treatments-xml/data/03/DD/87/03DD87C8FF8573081BB2FF7CFAD9963B.xml
2024-06-21 12:22:17 +02:00

336 lines
32 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="186" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="100" approvalRequired_for_treatments="80" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FF8573081BB2FF7CFAD9963B" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Macropus tibol Miklouho-Maclay 1885" docType="treatment" docVersion="3" lastPageNumber="338" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="337" updateTime="1674153641407" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Divljan, Anja</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B" httpUri="http://treatment.plazi.org/id/03DD87C8FF8573081BB2FF7CFAD9963B" lastPageId="61" lastPageNumber="338" pageId="60" pageNumber="337">
<subSubSection pageId="60" pageNumber="337" type="nomenclature">
<paragraph blockId="60.[896,1410,151,178]" box="[896,1410,151,178]" pageId="60" pageNumber="337">
<heading box="[896,1410,151,178]" centered="true" fontSize="11" level="2" pageId="60" pageNumber="337" reason="2">
<taxonomicName authority="Miklouho-Maclay, 1885 b" authorityName="Miklouho-Maclay" authorityYear="1885" box="[896,1410,151,178]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="60" pageNumber="337" phylum="Chordata" rank="species" species="tibol">
<emphasis bold="true" box="[896,1410,151,178]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[896,1083,151,178]" italics="true" pageId="60" pageNumber="337">Macropus tibol</emphasis>
<bibRefCitation author="Miklouho-Maclay, N" box="[1091,1410,151,177]" pageId="60" pageNumber="337" pagination="141 - 144" refId="ref139132" refString="Miklouho-Maclay, N. d. 1885 b. Notes on the zoology of the Maclay coast in New Guinea. II. On a new species of Macropus: Macropus tibol. Proceedings of the Linnean Society of New South Wales (series 1) 10 (2): 141 - 144, pl. 19." type="journal article" year="1885">Miklouho-Maclay, 1885b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="60.[869,1437,197,246]" pageId="60" pageNumber="337">
<emphasis box="[869,1162,198,219]" italics="true" pageId="60" pageNumber="337">Proc. Linn. Soc. N.S.W. (ser. 1)</emphasis>
10(2): 141, pl. 19, figs 111. (
<date box="[907,1036,224,246]" pageId="60" pageNumber="337" value="1885-07-31">31 July 1885</date>
).
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="337" type="description">
<paragraph blockId="60.[839,1293,267,292]" box="[839,1293,267,292]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,1013,267,292]" pageId="60" pageNumber="337">Common name</emphasis>
. New
<collectingCountry box="[1083,1160,268,292]" name="Guinea-Bissau" pageId="60" pageNumber="337">Guinea</collectingCountry>
Pademelon.
</paragraph>
<paragraph blockId="60.[839,1466,313,366]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,1017,313,337]" pageId="60" pageNumber="337">Current name</emphasis>
.
<taxonomicName authority="(Ramsay, 1877 e)" baseAuthorityName="Ramsay" baseAuthorityYear="1877" box="[1037,1460,313,337]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="60" pageNumber="337" phylum="Chordata" rank="species" species="browni">
<emphasis box="[1037,1246,313,337]" italics="true" pageId="60" pageNumber="337">Thylogale browni</emphasis>
(Ramsay, 1877e)
</taxonomicName>
, following
<bibRefCitation author="Flannery, T" box="[949,1125,342,366]" pageId="60" pageNumber="337" pagination="7 - 23" refId="ref128748" refString="Flannery, T. F. 1992. Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy 15: 7 - 23." type="journal article" year="1992">Flannery (1992)</bibRefCitation>
.
</paragraph>
<paragraph blockId="60.[837,1467,387,500]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,850,387,411]" pageId="60" pageNumber="337">?</emphasis>
<materialsCitation collectingDate="1909-02-04" collectorName="M Register" country="Guinea-Bissau" location="New" pageId="60" pageNumber="337" specimenCount="1" typeStatus="syntype">
<emphasis bold="true" box="[850,944,387,411]" pageId="60" pageNumber="337">
<typeStatus box="[850,944,387,411]" pageId="60" pageNumber="337">Syntype</typeStatus>
</emphasis>
. M.2031, unsexed study skin with extracted skull, entered in the
<collectorName box="[1005,1132,417,441]" pageId="60" pageNumber="337">M Register</collectorName>
as “?
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C9EFE4AFB369685" box="[1196,1251,417,441]" country="Guinea-Bissau" name="New" pageId="60" pageNumber="337">New</location>
<collectingCountry box="[1262,1346,417,441]" name="Guinea-Bissau" pageId="60" pageNumber="337">Guinea</collectingCountry>
”, received from Macleay Museum Committee in 1907 and registered
<date box="[840,1020,476,500]" pageId="60" pageNumber="337" value="1909-02-04">
<collectingDate box="[840,1020,476,500]" pageId="60" pageNumber="337" value="1909-02-04">4 February 1909</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="60.[837,1469,521,721]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,953,521,545]" pageId="60" pageNumber="337">Condition</emphasis>
. Cranium and dentaries largely intact but bone porous and crumbling; both dentaries fused to cranium. Cranium: most incisors have disintegrated and are lost; soft palate adhered to palate; hole in anterior of braincase roof and much of roof has been crushed inward. Study skin intact and in reasonable condition; claws of both manus corroded by salt; claw sheaths missing from left pes digits 4 and 6.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="337" type="materials_examined">
<paragraph blockId="60.[839,1467,742,796]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,984,742,766]" pageId="60" pageNumber="337">Type locality</emphasis>
. “
<materialsCitation country="Papua New Guinea" location="Finisterre Range" pageId="60" pageNumber="337" specimenCount="1" stateProvince="Madang">
Macleay coast” = north of
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091D38FD0DFA6F95C3" box="[1290,1466,742,767]" country="Papua New Guinea" name="Finisterre Range" pageId="60" pageNumber="337" stateProvince="Madang">Finisterre Range</location>
and east of
<collectingRegion box="[957,1047,772,796]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="337">Madang</collectingRegion>
,
<collectingRegion box="[1057,1248,772,796]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="337">Madang Province</collectingRegion>
,
<collectingCountry box="[1256,1464,772,796]" name="Papua New Guinea" pageId="60" pageNumber="337">Papua New Guinea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="61" lastPageNumber="338" pageId="60" pageNumber="337" type="discussion">
<paragraph blockId="60.[837,1469,817,2014]" pageId="60" pageNumber="337">
<emphasis bold="true" box="[839,961,817,841]" pageId="60" pageNumber="337">Comments</emphasis>
.
<materialsCitation country="Guinea-Bissau" county="Macleay Museum" location="The" municipality="The" pageId="60" pageNumber="337" specimenCount="1" typeStatus="syntype">
We cannot exclude the possibly that M.2031 is a
<typeStatus box="[884,965,847,871]" pageId="60" pageNumber="337">syntype</typeStatus>
but on balance it possibly is not. Although the dimensions of the skin and skull match those provided for
<specimenCount pageId="60" pageNumber="337" type="generic">one specimen</specimenCount>
in Miklouho-Maclays description (see below), the condition of the skull is inconsistent with the skull figured in the original description.
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C09FC2FFBB194E0" box="[1083,1124,964,988]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="The" pageId="60" pageNumber="337">The</location>
skull is badly corroded, encrusted with deposits and partly cleaned, in contrast to the cleaned skull illustrated by Miklouho-Maclay. The corroded skull is typical of specimens obtained from
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091CEEFBF7FA579308" box="[1244,1410,1052,1076]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Andrew Goldie" pageId="60" pageNumber="337">Andrew Goldie</location>
, who as far as we are aware, is the only collector of that period to store
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091BB6FBBCFC639352" box="[900,950,1111,1134]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="New" pageId="60" pageNumber="337">New</location>
<collectingCountry box="[959,1035,1110,1134]" name="Guinea-Bissau" pageId="60" pageNumber="337">Guinea</collectingCountry>
material in “brine” rather than alcohol.
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091B75FB9FFCA693B0" box="[839,883,1140,1164]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="One" pageId="60" pageNumber="337">One</location>
possibility is that the
<typeStatus box="[1111,1192,1140,1164]" pageId="60" pageNumber="337">syntype</typeStatus>
skins were initially stored in alcohol by
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091BE0FB7AFB459395" box="[978,1168,1169,1193]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Miklouho-Maclay" pageId="60" pageNumber="337">Miklouho-Maclay</location>
and later transferred to brine barrels with
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091BF3FB45FBD293FA" box="[961,1031,1198,1222]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Goldie" pageId="60" pageNumber="337">Goldie</location>
material at the
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091CAFFB45FA8293FA" box="[1181,1367,1198,1222]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Macleay Museum" pageId="60" pageNumber="337">Macleay Museum</location>
, although this seems unlikely to us.
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C74FB27FB4693D8" box="[1094,1171,1228,1252]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Further" pageId="60" pageNumber="337">Further</location>
, it seems unlikely that having extracted the brine corroded skull,
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C80FB02FAA5923D" box="[1202,1392,1257,1281]" country="Guinea-Bissau" county="Macleay Museum" municipality="The" name="Miklouho-Maclay" pageId="60" pageNumber="337">Miklouho-Maclay</location>
would have subsequently returned it to the preserving solution.
<collectingMunicipality box="[1426,1467,1286,1310]" pageId="60" pageNumber="337">The</collectingMunicipality>
documentation associated with the transfer of these specimens from the
<collectingCounty box="[939,1132,1345,1369]" pageId="60" pageNumber="337">Macleay Museum</collectingCounty>
in 1907 makes no mention of skulls, but this might have been an oversight
</materialsCitation>
.
</paragraph>
<paragraph blockId="60.[837,1469,817,2014]" pageId="60" pageNumber="337">
In his original account Miklouho-Maclay stated that he obtained
<specimenCount box="[941,1102,1433,1457]" pageId="60" pageNumber="337" type="generic">two specimens</specimenCount>
only, both males of similar body size. Evidently he provided illustrations of both
<typeStatus box="[1383,1467,1463,1487]" pageId="60" pageNumber="337">syntype</typeStatus>
skulls as revealed by his fig. 6 illustration of palatal ridges showing fully erupted rear molars, compared to incompletely erupted molars of the cleaned skull in his fig. 9. Miklouho- Maclay indicates that he shot the first specimen in 1871 and that his published illustrations and body measurements were taken of the second specimen the same day that it was killed by dogs in 1876. He states that both specimens are adult, but
<bibRefCitation author="Thomas, O" box="[837,1013,1697,1721]" pageId="60" pageNumber="337" refId="ref145409" refString="Thomas, O. 1888 a. Catalogue of the Marsupialia and Monotremata in the Collection of the British Museum (Natural History). London: British Museum (Natural History). xii + 401 pp. https: // doi. org / 10.5962 / bhl. title. 56181" type="book" year="1888">Thomas (1888a)</bibRefCitation>
regarded them as juvenile.
</paragraph>
<paragraph blockId="60.[837,1469,817,2014]" lastBlockId="61.[119,751,151,1994]" lastPageId="61" lastPageNumber="338" pageId="60" pageNumber="337">
<materialsCitation country="Papua New Guinea" county="The" lastPageId="61" lastPageNumber="338" location="Macleay Museum" municipality="Macleay's" pageId="60" pageNumber="337" specimenCount="1" typeStatus="syntype">
The
<specimenCount box="[915,1053,1727,1751]" pageId="60" pageNumber="337" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
, which are not listed by
<bibRefCitation author="Stanbury, P" box="[1298,1467,1726,1750]" pageId="60" pageNumber="337" pagination="462 - 463" refId="ref144751" refString="Stanbury, P. J. 1969. Type specimens in the Macleay Museum, University of Sydney. IV. Mammals. Proceedings of the Linnean Society of New South Wales 93 (3): 462 - 463." type="journal article" year="1969">Stanbury (1969)</bibRefCitation>
as being in the
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091BEDF937FB4891CF" box="[991,1181,1756,1780]" country="Papua New Guinea" county="The" municipality="Macleay's" name="Macleay Museum" pageId="60" pageNumber="337">Macleay Museum</location>
, do not appear to have been reported since the original description.A search of the MMUS collection by HEP and MMUS staff in 2016 failed to locate candidate specimens.
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C0FF8DFFAD69070" box="[1085,1283,1844,1868]" country="Papua New Guinea" county="The" municipality="Macleay's" name="Miklouho-Maclay" pageId="60" pageNumber="337">Miklouho-Maclay</location>
does not reveal whether either specimen reached
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF8573081BB2FF7CFAD9963B:8EAB6005FF8573091C90F8BAFB249055" box="[1186,1265,1873,1897]" country="Papua New Guinea" county="The" municipality="Macleay's" name="Sydney" pageId="60" pageNumber="337">Sydney</location>
after his field work in
<collectingCountry box="[864,1063,1902,1926]" name="Papua New Guinea" pageId="60" pageNumber="337">Papua New Guinea</collectingCountry>
but if so, it would likely have been held in his private collection given his habit of acknowledging specimens in
<collectingMunicipality box="[986,1093,1961,1985]" pageId="60" pageNumber="337">Macleays</collectingMunicipality>
private collection in his published works of other proposed mammal taxa.
<collectingCounty box="[1270,1311,1990,2014]" pageId="60" pageNumber="337">The</collectingCounty>
caption to his fig. 1 of the whole animal states that it was based on a sketch from life and a photograph of a stuffed specimen, implying that he did not necessarily have access to a study skin while preparing the description
</materialsCitation>
.
</paragraph>
<paragraph blockId="61.[119,751,151,1994]" pageId="61" pageNumber="338">
Although it is not known whether Miklouho-Maclays original specimens have survived and whether they left
<collectingCountry name="Australia" pageId="61" pageNumber="338">Australia</collectingCountry>
with him on returned to
<collectingCountry box="[412,481,327,351]" name="Russia" pageId="61" pageNumber="338">Russia</collectingCountry>
in 1886 (
<bibRefCitation author="Maclay, R" box="[574,717,327,352]" pageId="61" pageNumber="338" pagination="1846 - 1888" refId="ref137548" refString="Maclay, R. W. d. M. 1974. Mikluho-Maklai, Nicholai Nicholaievich (1846 - 1888). In Australian Dictionary of Biography, Canberra: National Centre of Biography, Australian National University. Online version, accessed 12 July 2017. http: // adb. anu. edu. au / biography / mikluho-maklai-nicholai-nicholaievich- 4198" type="book chapter" year="1974">Maclay, 1974</bibRefCitation>
), it is possible that the AM material contains one of his
<typeStatus box="[655,749,357,381]" pageId="61" pageNumber="338">syntypes</typeStatus>
of
<taxonomicName authorityName="Miklouho-Maclay" authorityYear="1885" box="[150,310,386,410]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="tibol">
<emphasis box="[150,310,386,410]" italics="true" pageId="61" pageNumber="338">Macropus tibol</emphasis>
</taxonomicName>
. The MMUS sent a number of containers of specimens in alcohol to the AM, which were registered on
<date box="[156,282,445,469]" pageId="61" pageNumber="338" value="1909-02-04">4 Feb 1909</date>
. Included were
<specimenCount box="[454,603,445,469]" pageId="61" pageNumber="338" type="generic">ten specimens</specimenCount>
of wallabies: M.2031,
<taxonomicName box="[221,411,474,498]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="browni">
<emphasis box="[221,411,474,498]" italics="true" pageId="61" pageNumber="338">Macropus browni</emphasis>
</taxonomicName>
; M.203233,
<taxonomicName baseAuthorityName="Schreber" baseAuthorityYear="1778" box="[568,749,474,498]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="brunii">
<emphasis box="[568,749,474,498]" italics="true" pageId="61" pageNumber="338">Thylogale brunii</emphasis>
</taxonomicName>
originally entered in the register as
<emphasis box="[498,696,503,527]" italics="true" pageId="61" pageNumber="338">
?
<taxonomicName box="[508,696,503,527]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="browni">Macropus browni</taxonomicName>
</emphasis>
, and M.203440,
<specimenCount box="[254,427,533,557]" pageId="61" pageNumber="338" type="generic">seven specimens</specimenCount>
originally entered as
<taxonomicName authorityName="Miklouho-Maclay" authorityYear="1885" class="Mammalia" family="Macropodidae" genus="Dorcopsis" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="macleayi">
<emphasis italics="true" pageId="61" pageNumber="338">Dorcopsis macleayi</emphasis>
</taxonomicName>
(now
<taxonomicName baseAuthorityName="Miklouho-Maclay" baseAuthorityYear="1885" box="[298,547,562,586]" class="Mammalia" family="Macropodidae" genus="Dorcopsulus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="macleayi">
<emphasis box="[298,547,562,586]" italics="true" pageId="61" pageNumber="338">Dorcopsulus macleayi</emphasis>
</taxonomicName>
). The original M Register entry states “possibly co-types” and “?New
<collectingCountry box="[655,735,591,615]" name="Guinea-Bissau" pageId="61" pageNumber="338">Guinea</collectingCountry>
”, meaning that cotypes are possibly included amongst the
<specimenCount pageId="61" pageNumber="338" type="generic">ten specimens</specimenCount>
. Given that
<taxonomicName baseAuthorityName="Miklouho-Maclay" baseAuthorityYear="1885" box="[353,584,650,674]" class="Mammalia" family="Macropodidae" genus="Dorcopsulus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="macleayi">
<emphasis box="[353,584,650,674]" italics="true" pageId="61" pageNumber="338">Dorcopsulus macleayi</emphasis>
</taxonomicName>
appears to have been described from a single specimen now in the MMUS (MMUS M381, see
<bibRefCitation author="Stanbury, P" box="[328,486,709,733]" pageId="61" pageNumber="338" pagination="462 - 463" refId="ref144751" refString="Stanbury, P. J. 1969. Type specimens in the Macleay Museum, University of Sydney. IV. Mammals. Proceedings of the Linnean Society of New South Wales 93 (3): 462 - 463." type="journal article" year="1969">Stanbury, 1969</bibRefCitation>
), any “co-types” are more likely to be amongst the specimens of
<taxonomicName authorityName="Gray" authorityYear="1837" box="[504,608,738,762]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="genus">
<emphasis box="[504,608,738,762]" italics="true" pageId="61" pageNumber="338">Thylogale</emphasis>
</taxonomicName>
: M.203133. The skulls in this series are badly decalcified by storage in brine rather than alcohol, as described by Miklouho-Maclay for material sent from Goldie in Macleays private collection.
</paragraph>
<paragraph blockId="61.[119,751,151,1994]" pageId="61" pageNumber="338">
The skull of M.2031 is decalcified but it closely resembles one of the skulls illustrated in the original description, being a young animal at the same stage of molar eruption. Further, M.2031 closely approximates the size of the skull illustrated by Miklouho-Maclay at nature size, taking into account likely distortion from the camera lucida drawing by Miklouho- Maclay, and likely shrinkage of the salt affected skull. Cranial measurements are not given in the original account but greatest skull length and zygomatic breadth, measured from the published hardcopy plate, are greater than M.2031: greatest skull length
<quantity box="[190,282,1149,1173]" metricMagnitude="-2" metricUnit="m" metricValue="8.03" pageId="61" pageNumber="338" unit="mm" value="80.3">80.3 mm</quantity>
vs 72.8; zygomatic breadth 44.6 vs
<quantity box="[650,745,1149,1173]" metricMagnitude="-2" metricUnit="m" metricValue="4.04" pageId="61" pageNumber="338" unit="mm" value="40.4">40.4 mm</quantity>
. However, unlike the skull illustrated in the original account to which M.2031 closely resembles, M.2031 retains the skin on the palate. Further, the mandible is now fused with the cranium.While this might suggest that M.2031 is not a
<typeStatus box="[663,745,1267,1291]" pageId="61" pageNumber="338">syntype</typeStatus>
, it is possible that the skin of the palate was loosened for the illustration, and that both the palate tissue and mandible have subsequently adhered to the salt affected skull. The study skin M.2301 is a close match with the dimensions given for
<specimenCount pageId="61" pageNumber="338" type="generic" typeStatus="syntype">one syntype</specimenCount>
in the original description considering the imprecise demarcation of the tail base in the study skin: nose tip to base of tail c.
<quantity box="[211,329,1471,1495]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" pageId="61" pageNumber="338" unit="in" value="13.5">13.5 inches</quantity>
vs.
<quantity box="[371,469,1471,1496]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="61" pageNumber="338" unit="in" value="13.0">13 inches</quantity>
for the
<typeStatus box="[550,634,1472,1496]" pageId="61" pageNumber="338">syntype</typeStatus>
; tail length
<quantity box="[121,260,1501,1525]" metricMagnitude="-1" metricUnit="m" metricValue="2.4765" metricValueMax="2.54" metricValueMin="2.413" pageId="61" pageNumber="338" unit="in" value="9.75" valueMax="10.0" valueMin="9.5">9.510 inches</quantity>
vs.
<quantity box="[296,397,1501,1525]" metricMagnitude="-1" metricUnit="m" metricValue="2.3622" pageId="61" pageNumber="338" unit="in" value="9.3">9.3 inches</quantity>
and ear length c.
<quantity box="[565,635,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="61" pageNumber="338" unit="mm" value="35.0">35 mm</quantity>
vs.
<quantity box="[672,745,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="61" pageNumber="338" unit="mm" value="34.0">34 mm</quantity>
.
</paragraph>
<paragraph blockId="61.[119,751,151,1994]" pageId="61" pageNumber="338">
Of the two
<taxonomicName authorityName="Gray" authorityYear="1837" box="[280,385,1530,1554]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="genus">
<emphasis box="[280,385,1530,1554]" italics="true" pageId="61" pageNumber="338">Thylogale</emphasis>
</taxonomicName>
specimens with associated skulls sent from MMUS, M.2031 is the only candidate
<typeStatus box="[638,719,1560,1584]" pageId="61" pageNumber="338">syntype</typeStatus>
of
<taxonomicName authorityName="Miklouho-Maclay" authorityYear="1885" box="[122,284,1589,1613]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="tibol">
<emphasis box="[122,284,1589,1613]" italics="true" pageId="61" pageNumber="338">Macropus tibol</emphasis>
</taxonomicName>
, if Miklouho-Maclays statement that both
<typeStatus box="[121,212,1619,1643]" pageId="61" pageNumber="338">syntypes</typeStatus>
are of similar size is correct. The other specimen, M.2032 (not listed by
<bibRefCitation author="Flannery, T" box="[357,535,1647,1672]" pageId="61" pageNumber="338" pagination="7 - 23" refId="ref128748" refString="Flannery, T. F. 1992. Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy 15: 7 - 23." type="journal article" year="1992">Flannery (1992))</bibRefCitation>
is a male study skin with a very badly decalcified and distorted skull that far exceeds the dimensions given by Miklouho-Maclay (nose tip to tail base =
<quantity box="[266,407,1735,1760]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.334" metricValueMin="5.08" pageId="61" pageNumber="338" unit="in" value="20.5" valueMax="21.0" valueMin="20.0">2021 inches</quantity>
vs
<quantity box="[445,549,1735,1760]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="61" pageNumber="338" unit="in" value="13.0">13 inches</quantity>
, tail length c.
<quantity metricMagnitude="-1" metricUnit="m" metricValue="3.429" pageId="61" pageNumber="338" unit="in" value="13.5">13.5 inches</quantity>
vs
<quantity box="[217,323,1765,1789]" metricMagnitude="-1" metricUnit="m" metricValue="2.3622" pageId="61" pageNumber="338" unit="in" value="9.3">9.3 inches</quantity>
). However, the stage of molar eruption and the presence of palatal skin ridges, which are firmly adhered to the skull unlike M.2031, are consistent with the illustration of one of the skulls in Miklouho-Maclays account but there is no strikingly individual feature such as damage that would secure the association.
</paragraph>
<paragraph blockId="61.[119,751,151,1994]" lastBlockId="61.[793,1422,151,264]" pageId="61" pageNumber="338">
The type series of the two other thylogales described by Miklouho-Maclay, both from New
<collectingCountry box="[480,558,1970,1994]" name="Guinea-Bissau" pageId="61" pageNumber="338">Guinea</collectingCountry>
, are accounted for:
<taxonomicName authority="Miklouho-Maclay, 1885 a" authorityName="Miklouho-Maclay" authorityYear="1885" box="[793,1272,151,176]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[793,991,152,176]" italics="true" pageId="61" pageNumber="338">Macropus gracilis</emphasis>
Miklouho-Maclay, 1885a
</taxonomicName>
(=
<taxonomicName baseAuthorityName="Schreber" baseAuthorityYear="1778" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="brunii">
<emphasis italics="true" pageId="61" pageNumber="338">Thylogale brunii</emphasis>
</taxonomicName>
) based on the
<typeStatus box="[1016,1108,181,205]" pageId="61" pageNumber="338">holotype</typeStatus>
skin and skull in the MMUS (
<bibRefCitation author="Stanbury, P" box="[802,967,210,234]" pageId="61" pageNumber="338" pagination="462 - 463" refId="ref144751" refString="Stanbury, P. J. 1969. Type specimens in the Macleay Museum, University of Sydney. IV. Mammals. Proceedings of the Linnean Society of New South Wales 93 (3): 462 - 463." type="journal article" year="1969">Stanbury, 1969</bibRefCitation>
) and
<taxonomicName authority="Miklouho-Maclay, 1885 a" authorityName="Miklouho-Maclay" authorityYear="1885" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="jukesii">
<emphasis box="[1030,1215,210,234]" italics="true" pageId="61" pageNumber="338">Macropus jukesii</emphasis>
Miklouho-Maclay, 1885a
</taxonomicName>
(=
<taxonomicName baseAuthorityName="Schreber" baseAuthorityYear="1778" box="[895,1072,240,264]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="61" pageNumber="338" phylum="Chordata" rank="species" species="brunii">
<emphasis box="[895,1072,240,264]" italics="true" pageId="61" pageNumber="338">Thylogale brunii</emphasis>
</taxonomicName>
), see above account.
</paragraph>
</subSubSection>
</treatment>
</document>