03C46AA69EFEF3FE1A62F1" ID-DOI="http://doi.org/10.5281/zenodo.5694784" ID-GBIF-Taxon="127696758" ID-Zenodo-Dep="5694784" LSID="urn:lsid:plazi:treatment:03DD87E12E03C46AA69EFEF3FE1A62F1" httpUri="http://treatment.plazi.org/id/03DD87E12E03C46AA69EFEF3FE1A62F1" lastPageId="6" lastPageNumber="101" pageId="5" pageNumber="100">
<subSubSection id="C36E657C2E03C469A69EFEF3FED6613A" pageId="5" pageNumber="100" type="nomenclature">
<paragraph id="8BCB36F72E03C469A69EFEF3FC93611B" blockId="5.[151,825,295,355]" box="[151,825,295,322]" pageId="5" pageNumber="100">
<heading id="D083819B2E03C469A69EFEF3FC93611B" bold="true" box="[151,825,295,322]" fontSize="11" level="1" pageId="5" pageNumber="100" reason="1">
<emphasis id="B900EAE52E03C469A69EFEF3FC93611B" bold="true" box="[151,825,295,322]" pageId="5" pageNumber="100">
<taxonomicName id="4C744D742E03C469A69EFEF3FD796118" ID-CoL="8Y7K3" box="[151,723,295,322]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="100" phylum="Arthropoda" rank="species" species="bergomensis" status="sp. nov." subGenus="Longipachylaelaps">
<emphasis id="B900EAE52E03C469A69EFEF3FE9D6118" bold="true" box="[151,311,295,321]" italics="true" pageId="5" pageNumber="100">Pachylaelaps</emphasis>
(
<emphasis id="B900EAE52E03C469A74EFEF3FD816118" bold="true" box="[327,555,295,321]" italics="true" pageId="5" pageNumber="100">Longipachylaelaps</emphasis>
)
<emphasis id="B900EAE52E03C469A435FEF3FD796118" bold="true" box="[572,723,295,321]" italics="true" pageId="5" pageNumber="100">bergomensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233579E2E03C469A4D2FEFCFC93611B" box="[731,825,296,322]" pageId="5" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB36F72E03C469A69EFE9EFED6613A" blockId="5.[151,825,295,355]" box="[151,380,330,355]" pageId="5" pageNumber="100">
(
<figureCitation id="134F2A722E03C469A696FE9EFF5F613A" box="[159,245,330,355]" captionStart="FIGURES 14 15" captionStartId="6.[151,263,1835,1858]" captionTargetBox="[158,1425,743,1809]" captionTargetId="figure@6.[151,1436,735,1814]" captionTargetPageId="6" captionText="FIGURES 14 15. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322030/files/figure.png" pageId="5" pageNumber="100">Figs 14</figureCitation>
¯26, 37, 54)
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E03C469A69EFE46FE186262" pageId="5" pageNumber="100" type="description">
<paragraph id="8BCB36F72E03C469A69EFE46FE186262" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<materialsCitation id="3B1C3CAA2E03C469A69EFE46FC3B624E" ID-GBIF-Occurrence="1434656173" collectingDate="2015-05-13" collectorName="Bosco di Astino &amp; P. Masan" country="Italy" county="Bosco di Astino" elevation="265" location="Specimens" municipality="Astino Valley" pageId="5" pageNumber="100" specimenCount="1" typeStatus="holotype">
<emphasis id="B900EAE52E03C469A69EFE46FE3261F2" bold="true" box="[151,408,402,427]" pageId="5" pageNumber="100">
<location id="8EAB602C2E03C469A69EFE46FEBF61F2" LSID="urn:lsid:plazi:treatment:03DD87E12E03C46AA69EFEF3FE1A62F1:8EAB602C2E03C469A69EFE46FEBF61F2" box="[151,277,402,427]" country="Italy" county="Bosco di Astino" municipality="Astino Valley" name="Specimens" pageId="5" pageNumber="100">Specimens</location>
examined.
</emphasis>
<typeStatus id="54CF88552E03C469A7A9FE46FDA261F2" box="[416,520,402,427]" pageId="5" pageNumber="100" type="holotype">Holotype</typeStatus>
female: North
<collectingCountry id="F36376672E03C469A4BEFE46FD4161F2" box="[695,747,402,427]" name="Italy" pageId="5" pageNumber="100">Italy</collectingCountry>
,
<location id="8EAB602C2E03C469A4F1FE47FC3D61F2" LSID="urn:lsid:plazi:treatment:03DD87E12E03C46AA69EFEF3FE1A62F1:8EAB602C2E03C469A4F1FE47FC3D61F2" box="[760,919,402,427]" country="Italy" county="Bosco di Astino" municipality="Astino Valley" name="Bergamo City" pageId="5" pageNumber="100">Bergamo City</location>
,
<collectingMunicipality id="6BAFAC8D2E03C469A5ADFE46FBEA61F2" box="[932,1088,402,427]" pageId="5" pageNumber="100">Astino Valley</collectingMunicipality>
,
<collectingCounty id="62AA4E7B2E03C469A245FE47FAAC61F2" box="[1100,1286,402,427]" pageId="5" pageNumber="100">
<collectorName id="268153212E03C469A245FE47FAAC61F2" box="[1100,1286,402,427]" pageId="5" pageNumber="100">Bosco di Astino</collectorName>
</collectingCounty>
, broadleaved deciduous forest (with
<taxonomicName id="4C744D742E03C469A7BAFE63FDF46197" box="[435,606,438,462]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="robur">
<emphasis id="B900EAE52E03C469A7BAFE63FDF46197" box="[435,606,438,462]" italics="true" pageId="5" pageNumber="100">Quercus robur</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A47CFE63FD5C6197" box="[629,758,439,462]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="petraea">
<emphasis id="B900EAE52E03C469A47CFE63FD236197" box="[629,649,439,462]" italics="true" pageId="5" pageNumber="100">Q</emphasis>
.
<emphasis id="B900EAE52E03C469A4A9FE63FD5C6197" box="[672,758,439,462]" italics="true" pageId="5" pageNumber="100">petraea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A505FE63FCD36197" box="[780,889,439,462]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="cerris">
<emphasis id="B900EAE52E03C469A505FE63FC8A6197" box="[780,800,439,462]" italics="true" pageId="5" pageNumber="100">Q</emphasis>
.
<emphasis id="B900EAE52E03C469A531FE63FCD36197" box="[824,889,439,462]" italics="true" pageId="5" pageNumber="100">cerris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A599FE63FBE66197" box="[912,1100,439,462]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="B900EAE52E03C469A599FE63FBE66197" box="[912,1100,439,462]" italics="true" pageId="5" pageNumber="100">Castanea sativa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A26AFE63FA876197" box="[1123,1325,438,462]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="betulus">
<emphasis id="B900EAE52E03C469A26AFE63FA876197" box="[1123,1325,438,462]" italics="true" pageId="5" pageNumber="100">Carpinus betulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A34AFE62FE9B61AA" class="Trematoda" family="Hemiuridae" genus="Robinia" kingdom="Animalia" order="Plagiorchiida" pageId="5" pageNumber="100" phylum="Platyhelminthes" rank="species" species="pseudoacacia">
<emphasis id="B900EAE52E03C469A34AFE62FE9B61AA" italics="true" pageId="5" pageNumber="100">Robinia pseudoacacia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A737FE0FFE2D61AA" box="[318,391,475,499]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="B900EAE52E03C469A737FE0FFE2D61AA" box="[318,391,475,499]" italics="true" pageId="5" pageNumber="100">Betula</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C744D742E03C469A7B5FE0FFD8B61AB" box="[444,545,475,498]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="B900EAE52E03C469A7B5FE0FFD8B61AB" box="[444,545,475,498]" italics="true" pageId="5" pageNumber="100">Fraxinus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C744D742E03C469A45FFE0FFCBC61AA" box="[598,790,475,499]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="avellana">
<emphasis id="B900EAE52E03C469A45FFE0FFCBC61AA" box="[598,790,475,499]" italics="true" pageId="5" pageNumber="100">Corylus avellana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A52DFE0FFC3361AB" box="[804,921,475,498]" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="B900EAE52E03C469A52DFE0FFC3361AB" box="[804,921,475,498]" italics="true" pageId="5" pageNumber="100">Crataegus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C744D742E03C469A5C4FE0FFB3D61AA" box="[973,1175,475,499]" class="Magnoliopsida" family="Adoxaceae" genus="Viburnum" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="lantana">
<emphasis id="B900EAE52E03C469A5C4FE0FFB3D61AA" box="[973,1175,475,499]" italics="true" pageId="5" pageNumber="100">Viburnum lantana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C744D742E03C469A2ACFE0FFA8161AB" box="[1189,1323,475,498]" class="Magnoliopsida" family="Cornaceae" genus="Cornus" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="mas">
<emphasis id="B900EAE52E03C469A2ACFE0FFA8161AB" box="[1189,1323,475,498]" italics="true" pageId="5" pageNumber="100">Cornus mas</emphasis>
</taxonomicName>
, ao.), leaf litter and soil detritus, altitude
<quantity id="4C8C9B122E03C469A7FAFE2AFD97624F" box="[499,573,510,535]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="5" pageNumber="100" unit="m" value="265.0">
<elevation id="0059D1C42E03C469A7FAFE2AFD97624F" box="[499,573,510,535]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="5" pageNumber="100" unit="m" value="265.0">265 m</elevation>
</quantity>
,
<date id="FFCA10372E03C469A440FE2BFD49624E" box="[585,739,510,535]" pageId="5" pageNumber="100" value="2015-05-13">
<collectingDate id="EF8EE9DF2E03C469A440FE2BFD49624E" box="[585,739,510,535]" pageId="5" pageNumber="100" value="2015-05-13">13 May 2015</collectingDate>
</date>
, coll.
<collectorName id="268153212E03C469A521FE2BFC3B624E" box="[808,913,511,535]" pageId="5" pageNumber="100">P. Mašán</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1C3CAA2E03C469A594FE2BFE046262" ID-GBIF-Occurrence="1434656172" collectingDate="2015-05-13" collectorName="Bosco di Astino &amp; P. Masan" country="Italy" county="Bosco di Astino" elevation="265" location="Specimens" municipality="Astino Valley" pageId="5" pageNumber="100" specimenCount="40" specimenCount-female="19" specimenCount-male="21" typeStatus="paratype">
<typeStatus id="54CF88552E03C469A594FE2BFBA4624E" box="[925,1038,511,535]" pageId="5" pageNumber="100" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D72FD7E2E03C469A210FE2BFB3E624E" box="[1049,1172,510,535]" pageId="5" pageNumber="100" type="female">19 females</specimenCount>
and
<specimenCount id="9D72FD7E2E03C469A2C5FE2BFA9E624E" box="[1228,1332,510,535]" pageId="5" pageNumber="100" type="male">21 males</specimenCount>
, with the same data as in
<typeStatus id="54CF88552E03C469A741FDF6FE046262" box="[328,430,546,571]" pageId="5" pageNumber="100" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E03C469A6CEFD92FBC6634A" pageId="5" pageNumber="100" type="diagnosis">
<paragraph id="8BCB36F72E03C469A6CEFD92FBC6634A" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFD92FEEB6206" bold="true" box="[199,321,582,607]" pageId="5" pageNumber="100">Diagnosis.</emphasis>
In female, tubiform structures of sperm induction system relatively large, with apices adjacent but separated, moderately narrowed toward apices, vase-like (
<figureCitation id="134F2A722E03C469A55DFDBEFC1962DA" box="[852,947,618,643]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 16</figureCitation>
,
<figureCitation id="134F2A722E03C469A5C1FDBEFBB862DA" box="[968,1042,618,643]" captionStart="FIGURES 23 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="5" pageNumber="100">2326</figureCitation>
). Male spermatodactyl expanded submedially, leaf-like, without distinct angustate subdistal or apical portion, less than double the length of the movable digit (
<figureCitation id="134F2A722E03C469A747FD66FE016292" box="[334,427,690,715]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 20</figureCitation>
,
<figureCitation id="134F2A722E03C469A7B7FD66FE746292" box="[446,478,690,715]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="5" pageNumber="100">54</figureCitation>
). Palptibia with two divergent petal-like projections: antiaxial projection longer, widened medially, rounded distally, with small subapical incision, and small lamellar prominence at its inner base; paraxial projection shorter, subtriangular, and tapered toward obtuse apex (
<figureCitation id="134F2A722E03C469A5D4FD2EFB9E634A" box="[989,1076,762,787]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 21</figureCitation>
,
<figureCitation id="134F2A722E03C469A249FD2FFBCA634A" box="[1088,1120,763,787]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="5" pageNumber="100">37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E03C46AA6CEFCCAFF5161AC" lastPageId="6" lastPageNumber="101" pageId="5" pageNumber="100" type="description">
<paragraph id="8BCB36F72E03C469A6CEFCCAFE71636E" blockId="5.[151,1437,402,2011]" box="[199,475,798,823]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFCCAFE71636E" bold="true" box="[199,475,798,823]" pageId="5" pageNumber="100">
Description.
<emphasis id="B900EAE52E03C469A756FCCAFE7E636F" bold="true" box="[351,468,798,822]" italics="true" pageId="5" pageNumber="100">FEMALE</emphasis>
.
</emphasis>
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEFC97FB4163B2" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFC97FE2C6302" box="[199,390,835,859]" italics="true" pageId="5" pageNumber="100">Dorsal idiosoma</emphasis>
(
<figureCitation id="134F2A722E03C469A790FC96FE456302" box="[409,495,834,859]" captionStart="FIGURES 14 15" captionStartId="6.[151,263,1835,1858]" captionTargetBox="[158,1425,743,1809]" captionTargetId="figure@6.[151,1436,735,1814]" captionTargetPageId="6" captionText="FIGURES 14 15. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322030/files/figure.png" pageId="5" pageNumber="100">Fig. 14</figureCitation>
). Dorsal shield 780925 µm long and 460560 µm wide, elongated and suboval (length/width 1.631.85), delicately and evenly reticulated on surface, bearing 30 pairs of dorsal setae. Dorsal setae uniform, smooth and needle-shaped, mostly subequal in length, relatively longer and mostly with tips reaching to or beyond insertions of following setae; setae z1 shortest. For length and spacing of some selected dorsal shield setae see
<tableCitation id="C6F6034C2E03C469A708FC06FEFE63B2" box="[257,340,978,1003]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1374,202,1650]" captionTargetPageId="12" captionText="TABLE 1. Measurements (in µm) of four species of Pachylaelaps (Longipachylaelaps)." httpUri="http://table.plazi.org/id/DF0B667F2E0AC460A69EFF4CFBEF60F6" pageId="5" pageNumber="100" tableUuid="DF0B667F2E0AC460A69EFF4CFBEF60F6">Table 1</tableCitation>
. Dorsolateral soft integument with seven pairs of marginal setae (r6, R1, R3R7).
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEFC22FEFB6552" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFC22FE2F6457" box="[199,389,1014,1038]" italics="true" pageId="5" pageNumber="100">Ventral idiosoma</emphasis>
(
<figureCitation id="134F2A722E03C469A79DFC22FE4C6456" box="[404,486,1014,1039]" captionStart="FIGURES 14 15" captionStartId="6.[151,263,1835,1858]" captionTargetBox="[158,1425,743,1809]" captionTargetId="figure@6.[151,1436,735,1814]" captionTargetPageId="6" captionText="FIGURES 14 15. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., female. 14. Dorsal idiosoma; 15. Ventral idiosoma. Not scaled." httpUri="https://zenodo.org/record/322030/files/figure.png" pageId="5" pageNumber="100">Fig. 15</figureCitation>
). Sternal shield 270290 µm long, slightly shorter than genitiventral shield (length of sternal shield/length of genitiventral shield 0.90.95). Genitiventral shield almost subequal in length and width (length 289318 µm, width 289310 µm, length/width 0.931.05). Anal shield subtriangular, 121143 µm long and 173213 µm wide (length/width 0.580.74); anus with three circum-anal setae situated close to posterior margin of shield. Peritremes normally developed, anterior tips reaching dorsal surface close to insertions of setae z1. Ventral shields with delicate reticulation on surface. Metapodal platelets minuscule, free on soft integument. Ventral soft integument with nine pairs of ventral setae (JV2JV5, ZV1ZV5). Ventral setae similar to those on dorsal idiosoma.
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEFAC3FA29652F" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFAC3FE526577" box="[199,504,1302,1326]" italics="true" pageId="5" pageNumber="100">Sperm induction structures</emphasis>
(
<figureCitation id="134F2A722E03C469A401FAC2FDCA6576" box="[520,608,1302,1327]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 16</figureCitation>
,
<figureCitation id="134F2A722E03C469A464FAC2FD126576" box="[621,696,1302,1327]" captionStart="FIGURES 23 26" captionStartId="8.[151,263,977,1000]" captionTargetBox="[151,1436,193,957]" captionTargetId="figure@8.[151,1436,193,957]" captionTargetPageId="8" captionText="FIGURES 23 26. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov., sperm induction structures, female, variant forms. Not scaled." httpUri="https://zenodo.org/record/322032/files/figure.png" pageId="5" pageNumber="100">2326</figureCitation>
). Tubiform structures of sperm induction system well developed, relatively large, with apices adjacent but separated, weakly sclerotised except short apical to subapical portions, moderately and regularly narrowed toward apices, with bases closely abutting anterior inner margin of coxae IV.
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEFA57FE2E662A" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEFA57FE7C65C3" box="[199,470,1411,1435]" italics="true" pageId="5" pageNumber="100">Gnathosomal structures</emphasis>
(
<figureCitation id="134F2A722E03C469A7EEFA56FDC765C2" box="[487,621,1410,1435]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 17, 18</figureCitation>
). Corniculi elongated and horn-like; laciniae densely pilose, as long as corniculi; deutosternum with six rows of denticles; subcapitular setae smooth and needle-shaped. Epistome with subtriangular and regularly narrowed base, narrow central neck and widened apical part densely crenulate anteriorly; basal part with denticulate lateral margins (
<figureCitation id="134F2A722E03C469A50BFA3AFCFD665E" box="[770,855,1518,1543]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Fig. 17</figureCitation>
). Fixed digit of chelicera with bifid terminal hook, one large distal tooth associated with pilus dentilis, and one additional distal denticle on lateral ridge; pilus dentilis well developed, curved and directed backward; movable digit of chelicera unidentate, with simple hook and one distal tooth (
<figureCitation id="134F2A722E03C469A72AF98EFEDD662A" box="[291,375,1626,1651]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEF9ABFF6966E2" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEF9ABFF5666CF" box="[199,252,1663,1686]" italics="true" pageId="5" pageNumber="100">Legs</emphasis>
. Leg setation normal for genus (Mašán, 2007). Tarsus II with two spur-like distal setae, pl1 and pl2 (
<figureCitation id="134F2A722E03C469A366F9AAFF1C66E2" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEF913FE1C675A" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEF913FEA66684" bold="true" box="[199,268,1735,1758]" italics="true" pageId="5" pageNumber="100">MALE</emphasis>
<emphasis id="B900EAE52E03C469A702F912FEB86686" bold="true" box="[267,274,1734,1759]" pageId="5" pageNumber="100">.</emphasis>
<emphasis id="B900EAE52E03C469A717F912FE756687" box="[286,479,1734,1758]" italics="true" pageId="5" pageNumber="100">Dorsal idiosoma</emphasis>
. Dorsal shield 720810 µm long and 425475 µm wide, elongated and suboval (length/width 1.621.71).
</paragraph>
<paragraph id="8BCB36F72E03C469A6CEF8DAFAF667CA" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEF8DAFE21677F" box="[199,395,1806,1830]" italics="true" pageId="5" pageNumber="100">Ventral idiosoma</emphasis>
. Sternal, genitiventral, peritrematal, metapodal and anal plates fused to form an entire holoventral shield bearing nine pairs of setae (st1st5, JV1JV3, ZV2), not including three circum-anal setae close to anus; shield with reticulate sculpture on surface. Dorsolateral and ventral soft integument with 13 pairs of setae (r6, R1, R3R7, JV4, JV5, Zv1, ZV3ZV5). Dorsal and ventral chaetotaxy and other characters as in female.
</paragraph>
<paragraph id="8BCB36F72E03C46AA6CEF84AFF45613C" blockId="5.[151,1437,402,2011]" lastBlockId="6.[151,1437,151,680]" lastPageId="6" lastPageNumber="101" pageId="5" pageNumber="100">
<emphasis id="B900EAE52E03C469A6CEF84AFE7E67EF" box="[199,468,1950,1974]" italics="true" pageId="5" pageNumber="100">Gnathosomal structures</emphasis>
(
<figureCitation id="134F2A722E03C469A7EDF84AFDCD67EE" box="[484,615,1950,1975]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="5" pageNumber="100">Figs 20, 21</figureCitation>
,
<figureCitation id="134F2A722E03C469A47DF84BFD3867EE" box="[628,658,1951,1975]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="5" pageNumber="100">37</figureCitation>
,
<figureCitation id="134F2A722E03C469A497F84AFD1467EE" box="[670,702,1950,1975]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="5" pageNumber="100">54</figureCitation>
). Cheliceral spermatodactyl 130150 µm in length, 22.25 times as long as movable digit, expanded submedially, slightly curved, leaf-like to scimitar-like, with smooth margins and relatively wide terminal portion (
<figureCitation id="134F2A722E00C46AA443FF43FD0F60E9" box="[586,677,151,176]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="6" pageNumber="101">Figs 20</figureCitation>
,
<figureCitation id="134F2A722E00C46AA4BDFF43FD7E60E9" box="[692,724,151,176]" captionStart="FIGURES 53 56" captionStartId="19.[151,263,1900,1923]" captionTargetBox="[151,1436,195,1879]" captionTargetId="figure@19.[151,1436,195,1879]" captionTargetPageId="19" captionText="FIGURES 53 56. Chelicerae, males, with variant forms of spermatodactyls. 53. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 54. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 55. Pachylaelaps (Longipachylaelaps) pantinii sp. nov.; 56. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov. Not scaled." httpUri="https://zenodo.org/record/322039/files/figure.png" pageId="6" pageNumber="101">54</figureCitation>
). Palptibiae slightly thickened basally, each bearing a pair of petal-like projections on proximal ventral surface, as in
<figureCitation id="134F2A722E00C46AA503FF68FCF4608D" box="[778,862,188,213]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="6" pageNumber="101">Figs 21</figureCitation>
and
<figureCitation id="134F2A722E00C46AA59CFF68FC1E608D" box="[917,948,188,212]" captionStart="FIGURES 36 39" captionStartId="11.[151,263,874,897]" captionTargetBox="[151,1436,193,853]" captionTargetId="figure@11.[151,1436,193,853]" captionTargetPageId="11" captionText="FIGURES 36 39. Palptibial projections, males. 36. Pachylaelaps (Longipachylaelaps) abludens sp. nov.; 37. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov.; 38. Pachylaelaps (Longipachylaelaps) marcovallei sp. nov.; 39. Pachylaelaps (Longipachylaelaps) pantinii sp. nov. Not scaled." httpUri="https://zenodo.org/record/322035/files/figure.png" pageId="6" pageNumber="101">37</figureCitation>
; projections relatively large, with divergent adjacent margins: antiaxial projection longer, with narrow base, wide medial expansion, rounded distal margin, small subapical incision and small lamellar prominence at its inner base; paraxial projection shorter, subtriangular, widened basally and tapered toward obtuse apex. Palpfemur with distinct papular tubercle on anterolateral basal surface.
</paragraph>
<paragraph id="8BCB36F72E00C46AA6CEFEA5FF5161AC" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="101">
<emphasis id="B900EAE52E00C46AA6CEFEA5FF5661D1" box="[199,252,369,392]" italics="true" pageId="6" pageNumber="101">Legs</emphasis>
. Medial segments of legs II spurred on their distal ventral surface: femur with one robust spur, genu and tibia each with a peg-like spur, as in
<figureCitation id="134F2A722E00C46AA424FE40FDD561F5" box="[557,639,404,429]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="6" pageNumber="101">Fig. 22</figureCitation>
. Femoral spur broadened in basal part, produced into narrowly rounded apex, with two small tubercles on axillary side (
<figureCitation id="134F2A722E00C46AA4B5FE63FCB86189" box="[700,786,439,464]" captionStart="FIGURES 16 22" captionStartId="7.[151,263,1834,1857]" captionTargetBox="[156,1429,201,1810]" captionTargetId="figure@7.[151,1436,193,1814]" captionTargetPageId="7" captionText="FIGURES 16 22. Pachylaelaps (Longipachylaelaps) bergomensis sp. nov. 16. Sperm induction structures, female; 17. Epistome, female; 18. Chelicera, female; 19. Tarsus II, female, mediodistal part; 20. Chelicera, male; 21. Palptibial projections, male; 22. Ventral spurs on leg II (femur, genu and tibia), male. Not scaled." httpUri="https://zenodo.org/record/322031/files/figure.png" pageId="6" pageNumber="101">Fig. 22</figureCitation>
). Terminal part of tarsus II with only one spur-like distal seta, pl1.
</paragraph>
</subSubSection>
<subSubSection id="C36E657C2E00C46AA6CEFE2BFE1A62F1" pageId="6" pageNumber="101" type="etymology">
<paragraph id="8BCB36F72E00C46AA6CEFE2BFC9E6264" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="101">
<emphasis id="B900EAE52E00C46AA6CEFE2BFEE46241" bold="true" box="[199,334,511,536]" pageId="6" pageNumber="101">Etymology.</emphasis>
The epithet of this species is derived from the Latin name “
<emphasis id="B900EAE52E00C46AA5FBFDD5FBC66241" box="[1010,1132,513,536]" italics="true" pageId="6" pageNumber="101">Bergomum</emphasis>
” (Bergamo) and alludes to the
<typeStatus id="54CF88552E00C46AA6C9FDF1FF5A6264" box="[192,240,549,573]" pageId="6" pageNumber="101">type</typeStatus>
locality situated in the urban area of Bergamo City.
</paragraph>
<paragraph id="8BCB36F72E00C46AA6CEFD93FE1A62F1" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="101">
<emphasis id="B900EAE52E00C46AA6CEFD93FE3C6239" bold="true" box="[199,406,583,608]" pageId="6" pageNumber="101">Taxonomic notes.</emphasis>
The new species may be distinguished from the other congeners by the characters discussed in the diagnosis (see above), and the key below. In addition, the species differs from the related species by the features listed in
<tableCitation id="C6F6034C2E00C46AA75EFD5BFE0762F1" box="[343,429,655,680]" captionStart="TABLE 2" captionStartId="16.[146,221,530,550]" captionText="TABLE 2. Comparative characteristics of selected features in the Pachylaelaps (Longipachylaelaps) species. SL = length of cheliceral spermatodactyl, MDL = length of movable digit of chelicera, ― = unknown adult stage,? = unknown character)." pageId="6" pageNumber="101">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>