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<mods:title id="DAC6D30145966B7710067B468D3B4D03">Taxonomic status of two enigmatic Mexican anoles: Anolis cumingii Peters 1863 and Anolis guentherii Bocourt 1873 (Reptilia, Squamata, Dactyloidae)</mods:title>
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<taxonomicName id="4C744D40FFA677352EC9FD55FE1E1EB1" authority="Peters" authorityName="Peters" box="[151,429,690,716]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA677352EC9FD55FE1E1EB1" bold="true" box="[151,429,690,716]" pageId="1" pageNumber="83">
<emphasis id="B900EAD1FFA677352EC9FD55FEE41EB1" bold="true" box="[151,343,690,716]" italics="true" pageId="1" pageNumber="83">Anolis cumingii</emphasis>
Peters
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<taxonomicName id="4C744D40FFA677352EC9FD1EFE5B1F72" authority="Peters 1863:140" authorityName="Peters" authorityPageNumber="140" authorityYear="1863" box="[151,488,761,783]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA677352EC9FD1EFE8B1F72" box="[151,312,761,783]" italics="true" pageId="1" pageNumber="83">Anolis cumingii</emphasis>
<treatmentCitation id="0AD510D2FFA677352F61FD1DFE5B1F72" author="Peters" box="[319,488,761,783]" page="140" pageId="1" pageNumber="83" year="1863">
<bibRefCitation id="EFE54B32FFA677352F61FD1DFE081F72" author="Peters" box="[319,443,761,783]" pageId="1" pageNumber="83" refString="Peters, W. C. H. (1863) Uber einige neue Arten der Saurier-Gattung Anolis. Monatsberichte der Koniglich Preußischen Akademie der Wissenschaften zu Berlin, 1863, 135 - 149." type="journal article" year="1863">Peters 1863</bibRefCitation>
:140
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;
<typeStatus id="54CF8861FFA677352FADFD1EFDFF1F72" box="[499,588,761,783]" pageId="1" pageNumber="83" type="holotype">holotype</typeStatus>
(ZMB 4105) from “
<collectingCountry id="F3637653FFA677352D7FFD1EFCC71F72" box="[801,884,761,783]" name="Mexico" pageId="1" pageNumber="83">Mexico</collectingCountry>
.”
<bibRefCitation id="EFE54B32FFA677352DD7FD1EFBEF1F72" author="Troschel" box="[905,1116,761,783]" pageId="1" pageNumber="83" refString="Troschel, F. H. (1864) Bericht uber die Leistungen in der Herpetologie wahrend des Jahres 1863. Archiv fur Naturgeschichte, 30 (2), 205 - 224." type="journal article" year="1864">Troschel (1864:213)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352A39FD1EFA941F72" author="Muller" box="[1127,1319,761,783]" pageId="1" pageNumber="83" refString="Muller, J. W. (1865) Reisen in die Vereinigten Staaten, Canada und Mexico. III. Beitrage zur Geschichte, Statistik und Zoologie von Mexico. Dritte Abteilung. Die Wirbelthiere Mexicos. III. Amphibia. Leipzig, Brockhaus." type="book" year="1865">Müller (1865:601)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352B6DFD1EFE9E1F53" author="Boulenger" pageId="1" pageNumber="83" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History). 2 nd ed., Vol. II. London (Trustees of the British Museum)." type="book" year="1885">Boulenger (1885:80)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352F66FCFFFE4B1F53" author="Gunther" box="[312,504,792,814]" pageId="1" pageNumber="83" refString="Gunther, A. C. L. G. (1885 - 1902) Biologia Centrali - Americana. Reptila and Batrachia. Porter, London." type="book" year="1885">Günther (1885:50)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352C5DFCFFFD631F53" author="Barbour" box="[515,720,792,814]" pageId="1" pageNumber="83" refString="Barbour, T. (1934) The anoles II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology, 77, 119 - 155." type="journal article" year="1934">Barbour (1934:129)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352C85FCFEFC461F53" author="Burt" box="[731,1013,792,814]" pageId="1" pageNumber="83" refString="Burt, C. E. &amp; Myers, G. S. (1942) Neotropical lizards in the collection of the Natural History Museum of Stanford University. Stanford University Publications University Series Biological Sciences, 8 (2), 273 - 324." type="journal article" year="1942">Burt and Myers (1942:291)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352A5EFCFFFA9A1F53" author="Smith" box="[1024,1321,792,814]" pageId="1" pageNumber="83" refString="Smith, H. M., &amp; Taylor, E. H. (1950 a) An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bulletin of the United States National Museum, 199, 1 - 253." type="journal article" year="1950" yearSuffix="a">Smith and Taylor (1950a:67)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352B6AFCFFFE3F1F31" author="Smith" pageId="1" pageNumber="83" refString="Smith, H. M., &amp; Taylor, E. H. (1950 b) Type localities of Mexican reptiles and amphibians. Kansas University Science Bulletin, 33 (8), 313 - 380." type="journal article" year="1950" yearSuffix="b">Smith and Taylor (1950b:320)</bibRefCitation>
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<taxonomicName id="4C744D40FFA677352EC9FCB2FEF11F16" box="[151,322,853,875]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cummingi">
<emphasis id="B900EAD1FFA677352EC9FCB2FEF11F16" box="[151,322,853,875]" italics="true" pageId="1" pageNumber="83">Anolis cummingi</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE54B32FFA677352F10FCB1FDBA1F16" author="Stuart" box="[334,521,853,875]" pageId="1" pageNumber="83" refString="Stuart, L. C. (1955) A brief review of the Guatemalan lizards of the genus Anolis. Miscellaneous Publication Museum of Zoology University of Michigan, 91, 1 - 31." type="journal article" year="1955">Stuart (1955:4,26)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352C4AFCB2FD5C1F16" author="Etheridge" box="[532,751,853,875]" pageId="1" pageNumber="83" refString="Etheridge, R. (1959) The Relationships of the Anoles (Reptilia: Sauria: Iguanidae). An Interpretation Based on Skeletal Morphology. Ph. D. Dissertation, University of Michigan, Ann Arbor, Michigan, USA." type="book" year="1959">Etheridge (1959:210)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352CA4FCB2FBF71F16" author="Fitch" box="[762,1092,853,875]" pageId="1" pageNumber="83" refString="Fitch, H. S. &amp; Henderson, R. W. (1973) A new anole (Reptilia: Iguanidae) from southern Veracruz, Mexico. Journal of Herpetology, 7 (2), 125 - 128." type="journal article" year="1973">Fitch and Henderson (1973:126)</bibRefCitation>
</paragraph>
<paragraph id="8BCB36C3FFA677352EC9FC93FD0D1FF7" blockId="1.[151,1436,761,937]" box="[151,702,884,906]" pageId="1" pageNumber="83">
<taxonomicName id="4C744D40FFA677352EC9FC93FEFA1FF7" box="[151,329,884,906]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cummingii">
<emphasis id="B900EAD1FFA677352EC9FC93FEFA1FF7" box="[151,329,884,906]" italics="true" pageId="1" pageNumber="83">Anolis cummingii</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE54B32FFA677352F08FC92FDA01FF7" author="Bocourt" box="[342,531,884,906]" pageId="1" pageNumber="83" refString="Bocourt, M. F. (1873) In A. Dumeril, Bocourt, M. - F. &amp; Mocquard, F. (eds.), Etudes sur les reptiles. In: Recherches zoologiques pour sevir a l´histoire de la faune de l´Amerique Centrale et du Mexique. Mission Scientifique au Mexique et dans l´Amerique Centrale, recherches Zool., part 3, sect. 1. Imprimerie au Nat.; Paris." type="book" year="1873">Bocourt (1873:89)</bibRefCitation>
,
<bibRefCitation id="EFE54B32FFA677352C40FC93FD0D1FF7" author="Cope" box="[542,702,884,906]" pageId="1" pageNumber="83" refString="Cope, E. D. (1887) Catalogue of Batrachians and reptiles of Central America and Mexico. Bulletin of the United States National Museum, 32, 1 - 98." type="journal article" year="1887">Cope (1887:31)</bibRefCitation>
</paragraph>
<paragraph id="8BCB36C3FFA677352EC9FC73FDC81FD4" blockId="1.[151,1436,761,937]" box="[151,635,914,937]" pageId="1" pageNumber="83">
<taxonomicName id="4C744D40FFA677352EC9FC73FE8C1FD4" box="[151,319,916,937]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA677352EC9FC73FE8C1FD4" box="[151,319,916,937]" italics="true" pageId="1" pageNumber="83">Norops cumingii</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE54B32FFA677352F12FC74FDC81FD4" author="Savage" box="[332,635,914,937]" pageId="1" pageNumber="83" refString="Savage, J. M. &amp; Guyer, C. (1989) Infrageneric classification and species composition of the anole genera, Anolis, Ctenonotus, Dactyloa, Norops and Semiurus (Sauria: Iguanidae). Amphibia - Reptilia, 10, 105 - 116." type="journal article" year="1989">Savage and Guyer (1989:111)</bibRefCitation>
</paragraph>
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<emphasis id="B900EAD1FFA677352EC9FC32FC181F93" bold="true" box="[151,939,981,1006]" pageId="1" pageNumber="83">
Description of the
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of
<taxonomicName id="4C744D40FFA677352FA1FC32FD0B1F93" box="[511,696,981,1006]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="83" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA677352FA1FC32FD0B1F93" bold="true" box="[511,696,981,1006]" italics="true" pageId="1" pageNumber="83">Anolis cumingii</emphasis>
</taxonomicName>
(ZMB 4105;
<figureCitation id="134F2A46FFA677352D06FC32FC281F93" box="[856,923,981,1006]" captionStart="FIGURE 1" captionStartId="2.[151,250,1320,1343]" captionTargetBox="[151,1435,327,1279]" captionTargetId="figure@2.[151,1436,327,1279]" captionTargetPageId="2" captionText="FIGURE 1. — Holotype of Anolis cumingii (ZMB 4105; SVL 46.0 mm)." httpUri="https://zenodo.org/record/282853/files/figure.png" pageId="1" pageNumber="83">Fig. 1</figureCitation>
).
</emphasis>
Adult male as indicated by shape of base of tail and size of dewlap; SVL 46.0 mm; tail length 102.0 mm, tail complete; tail slightly compressed in cross section, tail height
<quantity id="4C8C9B26FFA677352F35FBFAFE701848" box="[363,451,1053,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="83" unit="mm" value="2.5">2.5 mm</quantity>
, tail width
<quantity id="4C8C9B26FFA677352C61FBFAFD241848" box="[575,663,1053,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="83" unit="mm" value="2.1">2.1 mm</quantity>
; axilla to groin distance
<quantity id="4C8C9B26FFA677352DF6FBFAFBBC1848" box="[936,1039,1053,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="1" pageNumber="83" unit="mm" value="19.6">19.6 mm</quantity>
; head length 13.0 mm, head length/ SVL ratio 0.28; snout length 6.0 mm; head width
<quantity id="4C8C9B26FFA677352CA5FBA6FCE91824" box="[763,858,1089,1113]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="83" unit="mm" value="7.5">7.5 mm</quantity>
; longest toe of adpressed hind limb reaching to tympanum; shank length 11.0 mm, shank length/head length ratio 0.87. Scales on snout varying from faintly to strongly keeled; 5 postrostrals; 8 scales between nasals; 1 prenasal scale in contact with both rostral and first supralabial; most scales in distinct prefrontal depression smooth, some of them keeled; supraorbital semicircles well developed, separated by a minimum of one scale row; supraorbital disc composed of about 9 distinctly enlarged, faintly keeled scales; circumorbital row of small scales incomplete, thus, 1 (right)2 (left) enlarged supraorbitals in contact with supraorbital semicircles; a single large elongated superciliary; 2 rows of small keeled scales extending between enlarged supraorbitals and superciliary; no parietal depression; interparietal scale well developed, 1.4 x 1.0 mm (length x width), surrounded by scales of moderate size anteriorly and by small to moderate size scales posteriorly; 3 scales present between interparietal and supraorbital semicircles; canthal ridge distinct, composed of 6 canthal scales, with 3 larger posterior scales; 6 scales present between second canthals; 7 scales present between posterior canthals; 26 (right)27 (left) loreal scales in a maximum of 6 (right)5 (left) horizontal rows, with the scales mostly keeled; 5 subocular scales (anterior ones keeled) arranged in a single row; 7 (right)6 (left) supralabials to level below center of eye; 2 suboculars broadly in contact with supralabials; ear opening 0.7 x
<quantity id="4C8C9B26FFA677352F62F9DEFE271A2C" box="[316,404,1593,1617]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="83" unit="mm" value="0.7">0.7 mm</quantity>
(length x height), ratio tympanum height/parietal scale length 0.5; mental distinctly wider than long, completely divided medially, bordered posteriorly by 4 postmentals; 8 (right)7 (left) infralabials to level below center of eye; sublabials undifferentiated; small elongate keeled scales present on chin and throat; dewlap small, extending from level below oral ricti to a point
<quantity id="4C8C9B26FFA677352D3FF942FC171AC0" box="[865,932,1701,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="83" unit="mm" value="2.0">2 mm</quantity>
posterior to level of axilla; dorsum of body with weakly keeled scales with rounded posterior margins, granular on the neck and becoming more flattened posteriorly, 17 medial rows slightly enlarged, about 0.34 x
<quantity id="4C8C9B26FFA677352D62F90AFC111B78" box="[828,930,1773,1797]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="83" unit="mm" value="0.25">0.25 mm</quantity>
(length x width); 33 medial dorsal scales in one head length; 63 medial dorsal scales between levels of axilla and groin; lateral scales homogeneous, about 0.20 x
<quantity id="4C8C9B26FFA677352EF0F8D2FEA71B30" box="[174,276,1845,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="83" unit="mm" value="0.14">0.14 mm</quantity>
(length x width); ventrals at midbody keeled, mucronate, imbricate, about 0.64 x
<quantity id="4C8C9B26FFA677352A98F8D2FA9F1B30" box="[1222,1324,1845,1869]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="83" unit="mm" value="0.44">0.44 mm</quantity>
(length x width); 29 ventral scales in one head length, 43 ventral scales between levels of axilla and groin; 101 scales around midbody; caudal scales strongly keeled, without whorls of enlarged scales, although an indistinct division in segments is discernible; postcloacal scales not enlarged; no tube-like axillary pocket present; scales on dorsal surface of forelimb keeled, imbricate; digital pads dilated, dilated pad (width
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) almost three times the width of non-dilated scales on distal phalanx (
<quantity id="4C8C9B26FFA577362C0EFF70FD0B1CD2" box="[592,696,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="84" unit="mm" value="0.25">0.25 mm</quantity>
); 25 (right)26 (left) lamellae under phalanges IIIV of fourth toe; 7 (right)6 (left) scales under distal phalanx of fourth toe. Overall dorsal coloration pearl gray with a suggestion of olive brown on snout, neck, flank, and ventral surfaces; dewlap olive gray with a darker central area.
</paragraph>
<caption id="DF0B664BFFA577362EC9FACFFCC41942" box="[151,887,1320,1343]" httpUri="https://zenodo.org/record/282853/files/figure.png" pageId="2" pageNumber="84" targetBox="[151,1435,327,1279]" targetPageId="2">
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<emphasis id="B900EAD1FFA577362EC9FACFFEA01943" bold="true" box="[151,275,1320,1343]" pageId="2" pageNumber="84">FIGURE 1.</emphasis>
—Holotype of
<taxonomicName id="4C744D40FFA577362FF2FACEFDFF1942" box="[428,588,1321,1343]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA577362FF2FACEFDFF1942" box="[428,588,1321,1343]" italics="true" pageId="2" pageNumber="84">Anolis cumingii</emphasis>
</taxonomicName>
(ZMB 4105; SVL 46.0 mm).
</paragraph>
</caption>
<paragraph id="8BCB36C3FFA577362E99FA82FF4119BB" blockId="2.[151,1436,1381,1478]" pageId="2" pageNumber="84">
<emphasis id="B900EAD1FFA577362E99FA82FE551903" bold="true" box="[199,486,1381,1406]" pageId="2" pageNumber="84">Taxonomic conclusions:</emphasis>
A comparison of the
<typeStatus id="54CF8861FFA577362CB8FA81FCFB1903" box="[742,840,1382,1406]" pageId="2" pageNumber="84" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C744D40FFA577362D2DFA81FB951903" box="[883,1062,1382,1406]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA577362D2DFA81FB951903" box="[883,1062,1382,1406]" italics="true" pageId="2" pageNumber="84">Anolis cumingii</emphasis>
</taxonomicName>
with all other known species of anoles leads me to the conclusion that it is conspecific with
<taxonomicName id="4C744D40FFA577362D02FA6DFB2B19DF" authority="Hallowell 1856" authorityName="Hallowell" authorityYear="1856" box="[860,1176,1418,1442]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="sericeus">
<emphasis id="B900EAD1FFA577362D02FA6DFC6919DF" box="[860,986,1418,1442]" italics="true" pageId="2" pageNumber="84">A. sericeus</emphasis>
Hallowell 1856
</taxonomicName>
. Diagnostic characters include:
</paragraph>
<paragraph id="8BCB36C3FFA577362EC9FA11FDED1A38" blockId="2.[151,930,1526,1693]" pageId="2" pageNumber="84">(1) short legged (longest toe of adpressed hindlimb reaches to tympanum); (2) tympanum very small (less than one fourth the size of interparietal plate); (3) a single large and elongate superciliary;</paragraph>
<paragraph id="8BCB36C3FFA577362EC9F9A9FDEC1AFD" blockId="2.[151,930,1526,1693]" pageId="2" pageNumber="84">(4) ventral scales strongly keeled, imbricate, and mucronate. (5) no enlarged postcloacal scales in males;</paragraph>
<paragraph id="8BCB36C3FFA577362EC9F96FFD951AE0" blockId="2.[151,930,1526,1693]" box="[151,550,1672,1693]" pageId="2" pageNumber="84">(6) a moderate-sized dewlap in males</paragraph>
<paragraph id="8BCB36C3FFA577362E99F92DFDA91B33" blockId="2.[151,1436,1738,1942]" pageId="2" pageNumber="84">
Therefore, I herewith place
<taxonomicName id="4C744D40FFA577362C5CF92DFC8C1A9F" authority="Peters 1963" authorityName="Peters" authorityYear="1963" box="[514,831,1738,1762]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B900EAD1FFA577362C5CF92DFD001A9F" box="[514,691,1738,1762]" italics="true" pageId="2" pageNumber="84">Anolis cumingii</emphasis>
Peters
<quantity id="4C8C9B26FFA577362D59F92DFC8C1A9F" box="[775,831,1738,1762]" metricMagnitude="1" metricUnit="m" metricValue="4.98602" pageId="2" pageNumber="84" unit="in" value="1963.0">1963</quantity>
</taxonomicName>
in the synonymy of
<taxonomicName id="4C744D40FFA577362A74F92DFAED1A9F" authority="Hallowell 1856" authorityName="Hallowell" authorityYear="1856" box="[1066,1374,1738,1762]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="sericeus">
<emphasis id="B900EAD1FFA577362A74F92DFB151A9F" box="[1066,1190,1738,1762]" italics="true" pageId="2" pageNumber="84">A. sericeus</emphasis>
Hallowell 1856
</taxonomicName>
. This species is currently known from the Atlantic versant of the Mexican states of Tamaulipas, Hidalgo, San Luis Potosí, Veracruz, Tabasco, Campeche, Quintana Roo, and Yucatán as well as the extreme northern portion of Oaxaca (
<bibRefCitation id="EFE54B32FFA577362EA7F8D1FDBD1B33" author="Kohler" box="[249,526,1846,1870]" pageId="2" pageNumber="84" refString="Kohler, G. &amp; Vesely, M. (2010) A revision of the Anolis sericeus complex with the resurrection of A. wellbornae and the description of a new species (Squamata: Polychrotidae). Herpetologica, 66 (2), 207 - 228." type="journal article" year="2010">Köhler and Vesely 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BCB36C3FFA577362E99F8BDFAB21BEB" blockId="2.[151,1436,1738,1942]" pageId="2" pageNumber="84">
The only other species in the region that possibly can be confused with this species are
<taxonomicName id="4C744D40FFA577362AEDF8BDFAF91B0F" box="[1203,1354,1882,1906]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="unilobatus">
<emphasis id="B900EAD1FFA577362AEDF8BDFAF91B0F" box="[1203,1354,1882,1906]" italics="true" pageId="2" pageNumber="84">A. unilobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D40FFA577362BDAF8BDFEA61BEB" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="wellbornae">
<emphasis id="B900EAD1FFA577362BDAF8BDFEA61BEB" italics="true" pageId="2" pageNumber="84">A. wellbornae</emphasis>
</taxonomicName>
, which differ from
<taxonomicName id="4C744D40FFA577362FAEF899FDD81BEB" box="[496,619,1918,1942]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="84" phylum="Chordata" rank="species" species="sericeus">
<emphasis id="B900EAD1FFA577362FAEF899FDD81BEB" box="[496,619,1918,1942]" italics="true" pageId="2" pageNumber="84">A. sericeus</emphasis>
</taxonomicName>
by having a larger male dewlap (
<bibRefCitation id="EFE54B32FFA577362DBEF899FB461BEB" author="Kohler" box="[992,1269,1918,1942]" pageId="2" pageNumber="84" refString="Kohler, G. &amp; Vesely, M. (2010) A revision of the Anolis sericeus complex with the resurrection of A. wellbornae and the description of a new species (Squamata: Polychrotidae). Herpetologica, 66 (2), 207 - 228." type="journal article" year="2010">Köhler and Vesely 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>