215 lines
19 KiB
XML
215 lines
19 KiB
XML
<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="186" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="100" approvalRequired_for_treatments="80" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy & Divljan, Anja" docDate="2017" docId="03DD87C8FF9273191B18FEC1FC629011" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Phascolarctos cinereus subsp. victor Troughton 1935" docType="treatment" docVersion="3" lastPageNumber="320" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="320" updateTime="1674153641407" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Parnaby, Harry E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingleby, Sandy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Divljan, Anja</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-10-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>277</mods:start>
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<mods:end>420</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
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<mods:identifier type="ISSN">2201-4349</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
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<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FF9273191B18FEC1FC629011" httpUri="http://treatment.plazi.org/id/03DD87C8FF9273191B18FEC1FC629011" lastPageId="44" lastPageNumber="320" pageId="43" pageNumber="320">
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<subSubSection box="[810,1406,298,325]" pageId="43" pageNumber="320" type="nomenclature">
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<paragraph blockId="43.[810,1406,298,325]" box="[810,1406,298,325]" pageId="43" pageNumber="320">
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<heading box="[810,1406,298,325]" centered="true" fontSize="11" level="2" pageId="43" pageNumber="320" reason="2">
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<taxonomicName authority="Troughton, 1935 a" authorityName="Troughton" authorityYear="1935" box="[810,1406,298,325]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="43" pageNumber="320" phylum="Chordata" rank="subSpecies" species="cinereus" subSpecies="victor">
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<emphasis bold="true" box="[810,1406,298,325]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[810,1173,298,325]" italics="true" pageId="43" pageNumber="320">Phascolarctos cinereus victor</emphasis>
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<bibRefCitation author="Troughton, E" box="[1180,1406,298,324]" pageId="43" pageNumber="320" pagination="137 - 140" refId="ref147311" refString="Troughton, E. L. G. 1935 a. The southern race of the koala. Australian Naturalist 9 (6): 137 - 140." type="journal article" year="1935">Troughton, 1935a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[824,1268,350,372]" pageId="43" pageNumber="320" type="reference_group">
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<paragraph blockId="43.[824,1268,350,372]" box="[824,1268,350,372]" pageId="43" pageNumber="320">
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<emphasis box="[824,976,350,371]" italics="true" pageId="43" pageNumber="320">Aust. Naturalist</emphasis>
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. 9(6): 139. (
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<date box="[1095,1257,350,372]" pageId="43" pageNumber="320" value="1935-09">September 1935</date>
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).
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</paragraph>
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</subSubSection>
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<subSubSection box="[794,1052,398,423]" pageId="43" pageNumber="320" type="vernacular_names">
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<paragraph blockId="43.[794,1052,398,423]" box="[794,1052,398,423]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,968,398,423]" pageId="43" pageNumber="320">Common name</emphasis>
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. Koala.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="320" type="description">
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<paragraph blockId="43.[792,1422,449,591]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,957,449,473]" pageId="43" pageNumber="320">Current name</emphasis>
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.
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<taxonomicName authority="(Goldfuss, 1817)" baseAuthorityName="Goldfuss" baseAuthorityYear="1817" box="[973,1418,449,473]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="43" pageNumber="320" phylum="Chordata" rank="species" species="cinereus">
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<emphasis box="[973,1222,449,473]" italics="true" pageId="43" pageNumber="320">Phascolarctos cinereus</emphasis>
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(
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<bibRefCitation author="Goldfuss, G" box="[1239,1409,449,473]" pageId="43" pageNumber="320" refId="ref130367" refString="Goldfuss, G. A. 1817. Plates CLV Aa, CLV Ab. In Die Saugethiere in Abbildungen nach Natur, mit Beschreibungend. Part 65, ed. J. C. D. Schreiber, Erlange, Germany: Wolfgang Walther. [Plates only, no text.]" type="book" year="1817">Goldfuss, 1817</bibRefCitation>
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)
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</taxonomicName>
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. Most recent authors assume that previously recognized subspecies of the koala represent clinal variation, see Jackson & Groves (2015).
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<bibRefCitation author="Neaves, L & Frankham, S & Dennison, S & FitzGibbon, C & Flannagan, A & Gillett, E & Hynes, K & Handasyde, K & Helgen, K & Tsangaras, A & Greenwood, M" box="[987,1205,537,561]" pageId="43" pageNumber="320" pagination="0162207" refId="ref139724" refString="Neaves, L. E., G. J. Frankham, S. Dennison, S. FitzGibbon, C. Flannagan, A. Gillett, E. Hynes, K. Handasyde, K. M. Helgen, K. Tsangaras, A. D. Greenwood, M. D. B. Eldridge, and R. N. Johnson. 2016. Phylogeography of the Koala, (Phascolarctos cinereus), and harmonising data to inform conservation. PLoS ONE 11 (9): e 0162207. https: // doi. org / 10.1371 / journal. pone. 0162207" type="journal article" year="2016">
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Neaves
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<emphasis box="[1074,1128,537,561]" italics="true" pageId="43" pageNumber="320">et al.</emphasis>
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(2016)
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</bibRefCitation>
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did not find support for a Victorian subspecies.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="320" type="materials_examined">
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<paragraph blockId="43.[794,1423,617,788]" pageId="43" pageNumber="320">
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<materialsCitation collectingDate="1934-09" location="Noel Burnet" pageId="43" pageNumber="320" specimenCount="2" stateProvince="Victoria" typeStatus="holotype">
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<emphasis bold="true" box="[794,898,617,641]" pageId="43" pageNumber="320">
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<typeStatus box="[794,898,617,641]" pageId="43" pageNumber="320">Holotype</typeStatus>
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</emphasis>
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. M.5732 by original designation. [Adult male], skull, study skin (
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<figureCitation box="[996,1127,647,671]" captionStart-0="Figure 20" captionStart-1="Figure 21" captionStartId-0="44.[229,292,1527,1549]" captionStartId-1="45.[121,184,1952,1974]" captionTargetBox-0="[167,1467,151,1508]" captionTargetBox-1="[126,726,182,1900]" captionTargetId-0="figure-10@44.[167,1467,151,1508]" captionTargetId-1="figure-9@45.[121,748,176,1910]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionText-0="Figure 20. AM M.5732, holotype skin of Phascolarctos cinereus victor Troughton, 1935a. (Photography by Sally Cowan)." captionText-1="Figure 21. AM M.5732, holotype skull of Phascolarctos cinereus victor Troughton, 1935a. (Photography by Stuart Humphreys)." figureDoi-0="http://doi.org/10.5281/zenodo.5237846" figureDoi-1="http://doi.org/10.5281/zenodo.5237848" httpUri-0="https://zenodo.org/record/5237846/files/figure.png" httpUri-1="https://zenodo.org/record/5237848/files/figure.png" pageId="43" pageNumber="320">Figs 20–21</figureCitation>
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), from
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<collectingRegion box="[1212,1302,647,671]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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, presented by
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<location LSID="urn:lsid:plazi:treatment:03DD87C8FF9273191B18FEC1FC629011:8EAB6005FF92731E1B0FFD4FFC119580" box="[829,964,676,700]" name="Noel Burnet" pageId="43" pageNumber="320" stateProvince="Victoria">Noel Burnet</location>
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, registered in
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<date box="[1119,1299,676,700]" pageId="43" pageNumber="320" value="1934-09">
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<collectingDate box="[1119,1299,676,700]" pageId="43" pageNumber="320" value="1934-09">September 1934</collectingDate>
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</date>
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. Collection date not recorded. Original wooden tag contains the word “Booral” but there is no entry for locality, either in the register, or on the
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<typeStatus box="[988,1081,764,788]" pageId="43" pageNumber="320">holotype</typeStatus>
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specimen index card.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="320" type="description">
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<paragraph blockId="43.[792,1422,814,985]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,906,814,838]" pageId="43" pageNumber="320">Condition</emphasis>
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. Cranium missing both upper 2nd and 3rd incisors and 1st post-canine teeth, missing right upper canine, drilled wire holes in temporal bones and condyles; right dentary appears calcified (remodeled) underneath the last post-canine tooth. Study skin missing left top lip, ear tips torn (fur sparse around the ear tips).
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</paragraph>
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<paragraph blockId="43.[792,1421,1012,1183]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,1116,1012,1036]" pageId="43" pageNumber="320">Cranial measurements (mm)</emphasis>
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. M.5732: GL, 159.34; ConL, 152.30; BasL, 144.83; NasL, 42.71; NasB, 43.36; UC1–C1 (alv.), 16.58; DIL, 24.80; APV, 5.63; PPV, 9.05; PAL, 77.63; UPM (alv.), 7.92; UMR (alv.), 28.09; ZB, 90.23; POC, 30.55; MB, 64.95; DL (angl.), 119.10; DL (condyl.), 116.78; LPM (alv.), 5.82; LMR (alv.), 33.11.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="320" type="distribution">
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<paragraph blockId="43.[793,1423,1209,1791]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,942,1209,1233]" pageId="43" pageNumber="320">Type locality</emphasis>
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. We suggest that the
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<typeStatus box="[1185,1231,1210,1234]" pageId="43" pageNumber="320">type</typeStatus>
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locality is Koala Sanctuary, French Island,
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<collectingRegion box="[1083,1173,1239,1263]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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. The
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<typeStatus box="[1238,1284,1239,1263]" pageId="43" pageNumber="320">type</typeStatus>
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locality has generally been cited simply as
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<collectingRegion box="[1114,1202,1268,1292]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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, reflecting confusion about a more precise location.
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<bibRefCitation author="McKay, G" box="[1115,1284,1297,1322]" pageId="43" pageNumber="320" refId="ref138692" refString="McKay, G. M. 1988 a. Phascolarctidae. In Zoological Catalogue of Australia. Vol. 5. Mammalia, ed. D. W. Walton, pp. 51 - 52. Canberra: Australian Government Publishing Service." type="book" year="1988">McKay (1988a)</bibRefCitation>
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lists the
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<typeStatus box="[1375,1421,1298,1322]" pageId="43" pageNumber="320">type</typeStatus>
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locality as “Booral”,
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<collectingRegion box="[1018,1104,1327,1351]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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without further comment but presumably on the assumption that the label refers to a collecting locality. We have been unable to locate a place name of “Booral” in
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<collectingRegion box="[1018,1107,1415,1439]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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. Instead, we located evidence that Booral was the name of a captive male koala: “Booral and Burrendong” were a pair of koalas sent in early 1930 to Noel Burnet at Koala Park, Sydney, as a gift from the Victorian Government, from the French Island Sanctuary (
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<bibRefCitation author="Williams, J" box="[801,978,1561,1585]" pageId="43" pageNumber="320" pagination="13" refId="ref149828" refString="Williams, J. 1931. Little Gum-Baby. Real live bears have toy " teddies " for pals. Sunday Times (Perth, Western Australia): No. 1769, p. 13, Sunday 20 December." type="book chapter" year="1931">Williams, 1931</bibRefCitation>
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). The caption of a photograph of Noel Burnet, head of Koala Park, holding a koala states: “The photo shows the author with “Booral,” the largest koala in the establishment” (
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<bibRefCitation author="Burnet, N" box="[1005,1190,1649,1673]" pageId="43" pageNumber="320" refId="ref126006" refString="Burnet, N. 1932. The Blue Gum Family at Koala Park. Sydney: W. A. Pepperday. 35 pp." type="book" year="1932">Burnet, 1932: 29</bibRefCitation>
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). That image shows a clear frontal view of the snout area and hirsute ears; cited as diagnostic criteria in Troughton’s account. Further evidence derives from the etching titled “Booral (The Roue of Koala Park)” by artist Squire Morgan, dated c. 1932.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="320" type="materials_examined">
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<paragraph blockId="43.[793,1423,1812,1924]" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,805,1812,1836]" pageId="43" pageNumber="320">?</emphasis>
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<materialsCitation collectionCode="AM" location="French Island" pageId="43" pageNumber="320" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
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<emphasis bold="true" box="[804,910,1812,1836]" pageId="43" pageNumber="320">
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<typeStatus box="[804,910,1812,1836]" pageId="43" pageNumber="320">Paratype</typeStatus>
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</emphasis>
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. M.4841, male, skull, study skin, from
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<location LSID="urn:lsid:plazi:treatment:03DD87C8FF9273191B18FEC1FC629011:8EAB6005FF92731E1D71F8FFFCB59075" name="French Island" pageId="43" pageNumber="320" stateProvince="Victoria">French Island</location>
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,
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<collectingRegion box="[873,962,1841,1865]" country="Australia" name="Victoria" pageId="43" pageNumber="320">Victoria</collectingRegion>
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, registered at the
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<collectionCode box="[1146,1189,1842,1866]" country="Australia" name="Australian Museum" pageId="43" pageNumber="320" type="Museum">AM</collectionCode>
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</materialsCitation>
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<materialsCitation collectingDate="1930-02-13" collectionCode="SAM" location="South Australian Museum" pageId="43" pageNumber="320" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
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on
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<date box="[1228,1419,1841,1865]" pageId="43" pageNumber="320" value="1930-02-13">
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<collectingDate box="[1228,1419,1841,1865]" pageId="43" pageNumber="320" value="1930-02-13">13 February 1930</collectingDate>
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</date>
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, received from
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<location LSID="urn:lsid:plazi:treatment:03DD87C8FF9273191B18FEC1FC629011:8EAB6005FF92731E1B84F8A4FB05905A" box="[950,1232,1871,1895]" name="South Australian Museum" pageId="43" pageNumber="320" stateProvince="Victoria">South Australian Museum</location>
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, previously
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<collectionCode box="[1363,1421,1871,1895]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="43" pageNumber="320">SAM</collectionCode>
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</materialsCitation>
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M.2443.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="44" lastPageNumber="321" pageId="43" pageNumber="320" type="discussion">
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<paragraph blockId="43.[792,1421,1945,1999]" lastBlockId="44.[838,1467,1608,1837]" lastPageId="44" lastPageNumber="321" pageId="43" pageNumber="320">
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<emphasis bold="true" box="[794,917,1945,1969]" pageId="43" pageNumber="320">Comments</emphasis>
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. In his original description Troughton mentions
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<specimenCount box="[792,955,1975,1999]" pageId="43" pageNumber="320" type="generic">two specimens</specimenCount>
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only, “The
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<typeStatus box="[1089,1185,1975,1999]" pageId="43" pageNumber="320">holotype</typeStatus>
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male, M.
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<quantity box="[1291,1378,1975,1999]" metricMagnitude="2" metricUnit="m" metricValue="1.455928" pageId="43" pageNumber="320" unit="in" value="5732.0">5732 in</quantity>
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the Museum collection, and a young male received from
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<location LSID="urn:lsid:plazi:treatment:03DD87C8FF9273191B18FEC1FC629011:8EAB6005FF9573191894F98DFE9C9142" box="[166,329,1638,1662]" county="French Island" municipality="Burnet's Koala Park in Sydney" name="French Island" pageId="44" pageNumber="321">French Island</location>
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in 1930 …”. No specimen marked as
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<typeStatus box="[167,262,1667,1691]" pageId="44" pageNumber="321">paratype</typeStatus>
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has been located in the collection but M.4841 is a likely candidate, despite the absence of any designation as such on its original skull and skin tags. Nevertheless, it is a young animal as indicated by an incompletely erupted M3 and small body size. Troughton did not give the sex or locality of the
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<typeStatus box="[317,411,1814,1838]" pageId="44" pageNumber="321">holotype</typeStatus>
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and indicated that he was providing a preliminary diagnosis ahead of a more detailed account being prepared for “a popular work”—presumably his book (
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<bibRefCitation author="Troughton, E" box="[914,1112,1667,1691]" pageId="44" pageNumber="321" refId="ref147799" refString="Troughton, E. [L. G.] 1941. Furred Animals of Australia. First edition. Sydney: Angus & Robertson. xxvii + 374 pp." type="book" year="1941">Troughton, 1941</bibRefCitation>
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||
). It is possible that Troughton was unaware of the origin of the
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<typeStatus box="[1201,1297,1696,1720]" pageId="44" pageNumber="321">holotype</typeStatus>
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sent to the
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<collectionCode box="[1423,1466,1697,1721]" country="Australia" name="Australian Museum" pageId="44" pageNumber="321" type="Museum">AM</collectionCode>
|
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from
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<collectingMunicipality box="[904,1268,1726,1750]" pageId="44" pageNumber="321">Burnet’s Koala Park in Sydney</collectingMunicipality>
|
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; alternatively, he might have been aware that both specimens were from
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<collectingCounty box="[839,986,1784,1808]" pageId="44" pageNumber="321">French Island</collectingCounty>
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and overlooked the ambiguity of his original statement.
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</paragraph>
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</subSubSection>
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||
</treatment>
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</document> |