1601 lines
176 KiB
XML
1601 lines
176 KiB
XML
<document id="5694C1E8419854E61EF974343D3B1EB4" ID-DOI="10.5281/zenodo.276749" ID-GBIF-Dataset="ae745c92-3dd3-4a61-b9d4-49726d234367" ID-ISSN="1175-5326" ID-Zenodo-Dep="276749" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235161866" checkinUser="plazi" docAuthor="Neiber, Marco T., Vega-Luz, Ricardo, Vega-Luz, Rodolfo & Koenemann, Stefan" docDate="2011" docId="03DD87BDFA1EC26A8CF68E2D0535FC0F" docLanguage="en" docName="zt02757p046.pdf" docOrigin="Zootaxa 2757" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hemicycla (Adiverticula) diegoi Neiber, Vega-Luz, Vega-Luz & Koenemann, 2011, n. sp." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FFE4FFC5FA18C2678C618A3F0064FFD4" masterDocTitle="Hemicycla (Adiverticula) diegoi (Gastropoda: Pulmonata: Helicidae), a new species from Tenerife, Canary Islands, with a phylogenetic analysis of conchologically similar species in the genus Hemicycla Swainson, 1840" masterLastPageNumber="46" masterPageNumber="29" pageNumber="35" updateTime="1698239418362" updateUser="plazi">
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<mods:mods id="F28A360ABFFCF318443606A2C7A9CD4B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="411164CD849B46E0CE4E76AB24F82798">
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<mods:title id="0AEAED3D422B7A3A1F29EEB818E97CAD">Hemicycla (Adiverticula) diegoi (Gastropoda: Pulmonata: Helicidae), a new species from Tenerife, Canary Islands, with a phylogenetic analysis of conchologically similar species in the genus Hemicycla Swainson, 1840</mods:title>
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<mods:name id="90D5414D7C59D7A21D8213BDCCA1EC56" type="personal">
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<mods:role id="44E22197234D97E909CEDAE45C7151A5">
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<mods:roleTerm id="EA847A6CCCC3EE87E47C26CAF2EBE6AA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="57F8F1CF97EAAB5B6FDDA305E7014459">Neiber, Marco T.</mods:namePart>
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</mods:name>
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<mods:name id="0CE31622846902ED251FB8E88241FFE6" type="personal">
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<mods:roleTerm id="68DE971CBB64F04E6AF6A6803110BB07">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="615903FAA3499FA69BB2872096AE5859">Vega-Luz, Ricardo</mods:namePart>
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</mods:name>
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<mods:name id="08E1D54903BA30AC9B632BF616D34768" type="personal">
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<mods:roleTerm id="44894E27F02BDE738372224ECAC3ED20">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="28ED3ACF61D32AF9D19B55C9B7CF29CA">Vega-Luz, Rodolfo</mods:namePart>
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</mods:name>
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<mods:name id="FDA465D7367B1EAFB033F4D6E14A83FC" type="personal">
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<mods:roleTerm id="638D51F53232D4B25EB398D9DD1FE623">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="12A4F68E1AD566D09C6B0A38F0FCD830">Koenemann, Stefan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="79CFA72FCCC1C017907E1051E91D8F5E">text</mods:typeOfResource>
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<mods:title id="1353AF741AC6BA6BE5558A11C79359B4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="91EB880CE92D67B47DF52644E5928A6A">
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<mods:date id="EE88BEB4CC4B72D41B7D2D6786A6F550">2011</mods:date>
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<mods:detail id="AE7E2D96DD9F24349027AEA55ABB9A59" type="volume">
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<mods:number id="549795906BB89BE9BB62305734BFEC69">2757</mods:number>
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<mods:classification id="CDA481940ABB8A6F5FF1B9598782508D">journal article</mods:classification>
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<mods:identifier id="393BD17F3ED06097C06A04767F3BAC99" type="DOI">10.5281/zenodo.276749</mods:identifier>
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<mods:identifier id="4C561E84E68CBB1EF60C396048B2ED18" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5127B6A71A4BC14A18270FAA13C24BCC" type="Zenodo-Dep">276749</mods:identifier>
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<treatment id="03DD87BDFA1EC26A8CF68E2D0535FC0F" ID-DOI="http://doi.org/10.5281/zenodo.5615938" ID-GBIF-Taxon="119387965" ID-Zenodo-Dep="5615938" LSID="urn:lsid:plazi:treatment:03DD87BDFA1EC26A8CF68E2D0535FC0F" httpUri="http://treatment.plazi.org/id/03DD87BDFA1EC26A8CF68E2D0535FC0F" lastPageId="13" lastPageNumber="42" pageId="6" pageNumber="35">
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<subSubSection id="C36E6520FA1EC2618CF68E2D0204FBF8" box="[151,608,1042,1068]" pageId="6" pageNumber="35" type="nomenclature">
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<paragraph id="8BCB36ABFA1EC2618CF68E2D0204FBF8" blockId="6.[151,608,1042,1068]" box="[151,608,1042,1068]" pageId="6" pageNumber="35">
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||
<heading id="D08381C7FA1EC2618CF68E2D0204FBF8" bold="true" box="[151,608,1042,1068]" fontSize="11" level="1" pageId="6" pageNumber="35" reason="1">
|
||
<emphasis id="B900EAB9FA1EC2618CF68E2D0204FBF8" bold="true" box="[151,608,1042,1068]" pageId="6" pageNumber="35">
|
||
<taxonomicName id="4C744D28FA1EC2618CF68E2D0272FBF8" ID-CoL="3KMWB" box="[151,534,1042,1068]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="35" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA1EC2618CF68E2D017CFBF8" bold="true" box="[151,280,1042,1068]" italics="true" pageId="6" pageNumber="35">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA1EC2618D498E2D01D8FBF8" bold="true" box="[296,444,1042,1068]" italics="true" pageId="6" pageNumber="35">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA1EC2618DAC8E2D0272FBF8" bold="true" box="[461,534,1042,1068]" italics="true" pageId="6" pageNumber="35">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357C2FA1EC2618E7F8E2D0204FBF8" box="[542,608,1042,1068]" pageId="6" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA1EC2618CF68E6B02D0FA58" pageId="6" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8BCB36ABFA1EC2618CF68E6B0452FB5B" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CF68E6B015EFBB8" bold="true" box="[151,314,1107,1132]" pageId="6" pageNumber="35">
|
||
<typeStatus id="54CF8809FA1EC2618CF68E6B00B4FBB8" box="[151,208,1108,1132]" pageId="6" pageNumber="35">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
<collectingCountry id="F363763BFA1EC2618D278E6B01EDFBB8" box="[326,393,1108,1132]" name="Spain" pageId="6" pageNumber="35">Spain</collectingCountry>
|
||
, Canary Islands,Tenerife, Municipality of Buenavista del Norte, Teno, Los Roques, southwestern slopes, UTM (
|
||
<collectionCode id="ED65AE6EFA1EC2618DF78E48018CFB5B" box="[406,488,1143,1167]" pageId="6" pageNumber="35">MGRS</collectionCode>
|
||
) 28
|
||
<collectionCode id="ED65AE6EFA1EC2618E728E48022EFB5B" box="[531,586,1143,1167]" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="6" pageNumber="35">RCS</collectionCode>
|
||
1336, 100 to
|
||
<quantity id="4C8C9B4EFA1EC2618EBD8E480347FB5B" box="[732,803,1143,1167]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="35" unit="m" value="300.0">300 m</quantity>
|
||
above sea level (
|
||
<figureCitation id="134F2A2EFA1EC2618F858E48044DFB5B" box="[996,1065,1143,1167]" captionStart="FIGURE 2" captionStartId="8.[151,250,1607,1630]" captionTargetBox="[213,1364,613,1576]" captionTargetId="figure@8.[202,1371,609,1582]" captionTargetPageId="8" captionText="FIGURE 2. Distribution of Hemicycla (Adiverticula) diegoi n. sp. (․: modern record) and Hemicycla (Adiverticula) mascaensis (●: modern records, Ο: (sub-) fossil record) on Tenerife, Canary Island, Spain. The map is based on the topographical map sheets in the Series L, scale 1: 50000, Cartografía Militar de España for Tenerife, UTM (MGRS) coordinates, WGS- 84 ellipsoid and datum, zone 28 R, 100 x 100 km UTM-squares CS and DS with 5 km grid spacing. Symbols in the map correspond to 1 x 1 km UTM-squares." httpUri="https://zenodo.org/record/276751/files/figure.png" pageId="6" pageNumber="35">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA1EC2618CA68EA30481FA90" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CA68EA301B9FB60" bold="true" box="[199,477,1179,1204]" pageId="6" pageNumber="35">
|
||
<typeStatus id="54CF8809FA1EC2618CA68EA30166FB60" box="[199,258,1180,1204]" pageId="6" pageNumber="35">Type</typeStatus>
|
||
material (shells):
|
||
</emphasis>
|
||
<typeStatus id="54CF8809FA1EC2618D8B8EA30236FB60" box="[490,594,1180,1204]" pageId="6" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="134F2A2EFA1EC2618E068EA302D4FB60" box="[615,688,1180,1204]" captionStart="FIGURE 3" captionStartId="9.[151,250,1950,1973]" captionTargetBox="[151,1436,429,1934]" captionTargetId="figure@9.[151,1436,423,1935]" captionTargetPageId="9" captionText="FIGURE 3. Comparison of shells of Hemicycla (Adiverticula) diegoi n. sp. (holotype) (A – D) and Hemicycla (Adiverticula) mascaensis (E – H). A, E. Shell. B, F. Aperture. C, G. Last whorl sculpture. D, H. Protoconch. (A, E: scale bar = 10 mm; B, F: scale bar = 5 mm; C, D, G, H: scale bar = 2.5 mm)." httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
|
||
A–D):
|
||
<collectionCode id="ED65AE6EFA1EC2618F708EA30301FB60" LSID="urn:lsid:biocol.org:col:35275" box="[785,869,1180,1204]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="6" pageNumber="35">MNCN</collectionCode>
|
||
15.05/53.801 collected by R. Vega-Luz and B. Claveau at the
|
||
<typeStatus id="54CF8809FA1EC2618D5F8EFE010AFB0D" box="[318,366,1217,1241]" pageId="6" pageNumber="35">type</typeStatus>
|
||
locality,
|
||
<date id="FFCA106BFA1EC2618DB78E800213FB03" box="[470,631,1215,1239]" pageId="6" pageNumber="35" value="2009-03-02">2 March 2009</date>
|
||
.
|
||
<typeStatus id="54CF8809FA1EC2618EE38EFE028BFB0D" box="[642,751,1217,1241]" pageId="6" pageNumber="35" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D72FD22FA1EC2618E9F8E8003FDFB03" box="[766,921,1215,1239]" pageId="6" pageNumber="35" type="generic">13 specimens</specimenCount>
|
||
) collected at the
|
||
<typeStatus id="54CF8809FA1EC26188388EFE04EDFB0D" box="[1113,1161,1217,1241]" pageId="6" pageNumber="35">type</typeStatus>
|
||
locality by R. Vega-Luz and B. Claveau,
|
||
<date id="FFCA106BFA1EC2618D388EDB019BFB28" box="[345,511,1252,1276]" pageId="6" pageNumber="35" value="2009-03-02">2 March 2009</date>
|
||
:
|
||
<collectionCode id="ED65AE6EFA1EC2618E6C8EDB0205FB28" LSID="urn:lsid:biocol.org:col:35275" box="[525,609,1252,1276]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="6" pageNumber="35">MNCN</collectionCode>
|
||
15.05/53.802 (
|
||
<typeStatus id="54CF8809FA1EC2618F6E8EDB0315FB28" box="[783,881,1252,1276]" pageId="6" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
1),
|
||
<collectionCode id="ED65AE6EFA1EC2618FC28EDB038BFB28" box="[931,1007,1252,1276]" pageId="6" pageNumber="35">TFMC</collectionCode>
|
||
MT424 (
|
||
<typeStatus id="54CF8809FA1EC261883F8EDB04A4FB28" box="[1118,1216,1252,1276]" pageId="6" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
2),
|
||
<collectionCode id="ED65AE6EFA1EC26188938EDB055AFB28" box="[1266,1342,1252,1276]" pageId="6" pageNumber="35">TFMC</collectionCode>
|
||
MT425 (
|
||
<typeStatus id="54CF8809FA1EC2618CFF8F380165FACB" box="[158,257,1287,1311]" pageId="6" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
3),
|
||
<collectionCode id="ED65AE6EFA1EC2618D528F38010FFACB" LSID="urn:lsid:biocol.org:col:34838" box="[307,363,1287,1311]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="35">SMF</collectionCode>
|
||
335127 (
|
||
<typeStatus id="54CF8809FA1EC2618DBA8F38022CFACB" box="[475,584,1287,1311]" pageId="6" pageNumber="35" type="paratype">paratypes</typeStatus>
|
||
4 and 5), FLT (
|
||
<typeStatus id="54CF8809FA1EC2618F668F380310FACB" box="[775,884,1287,1311]" pageId="6" pageNumber="35" type="paratype">paratypes</typeStatus>
|
||
6 and 7), GMN 1944-XXVII-Hel (
|
||
<typeStatus id="54CF8809FA1EC26189708F380517FACB" box="[1297,1395,1287,1311]" pageId="6" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
8), GMN 1945-XXVII-Hel (
|
||
<typeStatus id="54CF8809FA1EC2618DD18F130277FA90" box="[432,531,1324,1348]" pageId="6" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
9), MRV (
|
||
<typeStatus id="54CF8809FA1EC2618EEA8F13029CFA90" box="[651,760,1324,1348]" pageId="6" pageNumber="35" type="paratype">paratypes</typeStatus>
|
||
10 and 11) and CBC (
|
||
<typeStatus id="54CF8809FA1EC2618F928F130404FA90" box="[1011,1120,1324,1348]" pageId="6" pageNumber="35" type="paratype">paratypes</typeStatus>
|
||
12 and 13).
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA1EC2618CA68F7002D0FA58" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CA68F7001D8FABC" bold="true" box="[199,444,1359,1384]" pageId="6" pageNumber="35">Additional material:</emphasis>
|
||
Five ethanol-preserved specimens, collected by R. Vega-Luz and B. Claveau at the
|
||
<typeStatus id="54CF8809FA1EC261890D8F6E05F8FABD" box="[1388,1436,1361,1385]" pageId="6" pageNumber="35">type</typeStatus>
|
||
locality,
|
||
<date id="FFCA106BFA1EC2618C968F4B01F3FA58" box="[247,407,1396,1420]" pageId="6" pageNumber="35" value="2009-03-02">2 March 2009</date>
|
||
, are deposited in
|
||
<collectionCode id="ED65AE6EFA1EC2618E3B8F4A02D4FA58" box="[602,688,1397,1420]" pageId="6" pageNumber="35">MTNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA1EC2618CA68FA70342FA00" pageId="6" pageNumber="35" type="etymology">
|
||
<paragraph id="8BCB36ABFA1EC2618CA68FA70342FA00" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CA68FA70135FA64" bold="true" box="[199,337,1432,1456]" pageId="6" pageNumber="35">Etymology:</emphasis>
|
||
The epithet was chosen in honour of Diego Luz, grandfather of the second and third authors, and a patron of the arts and sciences in the Punta de Teno area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA1EC2618CA68FE003ADF9C9" pageId="6" pageNumber="35" type="distribution">
|
||
<paragraph id="8BCB36ABFA1EC2618CA68FE003ADF9C9" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CA68FE00197FA2C" bold="true" box="[199,499,1503,1528]" pageId="6" pageNumber="35">Distribution and habitat:</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54CF8809FA1EC2618E948FDE0341FA2D" box="[757,805,1505,1529]" pageId="6" pageNumber="35">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="134F2A2EFA1EC2618FF18FDF03B1FA2C" box="[912,981,1504,1528]" captionStart="FIGURE 2" captionStartId="8.[151,250,1607,1630]" captionTargetBox="[213,1364,613,1576]" captionTargetId="figure@8.[202,1371,609,1582]" captionTargetPageId="8" captionText="FIGURE 2. Distribution of Hemicycla (Adiverticula) diegoi n. sp. (․: modern record) and Hemicycla (Adiverticula) mascaensis (●: modern records, Ο: (sub-) fossil record) on Tenerife, Canary Island, Spain. The map is based on the topographical map sheets in the Series L, scale 1: 50000, Cartografía Militar de España for Tenerife, UTM (MGRS) coordinates, WGS- 84 ellipsoid and datum, zone 28 R, 100 x 100 km UTM-squares CS and DS with 5 km grid spacing. Symbols in the map correspond to 1 x 1 km UTM-squares." httpUri="https://zenodo.org/record/276751/files/figure.png" pageId="6" pageNumber="35">Fig. 2</figureCitation>
|
||
), where it lives in an area with Canarian succulent scrub vegetation of the “cardonal” and “tabaibal amargo”
|
||
<typeStatus id="54CF8809FA1EC2618FF08C3A03A1F9C9" box="[913,965,1541,1565]" pageId="6" pageNumber="35">type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA1EC26D8CA68C1805FFF83A" lastPageId="10" lastPageNumber="39" pageId="6" pageNumber="35" type="description">
|
||
<paragraph id="8BCB36ABFA1EC2618CA68C18049EF9B0" blockId="6.[151,1436,1107,2032]" pageId="6" pageNumber="35">
|
||
<emphasis id="B900EAB9FA1EC2618CA68C18013FF994" bold="true" box="[199,347,1575,1600]" pageId="6" pageNumber="35">Description:</emphasis>
|
||
The soft body of the animal has a finely wrinkled, greyish brown epidermis dorsally, with two indistinct, darker dorsolateral stripes (
|
||
<figureCitation id="134F2A2EFA1EC2618E5E8C7302E6F9B0" box="[575,642,1612,1636]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="6" pageNumber="35">Fig. 4</figureCitation>
|
||
A). The sole of the foot is of a pale brown to tan colour.
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA1EC2608CA48C4F027BFEB0" blockId="6.[151,1436,1107,2032]" lastBlockId="7.[151,1436,151,356]" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35">
|
||
The shell (
|
||
<figureCitation id="134F2A2EFA1EC2618D5A8C4F0119F95C" box="[315,381,1648,1672]" captionStart="FIGURE 3" captionStartId="9.[151,250,1950,1973]" captionTargetBox="[151,1436,429,1934]" captionTargetId="figure@9.[151,1436,423,1935]" captionTargetPageId="9" captionText="FIGURE 3. Comparison of shells of Hemicycla (Adiverticula) diegoi n. sp. (holotype) (A – D) and Hemicycla (Adiverticula) mascaensis (E – H). A, E. Shell. B, F. Aperture. C, G. Last whorl sculpture. D, H. Protoconch. (A, E: scale bar = 10 mm; B, F: scale bar = 5 mm; C, D, G, H: scale bar = 2.5 mm)." httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
|
||
A–D) is dextral, small for the genus, depressed, with 3.50 to 3.75 whorls, which are separated by a slightly impressed suture. The umbilicus is open. The background colour of the shell is yellowish to tan, becoming paler towards the umbilical region. Three reddish brown to dark brown spiral bands are present above the periphery of the shell (the first just below the suture, the second in the middle between the suture and the whorl periphery and the third just above the periphery) and one or two below the periphery of the shell (one just below the periphery and another, which is, if present at all, very faint, on the underside). The first four bands are interrupted by irregular whitish blotches or zigzag lines. The protoconch (
|
||
<figureCitation id="134F2A2EFA1EC2618F048D7703CEF8B4" box="[869,938,1864,1888]" captionStart="FIGURE 3" captionStartId="9.[151,250,1950,1973]" captionTargetBox="[151,1436,429,1934]" captionTargetId="figure@9.[151,1436,423,1935]" captionTargetPageId="9" captionText="FIGURE 3. Comparison of shells of Hemicycla (Adiverticula) diegoi n. sp. (holotype) (A – D) and Hemicycla (Adiverticula) mascaensis (E – H). A, E. Shell. B, F. Aperture. C, G. Last whorl sculpture. D, H. Protoconch. (A, E: scale bar = 10 mm; B, F: scale bar = 5 mm; C, D, G, H: scale bar = 2.5 mm)." httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
|
||
D) is of a uniform, brown colour, coarsely granulated, with 1.25 to 1.50 whorls. The teleoconch is sculptured by numerous, very fine radial riblets and granules, which are predominant on the riblets, but also present in the interspaces. The granulation is especially well developed on the first teleoconch whorls, becoming less prominent to almost absent on the body whorl (
|
||
<figureCitation id="134F2A2EFA1EC261894E8D8B0515F818" box="[1327,1393,1972,1996]" captionStart="FIGURE 3" captionStartId="9.[151,250,1950,1973]" captionTargetBox="[151,1436,429,1934]" captionTargetId="figure@9.[151,1436,423,1935]" captionTargetPageId="9" captionText="FIGURE 3. Comparison of shells of Hemicycla (Adiverticula) diegoi n. sp. (holotype) (A – D) and Hemicycla (Adiverticula) mascaensis (E – H). A, E. Shell. B, F. Aperture. C, G. Last whorl sculpture. D, H. Protoconch. (A, E: scale bar = 10 mm; B, F: scale bar = 5 mm; C, D, G, H: scale bar = 2.5 mm)." httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="6" pageNumber="35">Fig. 3</figureCitation>
|
||
C). Both the radial riblets and the granulation disappear toward the umbilical region, letting the lower shell surface appear somewhat glossy. On the lower surface of the shell very fine spiral striae and malleations are visible, which are almost completely obscured by the radial riblets and granules on the upper surface of the shell. The body whorl is rounded at the periphery, declining only slightly towards the aperture. The peristome is white, outwardly reflected, and oval, except for a weak longitudinal thickening at the columellar margin. The upper and columellar margins converge towards each other at their insertion points and are connected by a very thin, translucent callus. For shell measurements see
|
||
<tableCitation id="C6F60310FA1FC2608DB28B73027EFEB0" box="[467,538,332,356]" captionStart="TABLE 3" captionStartId="7.[151,235,404,425]" captionTargetBox="[151,1437,514,1405]" captionTargetPageId="7" captionText="TABLE 3. Shell measurements (in mm) and univariate statistics for Hemicycla (Adiverticula) mascaensis and H. (A.) diegoi n." httpUri="http://table.plazi.org/id/DF0B6623FA1FC2608CF68BAB05F8FE7D" pageId="7" pageNumber="36" tableUuid="DF0B6623FA1FC2608CF68BAB05F8FE7D">Tab. 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0B6623FA1FC2608CF68BAB05F8FE7D" ID-Table-UUID="DF0B6623FA1FC2608CF68BAB05F8FE7D" box="[151,1436,403,426]" httpUri="http://table.plazi.org/id/DF0B6623FA1FC2608CF68BAB05F8FE7D" pageId="7" pageNumber="36" targetBox="[151,1437,514,1405]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8BCB36ABFA1FC2608CF68BAB05F8FE7D" blockId="7.[151,1437,403,487]" box="[151,1436,403,426]" pageId="7" pageNumber="36">
|
||
<emphasis id="B900EAB9FA1FC2608CF68BAB0160FE7D" bold="true" box="[151,260,404,425]" pageId="7" pageNumber="36">TABLE 3.</emphasis>
|
||
Shell measurements (in mm) and univariate statistics for
|
||
<taxonomicName id="4C744D28FA1FC2608F2A8BAB04A1FE7E" box="[843,1221,404,426]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA1FC2608F2A8BAB03D3FE7E" box="[843,951,404,426]" italics="true" pageId="7" pageNumber="36">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA1FC2608FA78BAB0427FE7E" box="[966,1091,404,426]" italics="true" pageId="7" pageNumber="36">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA1FC26088338BAA04A1FE7E" box="[1106,1221,405,426]" italics="true" pageId="7" pageNumber="36">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA1FC26088978BAA05E6FE7E" box="[1270,1410,404,426]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="n." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA1FC26088978BAA056BFE7E" box="[1270,1295,405,426]" italics="true" pageId="7" pageNumber="36">H.</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA1FC260897C8BAB0548FE7D" box="[1309,1324,404,425]" italics="true" pageId="7" pageNumber="36">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA1FC26089238BAB05E6FE7E" box="[1346,1410,404,426]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357C2FA1FC26089E68BAB05F8FE7D" box="[1415,1436,404,425]" pageId="7" pageNumber="36">n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA1FC2608CF68B8C05F8FE1C" blockId="7.[151,1437,403,487]" box="[151,1436,434,456]" pageId="7" pageNumber="36">
|
||
<emphasis id="B900EAB9FA1FC2608CF68B8C00D1FE1C" bold="true" box="[151,181,435,456]" pageId="7" pageNumber="36">sp.</emphasis>
|
||
for all ten shell characters defined in
|
||
<figureCitation id="134F2A2EFA1FC2608E598B8D0210FE1C" box="[568,628,434,456]" captionStart="FIGURE 1" captionStartId="5.[151,250,1428,1451]" captionTargetBox="[156,1430,673,1402]" captionTargetId="figure@5.[151,1436,663,1403]" captionTargetPageId="5" captionText="FIGURE 1. Shell and genital measurements for comparison of Hemicycla (Adiverticula) diegoi n. sp. (A – C) and Hemicycla (Adiverticula) mascaensis (D – E). A, B. Dorsal and lateral view of shell. Dmax = maximum diameter, Dmin = minimum diameter, SH = shell height, SpH = spire height, AH = aperture height, AB = aperture breadth, BWH = body whorl height. C, D. Aperture of shell. AWmax = maximum aperture width, AWmin = minimum aperture width, CW = callus width (C, for shell with open and D, for shell with covered umbilicus). E. Genital system. BC = length of bursa copulatrix, BCD = length of bursa copulatrix duct, DV = length of distal vagina, PV = length of proximal vagina, P = penis length, DEp = length of distal part of epiphallus (from the penis retractor muscle to the proximal end of the penis), PEp = length of proximal part of epiphallus (from the insertion point of the vas deferens to the penis retractor muscle), F = length of flagellum, Vsd = length of vas deferens, MG = length of mucus gland, DS = length of dart sac, SOv = length of spermoviduct, FOv = length of free oviduct, AlG = length of albumen gland." httpUri="https://zenodo.org/record/276750/files/figure.png" pageId="7" pageNumber="36">Fig. 1</figureCitation>
|
||
(Mean = mean value, Min = minimal value, Max = maximal value, SD = stan-
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA1FC2608CF68BEE05F9FE33" blockId="7.[151,1437,403,487]" box="[151,1437,465,487]" pageId="7" pageNumber="36">dard deviation, t = Welch’s statistic, edf = effective degrees of freedom and p = onesided p-value from Student’s t-distribution).</paragraph>
|
||
<paragraph id="8BCB36ABFA1FC2608CFE883D0381FAA8" pageId="7" pageNumber="36">
|
||
<table id="F974C40BFA1F3D988CF6883D05F9FAA9" box="[151,1437,514,1405]" gridcols="5" gridrows="21" pageId="7" pageNumber="36">
|
||
<tr id="354434E9FA1F3D988CF6883D05F9FD96" box="[151,1437,514,578]" gridrow="0" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF6883D0248FD96" box="[151,556,514,578]" gridcol="0" gridrow="0" pageId="7" pageNumber="36">Shell charac- Descriptive statistics ter Taxon Mean</th>
|
||
<th id="76955D95FA1F3D988EEB883D02D2FD96" box="[650,694,514,578]" gridcol="1" gridrow="0" pageId="7" pageNumber="36">Min</th>
|
||
<th id="76955D95FA1F3D988F40883D032BFD96" box="[801,847,514,578]" gridcol="2" gridrow="0" pageId="7" pageNumber="36">Max</th>
|
||
<th id="76955D95FA1F3D988FD8883D0381FD96" box="[953,997,514,578]" gridcol="3" gridrow="0" pageId="7" pageNumber="36">SD</th>
|
||
<th id="76955D95FA1F3D988831883D05F9FD96" box="[1104,1437,514,578]" gridcol="4" gridrow="0" pageId="7" pageNumber="36">Welch’s t-test t edf p</th>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6886A05F9FDBF" box="[151,1437,597,619]" gridrow="1" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF6886A0248FDBF" box="[151,556,597,619]" gridcol="0" gridrow="1" pageId="7" pageNumber="36">
|
||
Dmax
|
||
<taxonomicName id="4C744D28FA1FC2608D57886901ACFDBF" box="[310,456,598,619]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D57886901ACFDBF" box="[310,456,598,619]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
17.1
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB886A02D2FDBF" box="[650,694,597,619]" gridcol="1" gridrow="1" pageId="7" pageNumber="36">15.2</td>
|
||
<td id="76955D95FA1F3D988F40886A032BFDBF" box="[801,847,597,619]" gridcol="2" gridrow="1" pageId="7" pageNumber="36">18.6</td>
|
||
<td id="76955D95FA1F3D988FD8886A0381FDBF" box="[953,997,597,619]" gridcol="3" gridrow="1" pageId="7" pageNumber="36">0.67</td>
|
||
<td id="76955D95FA1F3D988831886A05F9FDBF" box="[1104,1437,597,619]" gridcol="4" gridrow="1" pageId="7" pageNumber="36">10.25 17.65 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6884105F9FD41" box="[151,1437,638,661]" gridrow="2" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF688410248FD41" box="[151,556,638,661]" gridcol="0" gridrow="2" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D5788BF01F1FD41" box="[310,405,639,661]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D5788BF012CFD41" box="[310,328,640,661]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D34884001F1FD41" box="[341,405,639,661]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA884001B0FD40" bold="true" box="[411,468,639,660]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA884001B0FD40" box="[411,468,639,660]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
14.4
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB884102D2FD41" box="[650,694,638,661]" gridcol="1" gridrow="2" pageId="7" pageNumber="36">12.3</td>
|
||
<td id="76955D95FA1F3D988F408841032BFD41" box="[801,847,638,661]" gridcol="2" gridrow="2" pageId="7" pageNumber="36">15.6</td>
|
||
<td id="76955D95FA1F3D988FD888410381FD41" box="[953,997,638,661]" gridcol="3" gridrow="2" pageId="7" pageNumber="36">0.95</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6889705F9FD6A" box="[151,1437,680,702]" gridrow="3" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF688970248FD6A" box="[151,556,680,702]" gridcol="0" gridrow="3" pageId="7" pageNumber="36">
|
||
Dmin
|
||
<taxonomicName id="4C744D28FA1FC2608D57889601ACFD6A" box="[310,456,681,702]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D57889601ACFD6A" box="[310,456,681,702]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
14.3
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB889702D2FD6A" box="[650,694,680,702]" gridcol="1" gridrow="3" pageId="7" pageNumber="36">12.8</td>
|
||
<td id="76955D95FA1F3D988F408897032BFD6A" box="[801,847,680,702]" gridcol="2" gridrow="3" pageId="7" pageNumber="36">15.5</td>
|
||
<td id="76955D95FA1F3D988FD888970381FD6A" box="[953,997,680,702]" gridcol="3" gridrow="3" pageId="7" pageNumber="36">0.59</td>
|
||
<td id="76955D95FA1F3D988831889705F9FD6A" box="[1104,1437,680,702]" gridcol="4" gridrow="3" pageId="7" pageNumber="36">10.59 18.49 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF688EE05F9FD33" box="[151,1437,721,743]" gridrow="4" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF688EE0248FD33" box="[151,556,721,743]" gridcol="0" gridrow="4" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D5788ED01F1FD33" box="[310,405,721,743]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D5788ED012CFD33" box="[310,328,722,743]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D3488EE01F1FD33" box="[341,405,721,743]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA88ED01B0FD33" bold="true" box="[411,468,722,743]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA88ED01B0FD33" box="[411,468,722,743]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
12.1
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB88EE02D2FD33" box="[650,694,721,743]" gridcol="1" gridrow="4" pageId="7" pageNumber="36">10.7</td>
|
||
<td id="76955D95FA1F3D988F4088EE032BFD33" box="[801,847,721,743]" gridcol="2" gridrow="4" pageId="7" pageNumber="36">13.2</td>
|
||
<td id="76955D95FA1F3D988FD888EE0381FD33" box="[953,997,721,743]" gridcol="3" gridrow="4" pageId="7" pageNumber="36">0.76</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF688C505F9FCC4" box="[151,1437,762,784]" gridrow="5" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF688C50248FCC4" box="[151,556,762,784]" gridcol="0" gridrow="5" pageId="7" pageNumber="36">
|
||
SH
|
||
<taxonomicName id="4C744D28FA1FC2608D5788C401ACFCC4" box="[310,456,763,784]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D5788C401ACFCC4" box="[310,456,763,784]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
9.2
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB88C502D2FCC4" box="[650,694,762,784]" gridcol="1" gridrow="5" pageId="7" pageNumber="36">7.7</td>
|
||
<td id="76955D95FA1F3D988F4088C5032BFCC4" box="[801,847,762,784]" gridcol="2" gridrow="5" pageId="7" pageNumber="36">10.5</td>
|
||
<td id="76955D95FA1F3D988FD888C50381FCC4" box="[953,997,762,784]" gridcol="3" gridrow="5" pageId="7" pageNumber="36">0.57</td>
|
||
<td id="76955D95FA1F3D98883188C505F9FCC4" box="[1104,1437,762,784]" gridcol="4" gridrow="5" pageId="7" pageNumber="36">18.77 30.52 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6891C05F9FCEE" box="[151,1437,803,826]" gridrow="6" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF6891C0248FCEE" box="[151,556,803,826]" gridcol="0" gridrow="6" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D57891A01F1FCEE" box="[310,405,804,826]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D57891A012CFCEE" box="[310,328,805,826]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D34891B01F1FCEE" box="[341,405,804,826]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA891B01B0FCED" bold="true" box="[411,468,804,825]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA891B01B0FCED" box="[411,468,804,825]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
6.8
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB891C02D2FCEE" box="[650,694,803,826]" gridcol="1" gridrow="6" pageId="7" pageNumber="36">6.1</td>
|
||
<td id="76955D95FA1F3D988F40891C032BFCEE" box="[801,847,803,826]" gridcol="2" gridrow="6" pageId="7" pageNumber="36">7.3</td>
|
||
<td id="76955D95FA1F3D988FD8891C0381FCEE" box="[953,997,803,826]" gridcol="3" gridrow="6" pageId="7" pageNumber="36">0.40</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6897205F9FCB7" box="[151,1437,845,867]" gridrow="7" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF689720248FCB7" box="[151,556,845,867]" gridcol="0" gridrow="7" pageId="7" pageNumber="36">
|
||
SpH
|
||
<taxonomicName id="4C744D28FA1FC2608D57897101ACFCB7" box="[310,456,846,867]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D57897101ACFCB7" box="[310,456,846,867]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
2.4
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB897202D2FCB7" box="[650,694,845,867]" gridcol="1" gridrow="7" pageId="7" pageNumber="36">1.2</td>
|
||
<td id="76955D95FA1F3D988F408972032BFCB7" box="[801,847,845,867]" gridcol="2" gridrow="7" pageId="7" pageNumber="36">3.5</td>
|
||
<td id="76955D95FA1F3D988FD889720381FCB7" box="[953,997,845,867]" gridcol="3" gridrow="7" pageId="7" pageNumber="36">0.53</td>
|
||
<td id="76955D95FA1F3D988831897205F9FCB7" box="[1104,1437,845,867]" gridcol="4" gridrow="7" pageId="7" pageNumber="36">9.48 35.81 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6894805F9FC59" box="[151,1437,887,909]" gridrow="8" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF689480248FC59" box="[151,556,887,909]" gridcol="0" gridrow="8" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D57894701F1FC59" box="[310,405,887,909]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578947012CFC59" box="[310,328,888,909]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D34894801F1FC59" box="[341,405,887,909]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA894801B0FC58" bold="true" box="[411,468,887,908]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA894801B0FC58" box="[411,468,887,908]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1.4
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB894802D2FC59" box="[650,694,887,909]" gridcol="1" gridrow="8" pageId="7" pageNumber="36">1.0</td>
|
||
<td id="76955D95FA1F3D988F408948032BFC59" box="[801,847,887,909]" gridcol="2" gridrow="8" pageId="7" pageNumber="36">1.8</td>
|
||
<td id="76955D95FA1F3D988FD889480381FC59" box="[953,997,887,909]" gridcol="3" gridrow="8" pageId="7" pageNumber="36">0.32</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF6899F05F9FC62" box="[151,1437,928,950]" gridrow="9" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF6899F0248FC62" box="[151,556,928,950]" gridcol="0" gridrow="9" pageId="7" pageNumber="36">
|
||
BWH
|
||
<taxonomicName id="4C744D28FA1FC2608D57899E01ACFC62" box="[310,456,929,950]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D57899E01ACFC62" box="[310,456,929,950]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
6.8
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB899F02D2FC62" box="[650,694,928,950]" gridcol="1" gridrow="9" pageId="7" pageNumber="36">6.0</td>
|
||
<td id="76955D95FA1F3D988F40899F032BFC62" box="[801,847,928,950]" gridcol="2" gridrow="9" pageId="7" pageNumber="36">7.4</td>
|
||
<td id="76955D95FA1F3D988FD8899F0381FC62" box="[953,997,928,950]" gridcol="3" gridrow="9" pageId="7" pageNumber="36">0.31</td>
|
||
<td id="76955D95FA1F3D988831899F05F9FC62" box="[1104,1437,928,950]" gridcol="4" gridrow="9" pageId="7" pageNumber="36">19.25 29.14 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF689F605F9FC0B" box="[151,1437,969,991]" gridrow="10" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF689F60248FC0B" box="[151,556,969,991]" gridcol="0" gridrow="10" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D5789F501F1FC0B" box="[310,405,969,991]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D5789F5012CFC0B" box="[310,328,970,991]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D3489F601F1FC0B" box="[341,405,969,991]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA89F501B0FC0B" bold="true" box="[411,468,970,991]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA89F501B0FC0B" box="[411,468,970,991]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
5.4
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB89F602D2FC0B" box="[650,694,969,991]" gridcol="1" gridrow="10" pageId="7" pageNumber="36">5.1</td>
|
||
<td id="76955D95FA1F3D988F4089F6032BFC0B" box="[801,847,969,991]" gridcol="2" gridrow="10" pageId="7" pageNumber="36">5.8</td>
|
||
<td id="76955D95FA1F3D988FD889F60381FC0B" box="[953,997,969,991]" gridcol="3" gridrow="10" pageId="7" pageNumber="36">0.23</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF689CD05F9FBDC" box="[151,1437,1010,1032]" gridrow="11" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF689CD0248FBDC" box="[151,556,1010,1032]" gridcol="0" gridrow="11" pageId="7" pageNumber="36">
|
||
AB
|
||
<taxonomicName id="4C744D28FA1FC2608D5789CC01ACFBDC" box="[310,456,1011,1032]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D5789CC01ACFBDC" box="[310,456,1011,1032]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
7.8
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB89CD02D2FBDC" box="[650,694,1010,1032]" gridcol="1" gridrow="11" pageId="7" pageNumber="36">6.9</td>
|
||
<td id="76955D95FA1F3D988F4089CD032BFBDC" box="[801,847,1010,1032]" gridcol="2" gridrow="11" pageId="7" pageNumber="36">8.9</td>
|
||
<td id="76955D95FA1F3D988FD889CD0381FBDC" box="[953,997,1010,1032]" gridcol="3" gridrow="11" pageId="7" pageNumber="36">0.42</td>
|
||
<td id="76955D95FA1F3D98883189CD05F9FBDC" box="[1104,1437,1010,1032]" gridcol="4" gridrow="11" pageId="7" pageNumber="36">9.01 16.65 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68E2405F9FBE6" box="[151,1437,1051,1074]" gridrow="12" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF68E240248FBE6" box="[151,556,1051,1074]" gridcol="0" gridrow="12" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D578E2201F1FBE6" box="[310,405,1052,1074]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578E22012CFBE6" box="[310,328,1053,1074]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D348E2301F1FBE6" box="[341,405,1052,1074]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA8E2301B0FBE5" bold="true" box="[411,468,1052,1073]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA8E2301B0FBE5" box="[411,468,1052,1073]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
6.2
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8E2402D2FBE6" box="[650,694,1051,1074]" gridcol="1" gridrow="12" pageId="7" pageNumber="36">5.1</td>
|
||
<td id="76955D95FA1F3D988F408E24032BFBE6" box="[801,847,1051,1074]" gridcol="2" gridrow="12" pageId="7" pageNumber="36">7.8</td>
|
||
<td id="76955D95FA1F3D988FD88E240381FBE6" box="[953,997,1051,1074]" gridcol="3" gridrow="12" pageId="7" pageNumber="36">0.69</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68E7A05F9FB8F" box="[151,1437,1093,1115]" gridrow="13" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF68E7A0248FB8F" box="[151,556,1093,1115]" gridcol="0" gridrow="13" pageId="7" pageNumber="36">
|
||
AH
|
||
<taxonomicName id="4C744D28FA1FC2608D578E7901ACFB8F" box="[310,456,1094,1115]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D578E7901ACFB8F" box="[310,456,1094,1115]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
4.2
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8E7A02D2FB8F" box="[650,694,1093,1115]" gridcol="1" gridrow="13" pageId="7" pageNumber="36">3.2</td>
|
||
<td id="76955D95FA1F3D988F408E7A032BFB8F" box="[801,847,1093,1115]" gridcol="2" gridrow="13" pageId="7" pageNumber="36">5.3</td>
|
||
<td id="76955D95FA1F3D988FD88E7A0381FB8F" box="[953,997,1093,1115]" gridcol="3" gridrow="13" pageId="7" pageNumber="36">0.55</td>
|
||
<td id="76955D95FA1F3D9888318E7A05F9FB8F" box="[1104,1437,1093,1115]" gridcol="4" gridrow="13" pageId="7" pageNumber="36">-0.14 23.21 0.445</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68E5105F9FB51" box="[151,1437,1134,1157]" gridrow="14" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF68E510248FB51" box="[151,556,1134,1157]" gridcol="0" gridrow="14" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D578E4F01F1FB51" box="[310,405,1135,1157]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578E4F012CFB51" box="[310,328,1136,1157]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D348E5001F1FB51" box="[341,405,1135,1157]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA8E5001B0FB50" bold="true" box="[411,468,1135,1156]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA8E5001B0FB50" box="[411,468,1135,1156]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
4.2
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8E5102D2FB51" box="[650,694,1134,1157]" gridcol="1" gridrow="14" pageId="7" pageNumber="36">3.5</td>
|
||
<td id="76955D95FA1F3D988F408E51032BFB51" box="[801,847,1134,1157]" gridcol="2" gridrow="14" pageId="7" pageNumber="36">4.9</td>
|
||
<td id="76955D95FA1F3D988FD88E510381FB51" box="[953,997,1134,1157]" gridcol="3" gridrow="14" pageId="7" pageNumber="36">0.50</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68EA705F9FB7A" box="[151,1437,1176,1198]" gridrow="15" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF68EA70248FB7A" box="[151,556,1176,1198]" gridcol="0" gridrow="15" pageId="7" pageNumber="36">
|
||
Awmax
|
||
<taxonomicName id="4C744D28FA1FC2608D578EA601ACFB7A" box="[310,456,1177,1198]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D578EA601ACFB7A" box="[310,456,1177,1198]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
10.3
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8EA702D2FB7A" box="[650,694,1176,1198]" gridcol="1" gridrow="15" pageId="7" pageNumber="36">9.0</td>
|
||
<td id="76955D95FA1F3D988F408EA7032BFB7A" box="[801,847,1176,1198]" gridcol="2" gridrow="15" pageId="7" pageNumber="36">11.9</td>
|
||
<td id="76955D95FA1F3D988FD88EA70381FB7A" box="[953,997,1176,1198]" gridcol="3" gridrow="15" pageId="7" pageNumber="36">0.59</td>
|
||
<td id="76955D95FA1F3D9888318EA705F9FB7A" box="[1104,1437,1176,1198]" gridcol="4" gridrow="15" pageId="7" pageNumber="36">14.41 19.29 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68EFE05F9FB03" box="[151,1437,1217,1239]" gridrow="16" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF68EFE0248FB03" box="[151,556,1217,1239]" gridcol="0" gridrow="16" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D578EFD01F1FB03" box="[310,405,1217,1239]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578EFD012CFB03" box="[310,328,1218,1239]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D348EFE01F1FB03" box="[341,405,1217,1239]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA8EFD01B0FB03" bold="true" box="[411,468,1218,1239]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA8EFD01B0FB03" box="[411,468,1218,1239]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
7.5
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8EFE02D2FB03" box="[650,694,1217,1239]" gridcol="1" gridrow="16" pageId="7" pageNumber="36">5.9</td>
|
||
<td id="76955D95FA1F3D988F408EFE032BFB03" box="[801,847,1217,1239]" gridcol="2" gridrow="16" pageId="7" pageNumber="36">8.2</td>
|
||
<td id="76955D95FA1F3D988FD88EFE0381FB03" box="[953,997,1217,1239]" gridcol="3" gridrow="16" pageId="7" pageNumber="36">0.71</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68ED505F9FAD4" box="[151,1437,1258,1280]" gridrow="17" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF68ED50248FAD4" box="[151,556,1258,1280]" gridcol="0" gridrow="17" pageId="7" pageNumber="36">
|
||
Awmin
|
||
<taxonomicName id="4C744D28FA1FC2608D578ED401ACFAD4" box="[310,456,1259,1280]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D578ED401ACFAD4" box="[310,456,1259,1280]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
8.3
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8ED502D2FAD4" box="[650,694,1258,1280]" gridcol="1" gridrow="17" pageId="7" pageNumber="36">7.2</td>
|
||
<td id="76955D95FA1F3D988F408ED5032BFAD4" box="[801,847,1258,1280]" gridcol="2" gridrow="17" pageId="7" pageNumber="36">9.7</td>
|
||
<td id="76955D95FA1F3D988FD88ED50381FAD4" box="[953,997,1258,1280]" gridcol="3" gridrow="17" pageId="7" pageNumber="36">0.50</td>
|
||
<td id="76955D95FA1F3D9888318ED505F9FAD4" box="[1104,1437,1258,1280]" gridcol="4" gridrow="17" pageId="7" pageNumber="36">13.72 24.38 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68F2C05F9FAFE" box="[151,1437,1299,1322]" gridrow="18" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF68F2C0248FAFE" box="[151,556,1299,1322]" gridcol="0" gridrow="18" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D578F2A01F1FAFE" box="[310,405,1300,1322]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578F2A012CFAFE" box="[310,328,1301,1322]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D348F2B01F1FAFE" box="[341,405,1300,1322]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA8F2B01B0FAFD" bold="true" box="[411,468,1300,1321]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA8F2B01B0FAFD" box="[411,468,1300,1321]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
6.5
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8F2C02D2FAFE" box="[650,694,1299,1322]" gridcol="1" gridrow="18" pageId="7" pageNumber="36">5.5</td>
|
||
<td id="76955D95FA1F3D988F408F2C032BFAFE" box="[801,847,1299,1322]" gridcol="2" gridrow="18" pageId="7" pageNumber="36">7.0</td>
|
||
<td id="76955D95FA1F3D988FD88F2C0381FAFE" box="[953,997,1299,1322]" gridcol="3" gridrow="18" pageId="7" pageNumber="36">0.43</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68F0205F9FA87" box="[151,1437,1341,1363]" gridrow="19" pageId="7" pageNumber="36">
|
||
<th id="76955D95FA1F3D988CF68F020248FA87" box="[151,556,1341,1363]" gridcol="0" gridrow="19" pageId="7" pageNumber="36">
|
||
CW
|
||
<taxonomicName id="4C744D28FA1FC2608D578F0101ACFA87" box="[310,456,1342,1363]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="mascaensis">
|
||
<emphasis id="B900EAB9FA1FC2608D578F0101ACFA87" box="[310,456,1342,1363]" italics="true" pageId="7" pageNumber="36">H. mascaensis</emphasis>
|
||
</taxonomicName>
|
||
6.3
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8F0202D2FA87" box="[650,694,1341,1363]" gridcol="1" gridrow="19" pageId="7" pageNumber="36">5.2</td>
|
||
<td id="76955D95FA1F3D988F408F02032BFA87" box="[801,847,1341,1363]" gridcol="2" gridrow="19" pageId="7" pageNumber="36">7.9</td>
|
||
<td id="76955D95FA1F3D988FD88F020381FA87" box="[953,997,1341,1363]" gridcol="3" gridrow="19" pageId="7" pageNumber="36">0.56</td>
|
||
<td id="76955D95FA1F3D9888318F0205F9FA87" box="[1104,1437,1341,1363]" gridcol="4" gridrow="19" pageId="7" pageNumber="36">25.17 37.24 <0.001</td>
|
||
</tr>
|
||
<tr id="354434E9FA1F3D988CF68F5905F9FAA9" box="[151,1437,1382,1405]" gridrow="20" pageId="7" pageNumber="36" rowspan-4="1">
|
||
<th id="76955D95FA1F3D988CF68F590248FAA9" box="[151,556,1382,1405]" gridcol="0" gridrow="20" pageId="7" pageNumber="36">
|
||
<taxonomicName id="4C744D28FA1FC2608D578F5701F1FAA9" box="[310,405,1383,1405]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="36" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov.">
|
||
<emphasis id="B900EAB9FA1FC2608D578F57012CFAA9" box="[310,328,1384,1405]" italics="true" pageId="7" pageNumber="36">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA1FC2608D348F5801F1FAA9" box="[341,405,1383,1405]" italics="true" pageId="7" pageNumber="36">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA1FC2608DFA8F5801B0FAA8" bold="true" box="[411,468,1383,1404]" pageId="7" pageNumber="36">
|
||
<taxonomicNameLabel id="A23357C2FA1FC2608DFA8F5801B0FAA8" box="[411,468,1383,1404]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
3.5
|
||
</th>
|
||
<td id="76955D95FA1F3D988EEB8F5902D2FAA9" box="[650,694,1382,1405]" gridcol="1" gridrow="20" pageId="7" pageNumber="36">2.7</td>
|
||
<td id="76955D95FA1F3D988F408F59032BFAA9" box="[801,847,1382,1405]" gridcol="2" gridrow="20" pageId="7" pageNumber="36">4.0</td>
|
||
<td id="76955D95FA1F3D988FD88F590381FAA9" box="[953,997,1382,1405]" gridcol="3" gridrow="20" pageId="7" pageNumber="36">0.33</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA1FC2608CA48F8103D4F826" blockId="7.[151,1436,1470,2035]" pageId="7" pageNumber="36">
|
||
Genital system (four specimens dissected): The nomenclature for the genital system follows Alonso and Ibáñez (2007). Especially, the terms “proximal” and “distal” refer to the position of structures in the genital system in relation to the hermaphroditic gland (see also
|
||
<bibRefCitation id="EFE54B5AFA1FC2608E0B8C390334F9CA" author="Giusti" box="[618,848,1542,1566]" pageId="7" pageNumber="45" refString="Giusti, F. & Lepri, A. (1981) Aspetti morfologici ed etologici dell'accoppiamento in alcune specie della famiglia Helicidae (Gastropoda: Polmonata). Lavori della Societa Malacologica Italiana, 17 - 18, 11 - 71." type="book chapter" year="1981">Giusti & Lepri 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE54B5AFA1FC2608F3A8C39040CF9CA" author="Giusti" box="[859,1128,1542,1566]" pageId="7" pageNumber="45" refString="Giusti, F. & Andreini, S. (1988) Morphological and ethological aspects of mating in two species of the family Helicidae (Gastropoda: Pulmonata): Theba pisana (Muller) and Helix aperta Born. Monitore Zoologico Italiano (Nuova Serie), 22, 331 - 363." type="journal article" year="1988">Giusti & Andreini 1988</bibRefCitation>
|
||
). The hermaphroditic gland is whitish; its duct is strongly undulating at first, but becomes straighter towards its insertion point at the base of the large, curved, milky-white albumen gland. Spermoviduct strongly folded; 3.2 to 3.4 times the length of free oviduct. The bursa copulatrix is rather small and rounded, its stalk is somewhat longer than penis and epiphallus together (
|
||
<figureCitation id="134F2A2EFA1FC2608D638CA90120F97A" box="[258,324,1686,1710]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="7" pageNumber="36">Fig. 4</figureCitation>
|
||
B); there is no diverticulum. The dart sac of one specimen contained a dart. It is
|
||
<quantity id="4C8C9B4EFA1FC26088B18CA90550F97A" box="[1232,1332,1686,1710]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="36" unit="mm" value="3.1">3.10 mm</quantity>
|
||
long and almost straight, with a four-bladed shaft forming a simple cross in transverse section, and a well-developed crown with twelve cusps (
|
||
<figureCitation id="134F2A2EFA1FC2608D0F8CE101D5F922" box="[366,433,1758,1782]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="7" pageNumber="36">Fig. 4</figureCitation>
|
||
D). One of the mucus glands with two terminal branches, the other with only one branch. The male parts of the genital system are characterised by a rather short flagellum that is somewhat shorter than the stalk of the bursa copulatrix, and slightly longer than penis and epiphallus together. The epiphallus is slender, with the penis retractor muscle attached to the epiphallus in its distal third. The penis is short and distinctly swollen (
|
||
<figureCitation id="134F2A2EFA1FC260890F8D7500C1F852" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="7" pageNumber="36">Fig. 4</figureCitation>
|
||
B). In the lumen of the penis, two penial papillae are present (the proximal one being much bigger than the distal penial papilla). The “ring zone”, a protuberant ring at the base of the distal penial papilla in the everted penis, is visible from outside. Between the distal penial papilla and the atrium, the contact organ is developed as an oblong tubercle (
|
||
<figureCitation id="134F2A2EFA1FC2608D618DE50126F826" box="[256,322,2010,2034]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="7" pageNumber="36">Fig. 4</figureCitation>
|
||
C). For measurements of the genital system see
|
||
<tableCitation id="C6F60310FA1FC2608F028DE503C8F826" box="[867,940,2010,2034]" captionStart="TABLE 4" captionStartId="8.[151,235,152,173]" captionTargetBox="[151,1436,304,532]" captionTargetPageId="8" captionText="TABLE 4. Genital measurements (in mm) for Hemicycla (Adiverticula) mascaensis and Hemicycla (Adiverticula) diegoi n. sp. for all 14 genital characters defined in Fig. 1 (Mean = mean value, Min = minimal value, Max = maximal value and n = number of specimens measured). For explanations of the abbreviations of the parts of the genital system, see Fig. 1." httpUri="http://table.plazi.org/id/DF0B6623FA10C26F8CF68AA704B2FF3E" pageId="7" pageNumber="36" tableUuid="DF0B6623FA10C26F8CF68AA704B2FF3E">Tab. 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0B6623FA10C26F8CF68AA704B2FF3E" ID-Table-UUID="DF0B6623FA10C26F8CF68AA704B2FF3E" httpUri="http://table.plazi.org/id/DF0B6623FA10C26F8CF68AA704B2FF3E" pageId="8" pageNumber="37" targetBox="[151,1436,304,532]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BCB36ABFA10C26F8CF68AA704B2FF3E" blockId="8.[151,1436,152,284]" pageId="8" pageNumber="37">
|
||
<emphasis id="B900EAB9FA10C26F8CF68AA70160FF79" bold="true" box="[151,260,152,173]" pageId="8" pageNumber="37">TABLE 4.</emphasis>
|
||
Genital measurements (in mm) for
|
||
<taxonomicName id="4C744D28FA10C26F8E0C8AA70382FF7A" box="[621,998,152,174]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="37" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA10C26F8E0C8AA702BDFF7A" box="[621,729,152,174]" italics="true" pageId="8" pageNumber="37">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA10C26F8E898AA70301FF7A" box="[744,869,152,174]" italics="true" pageId="8" pageNumber="37">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA10C26F8F128AA60382FF7A" box="[883,998,153,174]" italics="true" pageId="8" pageNumber="37">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA10C26F88768AA7053AFF7A" box="[1047,1374,152,174]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="37" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA10C26F88768AA704E7FF7A" box="[1047,1155,152,174]" italics="true" pageId="8" pageNumber="37">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA10C26F88F38AA7056BFF7A" box="[1170,1295,152,174]" italics="true" pageId="8" pageNumber="37">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA10C26F897F8AA7053AFF7A" box="[1310,1374,152,174]" italics="true" pageId="8" pageNumber="37">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA10C26F89028AA705F8FF79" bold="true" box="[1379,1436,152,173]" pageId="8" pageNumber="37">
|
||
<taxonomicNameLabel id="A23357C2FA10C26F89028AA705F8FF79" box="[1379,1436,152,173]" pageId="8" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
for all 14 genital characters defined in Fig. 1 (Mean = mean value, Min = minimal value, Max = maximal value and n = number of specimens measured). For explanations of the abbreviations of the parts of the genital system, see Fig. 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA10C26E8CFE8B3804D3FE53" blockId="8.[151,1436,152,284]" lastBlockId="9.[151,1437,151,391]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
|
||
Taxon BC fOv DS DV P DEp PEp MG SOv AlG BCD F Vsd PV Mandible and radula: The sickle-shaped mandible (
|
||
<figureCitation id="134F2A2EFA10C26F8F788D3B033BF8C8" box="[793,863,1796,1820]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="8" pageNumber="37">Fig. 4</figureCitation>
|
||
E) is
|
||
<quantity id="4C8C9B4EFA10C26F8FCA8D3B040AF8C8" box="[939,1134,1796,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.205" metricValueMax="1.23" metricValueMin="1.18" pageId="8" pageNumber="37" unit="mm" value="1.205" valueMax="1.23" valueMin="1.18">1.18 to 1.23 mm</quantity>
|
||
wide, of a reddish-brown colour, odontognathous, i.e., with three to four projecting ridges (four specimens dissected). The radula (one specimen dissected) is of the Helicinae-type with approximately 120 transverse rows of teeth with the following formula: C + 11 L + 24 M (C = central tooth, L = lateral teeth and M = marginal teeth). The central tooth of the radula is smaller than the first lateral teeth, triangular in shape, with a rounded tip and anterobasal grooves, but without ectocones. The first lateral teeth are broader than the central tooth, and possess a weakly developed ectocone. Anterobasal grooves are also clearly visible (
|
||
<figureCitation id="134F2A2EFA10C26F8EC58DE3028EF820" box="[676,746,2012,2036]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="8" pageNumber="37">Fig. 4</figureCitation>
|
||
H). In the following lateral teeth the size of the mesocone decreases, whereas the ectocone increases in size to appear clearly differentiated in the extreme lateral teeth. An endocone is not or hardly delineated in the first lateral teeth, but becomes more evident in the following lateral teeth to form a well developed triangular cusp in the first marginal teeth (
|
||
<figureCitation id="134F2A2EFA11C26E8F898AE00449FF23" box="[1000,1069,223,247]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="9" pageNumber="38">Fig. 4</figureCitation>
|
||
G–H). The anterobasal grooves gradually become less distinct towards the extreme lateral teeth and are hardly visible or absent in the marginal teeth. The ectocone and the endocone in the first marginal teeth are triangular in shape, whereas the mesocone is rounded (
|
||
<figureCitation id="134F2A2EFA11C26E8D658B73012EFEB0" box="[260,330,332,356]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="9" pageNumber="38">Fig. 4</figureCitation>
|
||
G). In the last marginal teeth, the endocone and the mesocone are of similar size, the ectocone being noticeably smaller and sometimes with an additional cusp on its outer margin (
|
||
<figureCitation id="134F2A2EFA11C26E882F8B5004F4FE53" box="[1102,1168,367,391]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="9" pageNumber="38">Fig. 4</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA10C26F8CFE8B0E05EAFDC0" pageId="8" pageNumber="37">
|
||
<table id="F974C40BFA103D988CF68B0F05F8FDC0" box="[151,1436,304,532]" gridcols="11" gridrows="5" pageId="8" pageNumber="37">
|
||
<tr id="354434E9FA103D988CF68B0F05F8FEBB" box="[151,1436,304,367]" gridrow="0" pageId="8" pageNumber="37">
|
||
<th id="76955D95FA103D988CF68B0F01D0FEBB" box="[151,436,304,367]" gridcol="0" gridrow="0" pageId="8" pageNumber="37">
|
||
<emphasis id="B900EAB9FA10C26F8CFE8B0E017BFE92" box="[159,287,305,326]" italics="true" pageId="8" pageNumber="37">H. mascaen-</emphasis>
|
||
Mean 2.39
|
||
<emphasis id="B900EAB9FA10C26F8CFE8B6F00DEFEB1" box="[159,186,336,357]" italics="true" pageId="8" pageNumber="37">sis</emphasis>
|
||
Min 2.31
|
||
</th>
|
||
<th id="76955D95FA103D988DB28B0F019BFEBB" box="[467,511,304,367]" gridcol="1" gridrow="0" pageId="8" pageNumber="37">1.22 1.08</th>
|
||
<th id="76955D95FA103D988E7F8B0F022EFEBB" box="[542,586,304,367]" gridcol="2" gridrow="0" pageId="8" pageNumber="37">3.68 3.61</th>
|
||
<th id="76955D95FA103D988E0B8B0F02F3FEBB" box="[618,663,304,367]" gridcol="3" gridrow="0" pageId="8" pageNumber="37">1.15 1.12</th>
|
||
<th id="76955D95FA103D988ED48B0F0286FEBB" box="[693,738,304,367]" gridcol="4" gridrow="0" pageId="8" pageNumber="37">2.27 2.10</th>
|
||
<th id="76955D95FA103D988F608B0F0349FEBB" box="[769,813,304,367]" gridcol="5" gridrow="0" pageId="8" pageNumber="37">1.32 1.23</th>
|
||
<th id="76955D95FA103D988F2C8B0F031DFEBB" box="[845,889,304,367]" gridcol="6" gridrow="0" pageId="8" pageNumber="37">2.05 1.81</th>
|
||
<th id="76955D95FA103D988FF98B0F03A1FEBB" box="[920,965,304,367]" gridcol="7" gridrow="0" pageId="8" pageNumber="37">2.86 2.81</th>
|
||
<th id="76955D95FA103D988F858B0F0474FEBB" box="[996,1040,304,367]" gridcol="8" gridrow="0" pageId="8" pageNumber="37">9.31 9.25</th>
|
||
<th id="76955D95FA103D9888518B0F0491FEBB" box="[1072,1269,304,367]" gridcol="9" gridrow="0" pageId="8" pageNumber="37">15.28 11.87 8.83 15.22 11.79 8.76</th>
|
||
<th id="76955D95FA103D9889748B0F05F8FEBB" box="[1301,1436,304,367]" gridcol="10" gridrow="0" pageId="8" pageNumber="37">11.18 1.29 11.05 1.21</th>
|
||
</tr>
|
||
<tr id="354434E9FA103D988CF68BBD05F8FE4C" box="[151,1436,386,408]" gridrow="1" pageId="8" pageNumber="37">
|
||
<th id="76955D95FA103D988CF68BBD01D0FE4C" box="[151,436,386,408]" gridcol="0" gridrow="1" pageId="8" pageNumber="37">n = 3 Max 2.47</th>
|
||
<td id="76955D95FA103D988DB28BBD019BFE4C" box="[467,511,386,408]" gridcol="1" gridrow="1" pageId="8" pageNumber="37">1.31</td>
|
||
<td id="76955D95FA103D988E7F8BBD022EFE4C" box="[542,586,386,408]" gridcol="2" gridrow="1" pageId="8" pageNumber="37">3.76</td>
|
||
<td id="76955D95FA103D988E0B8BBD02F3FE4C" box="[618,663,386,408]" gridcol="3" gridrow="1" pageId="8" pageNumber="37">1.20</td>
|
||
<td id="76955D95FA103D988ED48BBD0286FE4C" box="[693,738,386,408]" gridcol="4" gridrow="1" pageId="8" pageNumber="37">2.40</td>
|
||
<td id="76955D95FA103D988F608BBD0349FE4C" box="[769,813,386,408]" gridcol="5" gridrow="1" pageId="8" pageNumber="37">1.41</td>
|
||
<td id="76955D95FA103D988F2C8BBD031DFE4C" box="[845,889,386,408]" gridcol="6" gridrow="1" pageId="8" pageNumber="37">2.24</td>
|
||
<td id="76955D95FA103D988FF98BBD03A1FE4C" box="[920,965,386,408]" gridcol="7" gridrow="1" pageId="8" pageNumber="37">2.93</td>
|
||
<td id="76955D95FA103D988F858BBD0474FE4C" box="[996,1040,386,408]" gridcol="8" gridrow="1" pageId="8" pageNumber="37">9.36</td>
|
||
<td id="76955D95FA103D9888518BBD0491FE4C" box="[1072,1269,386,408]" gridcol="9" gridrow="1" pageId="8" pageNumber="37">15.34 11.94 8.94</td>
|
||
<td id="76955D95FA103D9889748BBD05F8FE4C" box="[1301,1436,386,408]" gridcol="10" gridrow="1" pageId="8" pageNumber="37">11.31 1.37</td>
|
||
</tr>
|
||
<tr id="354434E9FA103D988CF68B9405F8FE16" box="[151,1436,427,450]" gridrow="2" pageId="8" pageNumber="37">
|
||
<th id="76955D95FA103D988CF68B9401D0FE16" box="[151,436,427,450]" gridcol="0" gridrow="2" pageId="8" pageNumber="37">
|
||
<taxonomicName id="4C744D28FA10C26F8CFE8B920099FE16" box="[159,253,428,450]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="37" phylum="Mollusca" rank="species" species="diegoi">
|
||
<emphasis id="B900EAB9FA10C26F8CFE8B9200D5FE16" box="[159,177,429,450]" italics="true" pageId="8" pageNumber="37">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA10C26F8CDF8B930099FE16" box="[190,253,428,450]" italics="true" pageId="8" pageNumber="37">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA10C26F8D628B93017CFE15" bold="true" box="[259,280,428,449]" pageId="8" pageNumber="37">n.</emphasis>
|
||
Mean 2.00
|
||
</th>
|
||
<td id="76955D95FA103D988DB28B94019BFE16" box="[467,511,427,450]" gridcol="1" gridrow="2" pageId="8" pageNumber="37">2.59</td>
|
||
<td id="76955D95FA103D988E7F8B94022EFE16" box="[542,586,427,450]" gridcol="2" gridrow="2" pageId="8" pageNumber="37">2.86</td>
|
||
<td id="76955D95FA103D988E0B8B9402F3FE16" box="[618,663,427,450]" gridcol="3" gridrow="2" pageId="8" pageNumber="37">1.01</td>
|
||
<td id="76955D95FA103D988ED48B940286FE16" box="[693,738,427,450]" gridcol="4" gridrow="2" pageId="8" pageNumber="37">2.50</td>
|
||
<td id="76955D95FA103D988F608B940349FE16" box="[769,813,427,450]" gridcol="5" gridrow="2" pageId="8" pageNumber="37">1.81</td>
|
||
<td id="76955D95FA103D988F2C8B94031DFE16" box="[845,889,427,450]" gridcol="6" gridrow="2" pageId="8" pageNumber="37">3.16</td>
|
||
<td id="76955D95FA103D988FF98B9403A1FE16" box="[920,965,427,450]" gridcol="7" gridrow="2" pageId="8" pageNumber="37">3.12</td>
|
||
<td id="76955D95FA103D988F858B940474FE16" box="[996,1040,427,450]" gridcol="8" gridrow="2" pageId="8" pageNumber="37">8.72</td>
|
||
<td id="76955D95FA103D9888518B940491FE16" box="[1072,1269,427,450]" gridcol="9" gridrow="2" pageId="8" pageNumber="37">14.58 10.07 8.04</td>
|
||
<td id="76955D95FA103D9889748B9405F8FE16" box="[1301,1436,427,450]" gridcol="10" gridrow="2" pageId="8" pageNumber="37">9.91 0.28</td>
|
||
</tr>
|
||
<tr id="354434E9FA103D988CF68BEA05F8FE3F" box="[151,1436,469,491]" gridrow="3" pageId="8" pageNumber="37">
|
||
<th id="76955D95FA103D988CF68BEA01D0FE3F" box="[151,436,469,491]" gridcol="0" gridrow="3" pageId="8" pageNumber="37">
|
||
<emphasis id="B900EAB9FA10C26F8CFE8BEA00D9FE3E" bold="true" box="[159,189,469,490]" pageId="8" pageNumber="37">sp.</emphasis>
|
||
Min 1.39
|
||
</th>
|
||
<td id="76955D95FA103D988DB28BEA019BFE3F" box="[467,511,469,491]" gridcol="1" gridrow="3" pageId="8" pageNumber="37">2.45</td>
|
||
<td id="76955D95FA103D988E7F8BEA022EFE3F" box="[542,586,469,491]" gridcol="2" gridrow="3" pageId="8" pageNumber="37">2.42</td>
|
||
<td id="76955D95FA103D988E0B8BEA02F3FE3F" box="[618,663,469,491]" gridcol="3" gridrow="3" pageId="8" pageNumber="37">0.90</td>
|
||
<td id="76955D95FA103D988ED48BEA0286FE3F" box="[693,738,469,491]" gridcol="4" gridrow="3" pageId="8" pageNumber="37">2.30</td>
|
||
<td id="76955D95FA103D988F608BEA0349FE3F" box="[769,813,469,491]" gridcol="5" gridrow="3" pageId="8" pageNumber="37">1.79</td>
|
||
<td id="76955D95FA103D988F2C8BEA031DFE3F" box="[845,889,469,491]" gridcol="6" gridrow="3" pageId="8" pageNumber="37">2.73</td>
|
||
<td id="76955D95FA103D988FF98BEA03A1FE3F" box="[920,965,469,491]" gridcol="7" gridrow="3" pageId="8" pageNumber="37">3.03</td>
|
||
<td id="76955D95FA103D988F858BEA0474FE3F" box="[996,1040,469,491]" gridcol="8" gridrow="3" pageId="8" pageNumber="37">8.11</td>
|
||
<td id="76955D95FA103D9888518BEA0491FE3F" box="[1072,1269,469,491]" gridcol="9" gridrow="3" pageId="8" pageNumber="37">14.38 9.45 7.53</td>
|
||
<td id="76955D95FA103D9889748BEA05F8FE3F" box="[1301,1436,469,491]" gridcol="10" gridrow="3" pageId="8" pageNumber="37">9.26 0.19</td>
|
||
</tr>
|
||
<tr id="354434E9FA103D988CF68BC105F8FDC0" box="[151,1436,510,532]" gridrow="4" pageId="8" pageNumber="37">
|
||
<th id="76955D95FA103D988CF68BC101D0FDC0" box="[151,436,510,532]" gridcol="0" gridrow="4" pageId="8" pageNumber="37">n = 4 Max 2.27</th>
|
||
<td id="76955D95FA103D988DB28BC1019BFDC0" box="[467,511,510,532]" gridcol="1" gridrow="4" pageId="8" pageNumber="37">2.75</td>
|
||
<td id="76955D95FA103D988E7F8BC1022EFDC0" box="[542,586,510,532]" gridcol="2" gridrow="4" pageId="8" pageNumber="37">3.52</td>
|
||
<td id="76955D95FA103D988E0B8BC102F3FDC0" box="[618,663,510,532]" gridcol="3" gridrow="4" pageId="8" pageNumber="37">1.14</td>
|
||
<td id="76955D95FA103D988ED48BC10286FDC0" box="[693,738,510,532]" gridcol="4" gridrow="4" pageId="8" pageNumber="37">2.78</td>
|
||
<td id="76955D95FA103D988F608BC10349FDC0" box="[769,813,510,532]" gridcol="5" gridrow="4" pageId="8" pageNumber="37">1.82</td>
|
||
<td id="76955D95FA103D988F2C8BC1031DFDC0" box="[845,889,510,532]" gridcol="6" gridrow="4" pageId="8" pageNumber="37">3.60</td>
|
||
<td id="76955D95FA103D988FF98BC103A1FDC0" box="[920,965,510,532]" gridcol="7" gridrow="4" pageId="8" pageNumber="37">3.25</td>
|
||
<td id="76955D95FA103D988F858BC10474FDC0" box="[996,1040,510,532]" gridcol="8" gridrow="4" pageId="8" pageNumber="37">8.90</td>
|
||
<td id="76955D95FA103D9888518BC10491FDC0" box="[1072,1269,510,532]" gridcol="9" gridrow="4" pageId="8" pageNumber="37">14.81 10.74 8.80</td>
|
||
<td id="76955D95FA103D9889748BC105F8FDC0" box="[1301,1436,510,532]" gridcol="10" gridrow="4" pageId="8" pageNumber="37">10.45 0.41</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF0B6623FA10C26F8CF68C780135F90C" httpUri="https://zenodo.org/record/276751/files/figure.png" pageId="8" pageNumber="37" targetBox="[213,1364,613,1576]" targetPageId="8">
|
||
<paragraph id="8BCB36ABFA10C26F8CF68C780135F90C" blockId="8.[151,1436,1607,1752]" pageId="8" pageNumber="37">
|
||
<emphasis id="B900EAB9FA10C26F8CF68C780177F988" bold="true" box="[151,275,1607,1630]" pageId="8" pageNumber="37">FIGURE 2.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C744D28FA10C26F8DD48C780298F989" box="[437,764,1607,1629]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="37" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA10C26F8DD48C780245F989" box="[437,545,1607,1629]" italics="true" pageId="8" pageNumber="37">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA10C26F8E518C7802C9F989" box="[560,685,1607,1629]" italics="true" pageId="8" pageNumber="37">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA10C26F8EDD8C780298F989" box="[700,764,1607,1629]" italics="true" pageId="8" pageNumber="37">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA10C26F8F638C77035FF989" bold="true" box="[770,827,1608,1629]" pageId="8" pageNumber="37">
|
||
<taxonomicNameLabel id="A23357C2FA10C26F8F638C77035FF989" box="[770,827,1608,1629]" pageId="8" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(․: modern record) and
|
||
<taxonomicName id="4C744D28FA10C26F88558C7800D6F9A8" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="37" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA10C26F88558C7804C4F989" box="[1076,1184,1607,1629]" italics="true" pageId="8" pageNumber="37">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA10C26F88CE8C780548F989" box="[1199,1324,1607,1629]" italics="true" pageId="8" pageNumber="37">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA10C26F895A8C7700D6F9A8" italics="true" pageId="8" pageNumber="37">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
(●: modern records, Ο: (sub-)fossil record) on Tenerife, Canary Island, Spain. The map is based on the topographical map sheets in the Series L, scale 1:50000, Cartografía Militar de España for Tenerife, UTM (MGRS) coordinates, WGS-84 ellipsoid and datum, zone 28R, 100 x 100 km UTM-squares CS and DS with 5 km grid spacing. Symbols in the map correspond to1 x 1 km UTM-squares.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6623FA11C26E8CF68DA102FFF825" httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="9" pageNumber="38" targetBox="[151,1436,429,1934]" targetPageId="9">
|
||
<paragraph id="8BCB36ABFA11C26E8CF68DA102FFF825" blockId="9.[151,1436,1950,2034]" pageId="9" pageNumber="38">
|
||
<emphasis id="B900EAB9FA11C26E8CF68DA10170F867" bold="true" box="[151,276,1950,1973]" pageId="9" pageNumber="38">FIGURE 3.</emphasis>
|
||
Comparison of shells of
|
||
<taxonomicName id="4C744D28FA11C26E8E7D8DA10302F860" box="[540,870,1950,1972]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="38" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA11C26E8E7D8DA102ECF860" box="[540,648,1950,1972]" italics="true" pageId="9" pageNumber="38">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA11C26E8EF88DA10372F860" box="[665,790,1950,1972]" italics="true" pageId="9" pageNumber="38">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA11C26E8F468DA10302F860" box="[807,870,1950,1972]" italics="true" pageId="9" pageNumber="38">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA11C26E8F0F8DA003CDF860" bold="true" box="[878,937,1951,1972]" pageId="9" pageNumber="38">
|
||
<taxonomicNameLabel id="A23357C2FA11C26E8F0F8DA003CDF860" box="[878,937,1951,1972]" pageId="9" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype) (
|
||
<emphasis id="B900EAB9FA11C26E884A8DA10438F867" bold="true" box="[1067,1116,1950,1971]" pageId="9" pageNumber="38">A–D</emphasis>
|
||
) and
|
||
<taxonomicName id="4C744D28FA11C26E88FB8DA1016EF807" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="38" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA11C26E88FB8DA10562F860" box="[1178,1286,1950,1972]" italics="true" pageId="9" pageNumber="38">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA11C26E89778DA105F7F860" box="[1302,1427,1950,1972]" italics="true" pageId="9" pageNumber="38">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA11C26E8CF68D81016EF807" box="[151,266,1982,2003]" italics="true" pageId="9" pageNumber="38">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B900EAB9FA11C26E8D788D82012EF806" bold="true" box="[281,330,1981,2002]" pageId="9" pageNumber="38">E–H</emphasis>
|
||
).
|
||
<emphasis id="B900EAB9FA11C26E8D3E8D8201F2F806" bold="true" box="[351,406,1981,2002]" pageId="9" pageNumber="38">A, E.</emphasis>
|
||
Shell.
|
||
<emphasis id="B900EAB9FA11C26E8DBC8D82026AF806" bold="true" box="[477,526,1981,2002]" pageId="9" pageNumber="38">B, F.</emphasis>
|
||
Aperture.
|
||
<emphasis id="B900EAB9FA11C26E8E1D8D8302D6F807" bold="true" box="[636,690,1980,2003]" pageId="9" pageNumber="38">C, G.</emphasis>
|
||
Last whorl sculpture.
|
||
<emphasis id="B900EAB9FA11C26E8FF48D8203ABF806" bold="true" box="[917,975,1981,2002]" pageId="9" pageNumber="38">D, H.</emphasis>
|
||
Protoconch. (
|
||
<emphasis id="B900EAB9FA11C26E883F8D8204F2F806" bold="true" box="[1118,1174,1981,2002]" pageId="9" pageNumber="38">A, E:</emphasis>
|
||
scale bar = 10 mm;
|
||
<emphasis id="B900EAB9FA11C26E89078D8205F8F806" bold="true" box="[1382,1436,1981,2002]" pageId="9" pageNumber="38">B, F:</emphasis>
|
||
scale bar = 5mm;
|
||
<emphasis id="B900EAB9FA11C26E8D2D8DE401A7F825" bold="true" box="[332,451,2011,2034]" pageId="9" pageNumber="38">C, D, G, H:</emphasis>
|
||
scale bar = 2.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6623FA12C26D8CF68CC90555F8BC" httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="10" pageNumber="39" targetBox="[151,1436,193,1760]" targetPageId="10">
|
||
<paragraph id="8BCB36ABFA12C26D8CF68CC90555F8BC" blockId="10.[151,1437,1782,1897]" pageId="10" pageNumber="39">
|
||
<emphasis id="B900EAB9FA12C26D8CF68CC90176F8D8" bold="true" box="[151,274,1782,1805]" pageId="10" pageNumber="39">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C744D28FA12C26D8D798CC8023BF8D9" box="[280,607,1783,1805]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA12C26D8D798CC801E0F8D9" box="[280,388,1783,1805]" italics="true" pageId="10" pageNumber="39">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA12C26D8DF28CC80274F8D9" box="[403,528,1783,1805]" italics="true" pageId="10" pageNumber="39">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA12C26D8E7E8CC8023BF8D9" box="[543,607,1783,1805]" italics="true" pageId="10" pageNumber="39">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA12C26D8E058CC802D8F8D8" bold="true" box="[612,700,1783,1804]" pageId="10" pageNumber="39">
|
||
<taxonomicNameLabel id="A23357C2FA12C26D8E058CC802F9F8D8" box="[612,669,1783,1804]" pageId="10" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
A.
|
||
</emphasis>
|
||
living animal from the type locality (scale bar = 10 mm).
|
||
<emphasis id="B900EAB9FA12C26D89648CC80578F8D8" bold="true" box="[1285,1308,1783,1804]" pageId="10" pageNumber="39">B.</emphasis>
|
||
Genital system (scale bar = 5 mm).
|
||
<emphasis id="B900EAB9FA12C26D8DF98D2A01D4F8F8" bold="true" box="[408,432,1813,1836]" pageId="10" pageNumber="39">C.</emphasis>
|
||
Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep).
|
||
<emphasis id="B900EAB9FA12C26D8F4E8D0B032CF89D" bold="true" box="[815,840,1844,1865]" pageId="10" pageNumber="39">D.</emphasis>
|
||
Dart (scale bar = 1 mm).
|
||
<emphasis id="B900EAB9FA12C26D882E8D0B0403F89D" bold="true" box="[1103,1127,1844,1865]" pageId="10" pageNumber="39">E.</emphasis>
|
||
Mandible (scale bar = 1 mm).
|
||
<emphasis id="B900EAB9FA12C26D8CF68D6C00A3F8BC" bold="true" box="[151,199,1875,1896]" pageId="10" pageNumber="39">F–H</emphasis>
|
||
. Radula teeth (scale bar = 10 μm): marginal teeth (
|
||
<emphasis id="B900EAB9FA12C26D8EAD8D6C02BFF8BC" bold="true" box="[716,731,1875,1896]" pageId="10" pageNumber="39">F</emphasis>
|
||
), outer lateral teeth (
|
||
<emphasis id="B900EAB9FA12C26D8FD18D6D03A0F8BD" bold="true" box="[944,964,1874,1897]" pageId="10" pageNumber="39">G</emphasis>
|
||
), central and inner lateral teeth (
|
||
<emphasis id="B900EAB9FA12C26D896E8D6C0547F8BC" bold="true" box="[1295,1315,1875,1896]" pageId="10" pageNumber="39">H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA12C26D8CA48DB005FFF83A" blockId="10.[151,1436,1935,2030]" pageId="10" pageNumber="39">
|
||
<emphasis id="B900EAB9FA12C26D8CA48DB00261F87C" bold="true" box="[197,517,1935,1960]" pageId="10" pageNumber="39">Comparison and remarks.</emphasis>
|
||
<taxonomicName id="4C744D28FA12C26D8E6E8DAF036FF87C" box="[527,779,1935,1960]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA12C26D8E6E8DAF02E3F87C" box="[527,647,1936,1960]" italics="true" pageId="10" pageNumber="39">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA12C26D8EFB8DAF02CFF87C" box="[666,683,1936,1960]" italics="true" pageId="10" pageNumber="39">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D8EA48DAF036FF87C" box="[709,779,1936,1960]" italics="true" pageId="10" pageNumber="39">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA12C26D8F748DAF0332F87C" bold="true" box="[789,854,1936,1960]" pageId="10" pageNumber="39">
|
||
<taxonomicNameLabel id="A23357C2FA12C26D8F748DAF0332F87C" box="[789,854,1936,1960]" pageId="10" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs anatomically from
|
||
<taxonomicName id="4C744D28FA12C26D88EC8DAE055CF87C" box="[1165,1336,1935,1960]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="inutilis" subGenus="H.">
|
||
<emphasis id="B900EAB9FA12C26D88EC8DAE04C5F87C" box="[1165,1185,1937,1960]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D88DD8DAE04B4F87C" box="[1212,1232,1937,1960]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D888B8DAF055CF87C" box="[1258,1336,1936,1960]" italics="true" pageId="10" pageNumber="39">inutilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D28FA12C26D89288DAE0177F81F" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="pouchadan" subGenus="H.">
|
||
<emphasis id="B900EAB9FA12C26D89288DAE0539F87C" box="[1353,1373,1937,1960]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D89168DAE05EFF87C" box="[1399,1419,1937,1960]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D8CF68D8C0177F81F" box="[151,275,1971,1995]" italics="true" pageId="10" pageNumber="39">pouchadan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D28FA12C26D8D408D8B0194F81F" box="[289,496,1971,1995]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="saponacea" subGenus="H.">
|
||
<emphasis id="B900EAB9FA12C26D8D408D8B0151F81F" box="[289,309,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D8D2C8D8B0105F81F" box="[333,353,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D8D198D8B0194F81F" box="[376,496,1972,1995]" italics="true" pageId="10" pageNumber="39">saponacea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D28FA12C26D8D9F8D8B02D3F81F" box="[510,695,1971,1995]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="berkeleii" subGenus="H.">
|
||
<emphasis id="B900EAB9FA12C26D8D9F8D8B0276F81F" box="[510,530,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D8E4B8D8B025AF81F" box="[554,574,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D8E348D8C02D3F81F" box="[597,695,1971,1995]" italics="true" pageId="10" pageNumber="39">berkeleii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B900EAB9FA12C26D8EA48D8B02BDF81F" box="[709,729,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D8E908D8B0361F81F" box="[753,773,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.) cf.
|
||
<emphasis id="B900EAB9FA12C26D8F218D8B03A2F81F" box="[832,966,1972,1995]" italics="true" pageId="10" pageNumber="39">paivanopsis</emphasis>
|
||
and
|
||
<taxonomicName id="4C744D28FA12C26D8F9D8D8B051DF81F" box="[1020,1401,1971,1995]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA12C26D8F9D8D8B0474F81F" box="[1020,1040,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D88498D8B0458F81F" box="[1064,1084,1972,1995]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D88358D8C051DF81F" box="[1108,1401,1971,1995]" italics="true" pageId="10" pageNumber="39">quadricincta quadricincta</emphasis>
|
||
</taxonomicName>
|
||
by the lack of a diverticulum on the stalk of the bursa copulatrix. Conchologically,
|
||
<taxonomicName id="4C744D28FA12C26D88368DE80562F83A" box="[1111,1286,2006,2030]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="39" phylum="Mollusca" rank="species" species="inutilis" subGenus="H.">
|
||
<emphasis id="B900EAB9FA12C26D88368DE8040FF83A" box="[1111,1131,2007,2030]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D88E98DE804F8F83A" box="[1160,1180,2007,2030]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA12C26D88D98DE90562F83A" box="[1208,1286,2006,2030]" italics="true" pageId="10" pageNumber="39">inutilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B900EAB9FA12C26D89268DE8053FF83A" box="[1351,1371,2007,2030]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA12C26D89168DE805EFF83A" box="[1399,1419,2007,2030]" italics="true" pageId="10" pageNumber="39">H</emphasis>
|
||
.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA13C26C8CF68AA70442FBCD" pageId="11" pageNumber="40" type="reference_group">
|
||
<paragraph id="8BCB36ABFA13C26C8CF68AA7048CFC5A" blockId="11.[151,1437,151,2027]" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C744D28FA13C26C8CF68AA70177FF64" ID-CoL="3KMX8" box="[151,275,152,176]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="pouchadan">
|
||
<emphasis id="B900EAB9FA13C26C8CF68AA70177FF64" box="[151,275,152,176]" italics="true" pageId="11" pageNumber="40">pouchadan</emphasis>
|
||
</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="4C744D28FA13C26C8DC58AA60227FF64" ID-CoL="3KMWB" box="[420,579,151,176]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8DC58AA601DCFF64" box="[420,440,153,176]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8DB38AA70187FF64" box="[466,483,152,176]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8D9C8AA70227FF64" box="[509,579,152,176]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8E2C8AA702EAFF64" bold="true" box="[589,654,152,176]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8E2C8AA702EAFF64" box="[589,654,152,176]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by their ribbed shells, and the latter also by its completely covered umbilicus.
|
||
<taxonomicName id="4C744D28FA13C26C8D758A8301ABFF07" ID-CoL="3KMW7" box="[276,463,186,211]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="berkeleii" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8D758A83014CFF07" box="[276,296,188,211]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8D208A830131FF07" box="[321,341,188,211]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8D0C8A8401ABFF07" box="[365,463,187,211]" italics="true" pageId="11" pageNumber="40">berkeleii</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C744D28FA13C26C8E078A830366FF07" ID-CoL="3KMWB" box="[614,770,186,211]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8E078A83021EFF07" box="[614,634,188,211]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8EF28A8402C0FF07" box="[659,676,187,211]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8EDD8A840366FF07" box="[700,770,187,211]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8F6B8A84032EFF07" bold="true" box="[778,842,187,211]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8F6B8A84032EFF07" box="[778,842,187,211]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by possessing a prominently keeled and much more coarsely granulated shell, and by the covered umbilicus. The shells of some forms of
|
||
<taxonomicName id="4C744D28FA13C26C88318AE0057BFF22" ID-CoL="3KMXC" box="[1104,1311,222,246]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="saponacea" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C88318AE00400FF22" box="[1104,1124,223,246]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C881A8AE004EBFF22" box="[1147,1167,223,246]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88C68AE0057BFF22" box="[1191,1311,223,246]" italics="true" pageId="11" pageNumber="40">saponacea</emphasis>
|
||
</taxonomicName>
|
||
are similar in size compared to
|
||
<taxonomicName id="4C744D28FA13C26C8D1B8B3D0271FECD" ID-CoL="3KMWB" box="[378,533,256,281]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8D1B8B3D01EAFECD" box="[378,398,258,281]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8DC78B3E01D3FECD" box="[422,439,257,281]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8DAE8B3E0271FECD" box="[463,533,257,281]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8E7D8B3E023FFECD" bold="true" box="[540,603,257,281]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8E7D8B3E023FFECD" box="[540,603,257,281]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but can be readily separated from the new species by its completely covered umbilicus, and by lacking irregular white blotches or stripes that interrupt the otherwise similar banding pattern of five darker brown bands.
|
||
<taxonomicName id="4C744D28FA13C26C8E7D8B79039BFE8A" ID-CoL="7TW86" box="[540,1023,326,351]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA13C26C8E7D8B7902F0FE8A" box="[540,660,326,350]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C8EC68B7702DFFE8B" box="[679,699,328,351]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8EB78B79039BFE8A" box="[726,1023,326,350]" italics="true" pageId="11" pageNumber="40">quadricincta quadricincta</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by its keeled, finely ribbed and larger shell.
|
||
<taxonomicName id="4C744D28FA13C26C8D878B56023AFE55" ID-CoL="7P4ZH" box="[486,606,361,385]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA13C26C8D878B56023AFE55" box="[486,606,361,385]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C8E0F8B5502E6FE55" box="[622,642,362,385]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.) cf.
|
||
<emphasis id="B900EAB9FA13C26C8EDD8B550326FE55" box="[700,834,362,385]" italics="true" pageId="11" pageNumber="40">paivanopsis</emphasis>
|
||
exhibits a colour pattern which is in all respects identical to that of
|
||
<taxonomicName id="4C744D28FA13C26C8D258BB20187FE70" ID-CoL="3KMWB" box="[324,483,396,420]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8D258BB2013CFE70" box="[324,344,397,420]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8D138BB301E7FE70" box="[370,387,396,420]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8DFC8BB30187FE70" box="[413,483,396,420]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8D8D8BB20249FE71" bold="true" box="[492,557,397,421]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8D8D8BB20249FE71" box="[492,557,397,421]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but possesses an angular to keeled last whorl.
|
||
<taxonomicName id="4C744D28FA13C26C88348BB30535FE70" ID-CoL="3KMWB" box="[1109,1361,396,420]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C88348BB304A9FE70" box="[1109,1229,396,420]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C88818BB30495FE70" box="[1248,1265,396,420]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C896A8BB30535FE70" box="[1291,1361,396,420]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C893B8BB205F8FE71" bold="true" box="[1370,1436,397,421]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C893B8BB205F8FE71" box="[1370,1436,397,421]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares the absence of the diverticulum on the stalk of the bursa copulatrix with
|
||
<taxonomicName id="4C744D28FA13C26C88448B8F04B1FE13" ID-CoL="3KMX9" box="[1061,1237,431,455]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="pouchet" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C88448B8F045DFE13" box="[1061,1081,432,455]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88338B900407FE13" box="[1106,1123,431,455]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C881D8B9004B1FE13" box="[1148,1237,431,455]" italics="true" pageId="11" pageNumber="40">pouchet</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C896F8B8F008EFE3E" ID-CoL="3KMWX" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C896F8B8F0546FE13" box="[1294,1314,432,455]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C895D8B900529FE13" box="[1340,1357,431,455]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C89048B8F008EFE3E" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
. Conchologically,
|
||
<taxonomicName id="4C744D28FA13C26C8DA28BEC0200FE3E" ID-CoL="3KMWB" box="[451,612,466,490]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8DA28BEC01B3FE3E" box="[451,471,467,490]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8D938BED0267FE3E" box="[498,515,466,490]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8E7F8BED0200FE3E" box="[542,612,466,490]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8E0F8BEC02D4FE3F" bold="true" box="[622,688,467,491]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8E0F8BEC02D4FE3F" box="[622,688,467,491]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C744D28FA13C26C8F2E8BEC0466FE3E" ID-CoL="3KMX9" box="[847,1026,466,490]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="pouchet" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F2E8BEC0307FE3E" box="[847,867,467,490]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F1C8BED03EAFE3E" box="[893,910,466,490]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FC88BED0466FE3E" box="[937,1026,466,490]" italics="true" pageId="11" pageNumber="40">pouchet</emphasis>
|
||
</taxonomicName>
|
||
by its much smaller size and open umbilicus. Morphologically,
|
||
<taxonomicName id="4C744D28FA13C26C8DBB8BC902C8FDD9" ID-CoL="3KMWX" box="[474,684,501,525]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8DBB8BC90192FDD9" box="[474,502,502,525]" italics="true" pageId="11" pageNumber="40">H.</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C8E648BCA0272FDD9" box="[517,534,501,525]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8E4C8BC902C8FDD9" box="[557,684,502,525]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
is closest to
|
||
<taxonomicName id="4C744D28FA13C26C8F5A8BC903B7FDD9" ID-CoL="3KMWB" box="[827,979,501,525]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F5A8BC9032BFDD9" box="[827,847,502,525]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F048BCA0312FDD9" box="[869,886,501,525]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FEC8BCA03B7FDD9" box="[909,979,501,525]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8FBB8BCA047CFDD9" bold="true" box="[986,1048,501,525]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8FBB8BCA047CFDD9" box="[986,1048,501,525]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
However, the shell of
|
||
<taxonomicName id="4C744D28FA13C26C89728BC9008EFDE4" ID-CoL="3KMWX" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C89728BC90543FDD9" box="[1299,1319,502,525]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C895F8BCA052BFDD9" box="[1342,1359,501,525]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C89048BC9008EFDE4" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
is larger, the apertural margins are less converging, the white blotches interrupting the brown spiral bands are absent or very faint, the protoconch is less coarsely granulated, the teleoconch sculpture of fine transverse riblets is more prominent, and the umbilicus is completely covered or rarely visible as a narrow slit. Compared to
|
||
<taxonomicName id="4C744D28FA13C26C89E088600174FD4D" ID-CoL="3KMWB" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C89E0886005F1FDA2" box="[1409,1429,607,630]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8CC188BE00D5FD4D" box="[160,177,641,665]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8CAB88BE0174FD4D" box="[202,272,641,665]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8D7988BE0133FD4D" bold="true" box="[280,343,641,665]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8D7988BE0133FD4D" box="[280,343,641,665]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C744D28FA13C26C8D0688BD0259FD4D" ID-CoL="3KMWX" box="[359,573,641,665]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8D0688BD011FFD4D" box="[359,379,642,665]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8DF588BE01C1FD4D" box="[404,421,641,665]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8DDC88BD0259FD4D" box="[445,573,642,665]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
possesses a somewhat angular aperture (
|
||
<figureCitation id="134F2A2EFA13C26C886F88BE0436FD4D" box="[1038,1106,641,665]" captionStart="FIGURE 3" captionStartId="9.[151,250,1950,1973]" captionTargetBox="[151,1436,429,1934]" captionTargetId="figure@9.[151,1436,423,1935]" captionTargetPageId="9" captionText="FIGURE 3. Comparison of shells of Hemicycla (Adiverticula) diegoi n. sp. (holotype) (A – D) and Hemicycla (Adiverticula) mascaensis (E – H). A, E. Shell. B, F. Aperture. C, G. Last whorl sculpture. D, H. Protoconch. (A, E: scale bar = 10 mm; B, F: scale bar = 5 mm; C, D, G, H: scale bar = 2.5 mm)." httpUri="https://zenodo.org/record/276752/files/figure.png" pageId="11" pageNumber="40">Fig. 3</figureCitation>
|
||
F). Furthermore, the shell of
|
||
<taxonomicName id="4C744D28FA13C26C8CF6889A0150FD68" ID-CoL="3KMWB" box="[151,308,676,700]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8CF6889A00CFFD68" box="[151,171,677,700]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8CA5889B00B1FD68" box="[196,213,676,700]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8C8F889B0150FD68" box="[238,308,676,700]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8D5C889B011AFD68" bold="true" box="[317,382,676,700]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8D5C889B011AFD68" box="[317,382,676,700]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is more depressed than that of
|
||
<taxonomicName id="4C744D28FA13C26C8E89889A03A4FD68" ID-CoL="3KMWX" box="[744,960,676,700]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8E89889A0298FD68" box="[744,764,677,700]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F74889B0342FD68" box="[789,806,676,700]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8F21889A03A4FD68" box="[832,960,677,700]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFE54B5AFA13C26C8FAE889B04D4FD68" box="[975,1200,676,700]" pageId="11" pageNumber="45" refString="Ibanez, M., Groh, K., Alonso, M. R. & Cavero, E. (1988) Revision of the genus Hemicycla Swainson, 1840 (Mollusca, Helicidae) from Tenerife: Adiverticula n. subgen. and description of three new taxa. Bulletin du Museum national d'Histoire naturelle, Paris, Section A, Zoologie, Biologie et Ecologie Animales, 4 e Serie, 10, 309 - 326." type="journal article">
|
||
Ibáñez
|
||
<emphasis id="B900EAB9FA13C26C8842889A0432FD68" box="[1059,1110,676,700]" italics="true" pageId="11" pageNumber="40">et al</emphasis>
|
||
. (1988)
|
||
</bibRefCitation>
|
||
state that the mucus glands are undivided in
|
||
<taxonomicName id="4C744D28FA13C26C8DC288F70211FD0B" ID-CoL="3KMWX" box="[419,629,711,735]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8DC288F701D3FD0B" box="[419,439,712,735]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8DAF88F801BBFD0B" box="[462,479,711,735]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8D9788F70211FD0B" box="[502,629,712,735]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
, which can be confirmed for the three dissected specimens in this study, whereas
|
||
<taxonomicName id="4C744D28FA13C26C8C9D88D401FEFCD6" ID-CoL="3KMWB" box="[252,410,745,770]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8C9D88D40174FCD6" box="[252,272,747,770]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8D4888D5015EFCD6" box="[297,314,746,770]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8D3588D501FEFCD6" box="[340,410,746,770]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8DC288D50180FCD6" bold="true" box="[419,484,746,770]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8DC288D50180FCD6" box="[419,484,746,770]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has one mucus gland with two terminal branches and the other simple. Otherwise, there are no substantial differences in the genital systems of both species (
|
||
<tableCitation id="C6F60310FA13C26C8F958933045BFCF0" box="[1012,1087,780,804]" captionStart="TABLE 4" captionStartId="8.[151,235,152,173]" captionTargetBox="[151,1436,304,532]" captionTargetPageId="8" captionText="TABLE 4. Genital measurements (in mm) for Hemicycla (Adiverticula) mascaensis and Hemicycla (Adiverticula) diegoi n. sp. for all 14 genital characters defined in Fig. 1 (Mean = mean value, Min = minimal value, Max = maximal value and n = number of specimens measured). For explanations of the abbreviations of the parts of the genital system, see Fig. 1." httpUri="http://table.plazi.org/id/DF0B6623FA10C26F8CF68AA704B2FF3E" pageId="11" pageNumber="40" tableUuid="DF0B6623FA10C26F8CF68AA704B2FF3E">Tab. 4</tableCitation>
|
||
,
|
||
<figureCitation id="134F2A2EFA13C26C882F893304F6FCF0" box="[1102,1170,780,804]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="11" pageNumber="40">Fig. 4</figureCitation>
|
||
B–C and
|
||
<figureCitation id="134F2A2EFA13C26C896A8933052BFCF0" box="[1291,1359,780,804]" captionStart="FIGURE 5" captionStartId="12.[151,250,1677,1700]" captionTargetBox="[164,1411,176,1651]" captionTargetId="figure@12.[164,1412,167,1651]" captionTargetPageId="12" captionText="FIGURE 5. Hemicycla (Adiverticula) mascaensis. A. Living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276754/files/figure.png" pageId="11" pageNumber="40">Fig. 5</figureCitation>
|
||
B–C). Moreover, there are no significant differences in the dart, radula and mandible of both species (
|
||
<figureCitation id="134F2A2EFA13C26C88DB89100498FC93" box="[1210,1276,815,839]" captionStart="FIGURE 4" captionStartId="10.[151,250,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@10.[151,1436,193,1761]" captionTargetPageId="10" captionText="FIGURE 4. Hemicycla (Adiverticula) diegoi n. sp. A. living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276753/files/figure.png" pageId="11" pageNumber="40">Fig. 4</figureCitation>
|
||
D–H and
|
||
<figureCitation id="134F2A2EFA13C26C890E891000C1FCBE" captionStart="FIGURE 5" captionStartId="12.[151,250,1677,1700]" captionTargetBox="[164,1411,176,1651]" captionTargetId="figure@12.[164,1412,167,1651]" captionTargetPageId="12" captionText="FIGURE 5. Hemicycla (Adiverticula) mascaensis. A. Living animal from the type locality (scale bar = 10 mm). B. Genital system (scale bar = 5 mm). C. Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep). D. Dart (scale bar = 1 mm). E. Mandible (scale bar = 1 mm). F – H. Radula teeth (scale bar = 10 μm): marginal teeth (F), outer lateral teeth (G), central and inner lateral teeth (H)." httpUri="https://zenodo.org/record/276754/files/figure.png" pageId="11" pageNumber="40">Fig. 5</figureCitation>
|
||
D–H).
|
||
<bibRefCitation id="EFE54B5AFA13C26C8C98896D01AAFCBE" author="Ibanez" box="[249,462,850,875]" pageId="11" pageNumber="45" refString="Ibanez, M, Siverio, F., Alonso, M. R. & Ponte-Lira, C. E. (2006) Two Canariella species (Gastropoda: Helicodea: Hygromiidae) endemic to the Northwest Tenerife (Canary Islands). Zootaxa, 1258, 33 - 45." type="journal article" year="2006">
|
||
Ibáñez
|
||
<emphasis id="B900EAB9FA13C26C8D2B896B011FFCBF" box="[330,379,851,875]" italics="true" pageId="11" pageNumber="40">et al</emphasis>
|
||
. (2006
|
||
</bibRefCitation>
|
||
: 43) report
|
||
<taxonomicName id="4C744D28FA13C26C8E2C896C038AFCBF" ID-CoL="3KMWX" box="[589,1006,850,875]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8E2C896C02A1FCBF" box="[589,709,851,875]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C8EB4896C0304FCBF" box="[725,864,851,875]" italics="true" pageId="11" pageNumber="40">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA13C26C8F0E896B038AFCBF" box="[879,1006,852,875]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
from the Teno Bajo area, without providing further explanation. This record is here considered to represent
|
||
<taxonomicName id="4C744D28FA13C26C8FCD894904C6FC5A" ID-CoL="3KMWB" box="[940,1186,885,910]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8FCD89490440FC5A" box="[940,1060,886,910]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA13C26C885589490428FC5A" box="[1076,1100,886,910]" italics="true" pageId="11" pageNumber="40">A.</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA13C26C883D894904C6FC5A" box="[1116,1186,886,910]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C88C88949048CFC5A" bold="true" box="[1193,1256,886,910]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C88C88949048CFC5A" box="[1193,1256,886,910]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA13C26C8CA689A70442FBCD" blockId="11.[151,1437,151,2027]" pageId="11" pageNumber="40">
|
||
<emphasis id="B900EAB9FA13C26C8CA689A701EBFC65" bold="true" box="[199,399,920,945]" pageId="11" pageNumber="40">Genetic analysis.</emphasis>
|
||
A fragment of 661 bp was obtained for COI, with an average nucleotide composition for the coding strand of 23.8% A, 43.2% T, 14% C and 18.9% G. Of a total of 661 bp, 429 (64.9%) were conserved sites, 59 (8.9%) singletons within the data set, and 173 (26.2%) parsimony informative sites. The COI fragments are relatively A-T-rich, as has been observed in other helicid taxa (
|
||
<bibRefCitation id="EFE54B5AFA13C26C8F588E3E047EFBCD" author="Elejalde" box="[825,1050,1025,1049]" lastPageId="11" lastPageNumber="45" pageId="11" pageNumber="44" refString="Elejalde, M. A., Munoz, B., Arrebola, J. R. & Gomez-Moliner, B. J. (2005) Phylogenetic relationships of Iberus gualtieranus and Iberus alonensis (Gastropoda: Helicidae) based on partial mitochondrial 16 S rRNA and COI gene sequences. Journal of Molluscan Studies, 71, 349 - 355." type="journal article" year="2005">
|
||
Elejalde
|
||
<emphasis id="B900EAB9FA13C26C8FFF8E3D03ABFBCD" box="[926,975,1025,1049]" italics="true" pageId="11" pageNumber="40">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6520FA13C26A8CA48E1B0535FC0F" lastPageId="13" lastPageNumber="42" pageId="11" pageNumber="40" type="description">
|
||
<paragraph id="8BCB36ABFA13C26C8CA48E1B04FFF9D6" blockId="11.[151,1437,151,2027]" pageId="11" pageNumber="40">
|
||
The Bayesian analysis reveals two major lineages (
|
||
<figureCitation id="134F2A2EFA13C26C8F648E1B032FFBE8" box="[773,843,1060,1084]" captionStart="FIGURE 6" captionStartId="13.[151,250,1858,1881]" captionTargetBox="[369,1224,1085,1814]" captionTargetId="figure@13.[356,1230,1079,1821]" captionTargetPageId="13" captionText="FIGURE 6. Bayesian tree based on COI sequences of 19 specimens from 10 different taxa in the genus Hemicycla and two outgroup taxa. Specimen abbreviations correspond to those listed in Tab. 2. The bold horizontal lines group the Hemicycla species in the two currently recognised subgenera Hemicycla s. str. and Adiverticula. Numbers at nodes indicate posterior probability values as percentages; branch lengths are measured in nucleotide substitutions per site (scale bar)." httpUri="https://zenodo.org/record/276755/files/figure.png" pageId="11" pageNumber="40">Fig. 6</figureCitation>
|
||
). One lineage, containing
|
||
<taxonomicName id="4C744D28FA13C26C88158E1A0540FBE8" box="[1140,1316,1060,1084]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="plicaria" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C88158E1A04ECFBE8" box="[1140,1160,1061,1084]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88C18E1A04D0FBE8" box="[1184,1204,1061,1084]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88AD8E1B0540FBE8" box="[1228,1316,1060,1084]" italics="true" pageId="11" pageNumber="40">plicaria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CF8809FA13C26C89558E1B0502FBE8" box="[1332,1382,1060,1084]" pageId="11" pageNumber="40">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C744D28FA13C26C8D398E7801B4FB8B" box="[344,464,1095,1119]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA13C26C8D398E7801B4FB8B" box="[344,464,1095,1119]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
</taxonomicName>
|
||
) and all the samples of species with a diverticulum on the stalk of the bursa copulatrix, is only weakly supported by a posterior probability value of 58. The other lineage, containing all samples of species without a diverticulum on the stalk of the bursa copulatrix (which correspond to
|
||
<taxonomicName id="4C744D28FA13C26C887E8EB204F3FB71" box="[1055,1175,1165,1189]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA13C26C887E8EB204F3FB71" box="[1055,1175,1165,1189]" italics="true" pageId="11" pageNumber="40">Hemicycla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C744D28FA13C26C88C68EB20556FB71" box="[1191,1330,1165,1189]" class="Gastropoda" family="Helicidae" genus="Adiverticula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA13C26C88C68EB20556FB71" box="[1191,1330,1165,1189]" italics="true" pageId="11" pageNumber="40">Adiverticula</emphasis>
|
||
</taxonomicName>
|
||
)), is supported by a moderately high posterior probability value of 92. Because of these results and the presence of a diverticulum on the stalk of the bursa copulatrix,
|
||
<taxonomicName id="4C744D28FA13C26C8EF98EEB0332FB3F" box="[664,854,1235,1259]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="berkeleii" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8EF98EEB02C8FB3F" box="[664,684,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8EA78EEB02BEFB3F" box="[710,730,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8E958EEC0332FB3F" box="[756,854,1235,1259]" italics="true" pageId="11" pageNumber="40">berkeleii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D28FA13C26C8F078EEB048DFB3F" box="[870,1257,1235,1259]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA13C26C8F078EEB031EFB3F" box="[870,890,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8FF58EEB03CCFB3F" box="[916,936,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FA08EEC048DFB3F" box="[961,1257,1235,1259]" italics="true" pageId="11" pageNumber="40">quadricincta quadricincta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B900EAB9FA13C26C89458EEB055CFB3F" box="[1316,1336,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C89308EEB0501FB3F" box="[1361,1381,1236,1259]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.) cf.
|
||
<emphasis id="B900EAB9FA13C26C8CF68EC80179FADA" box="[151,285,1271,1294]" italics="true" pageId="11" pageNumber="40">paivanopsis</emphasis>
|
||
(which were not assigned to any subgenus by Alonso & Ibáñez 2007), are placed in
|
||
<emphasis id="B900EAB9FA13C26C88808EC905F8FADA" box="[1249,1436,1270,1294]" italics="true" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C744D28FA13C26C88808EC9053DFADA" box="[1249,1369,1270,1294]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus" sensu="stricto">Hemicycla</taxonomicName>
|
||
<taxonomicNameLabel id="A23357C2FA13C26C89008EC805F8FADA" box="[1377,1436,1271,1294]" pageId="11" pageNumber="40" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Pairwise p-genetic distances between the
|
||
<emphasis id="B900EAB9FA13C26C8E0F8F26034EFAE5" box="[622,810,1305,1329]" italics="true" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C744D28FA13C26C8E0F8F260282FAE5" box="[622,742,1305,1329]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus" sensu="stricto">Hemicycla</taxonomicName>
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8E8F8F25034EFAE5" box="[750,810,1306,1329]" pageId="11" pageNumber="40" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the
|
||
<emphasis id="B900EAB9FA13C26C8FEC8F2503C5FAE5" box="[909,929,1306,1329]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<taxonomicName id="4C744D28FA13C26C8FDB8F260421FAE5" box="[954,1093,1305,1329]" class="Gastropoda" family="Helicidae" genus="Adiverticula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA13C26C8FDB8F260421FAE5" box="[954,1093,1305,1329]" italics="true" pageId="11" pageNumber="40">Adiverticula</emphasis>
|
||
</taxonomicName>
|
||
) lineages range from 0.101 to 0.141. The
|
||
<emphasis id="B900EAB9FA13C26C8D7B8F0301B3FA80" box="[282,471,1340,1364]" italics="true" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C744D28FA13C26C8D7B8F0301F6FA80" box="[282,402,1340,1364]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="genus" sensu="stricto">Hemicycla</taxonomicName>
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8DFB8F0201B3FA80" box="[410,471,1341,1364]" pageId="11" pageNumber="40" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lineage contains four clades grouped in a polytomy. The first clade contains
|
||
<taxonomicName id="4C744D28FA13C26C892B8F02011EFAA2" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="bidentalis" subGenus="H." subSpecies="bidentalis">
|
||
<emphasis id="B900EAB9FA13C26C892B8F02053AFA80" box="[1354,1374,1341,1364]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C89168F0205EFFA80" box="[1399,1419,1341,1364]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8CF68F61011EFAA2" box="[151,378,1374,1398]" italics="true" pageId="11" pageNumber="40">bidentalis bidentalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D28FA13C26C8DEB8F5F024BFAA2" box="[394,559,1374,1399]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="inutilis" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8DEB8F5F01FAFAA3" box="[394,414,1376,1399]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8DD48F5F01ADFAA3" box="[437,457,1376,1399]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8D808F61024BFAA2" box="[481,559,1374,1398]" italics="true" pageId="11" pageNumber="40">inutilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C8E078F5F035EFAA2" box="[614,826,1374,1399]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="pouchadan" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8E078F5F021EFAA3" box="[614,634,1376,1399]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8EF38F5F02C2FAA3" box="[658,678,1376,1399]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8EDF8F61035EFAA2" box="[702,826,1374,1398]" italics="true" pageId="11" pageNumber="40">pouchadan</emphasis>
|
||
</taxonomicName>
|
||
from Tenerife; it is supported by a posterior probability value of 82. The second clade includes
|
||
<emphasis id="B900EAB9FA13C26C8E1C8FBD02F5FA4D" box="[637,657,1410,1433]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8EC88FBD02D9FA4D" box="[681,701,1410,1433]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.) cf.
|
||
<emphasis id="B900EAB9FA13C26C8E9A8FBD03E5FA4D" box="[763,897,1410,1433]" italics="true" pageId="11" pageNumber="40">paivanopsis</emphasis>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C8FD88FBD055CFA4D" box="[953,1336,1409,1433]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA13C26C8FD88FBD03A9FA4D" box="[953,973,1410,1433]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F848FBD039DFA4D" box="[997,1017,1410,1433]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88738FBE055CFA4D" box="[1042,1336,1409,1433]" italics="true" pageId="11" pageNumber="40">quadricincta quadricincta</emphasis>
|
||
</taxonomicName>
|
||
from La Gomera, with a support value of 88. The third clade comprises
|
||
<taxonomicName id="4C744D28FA13C26C8F3E8F99044AFA69" box="[863,1070,1444,1469]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="saponacea" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8F3E8F990317FA69" box="[863,883,1446,1469]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8FEA8F9903FBFA69" box="[907,927,1446,1469]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FD78F99044AFA69" box="[950,1070,1446,1469]" italics="true" pageId="11" pageNumber="40">saponacea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C88048F99057BFA69" box="[1125,1311,1444,1469]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="berkeleii" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C88048F99041DFA69" box="[1125,1145,1446,1469]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88F08F9904C1FA69" box="[1169,1189,1446,1469]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88DC8F9A057BFA69" box="[1213,1311,1445,1469]" italics="true" pageId="11" pageNumber="40">berkeleii</emphasis>
|
||
</taxonomicName>
|
||
from Gran Canaria, with a support value of 100. Specimens assigned to
|
||
<taxonomicName id="4C744D28FA13C26C8F2F8FF7047BFA0B" box="[846,1055,1479,1503]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="saponacea" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8F2F8FF70306FA0B" box="[846,866,1480,1503]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F1A8FF703EBFA0B" box="[891,911,1480,1503]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FC68FF7047BFA0B" box="[935,1055,1480,1503]" italics="true" pageId="11" pageNumber="40">saponacea</emphasis>
|
||
</taxonomicName>
|
||
on the basis of shell morphology were not resolved as a monophyletic group. The fourth clade contains only
|
||
<taxonomicName id="4C744D28FA13C26C8F858FD404F0F9D6" box="[996,1172,1514,1538]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="plicaria" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8F858FD4039CF9D6" box="[996,1016,1515,1538]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C886E8FD40447F9D6" box="[1039,1059,1515,1538]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C885A8FD504F0F9D6" box="[1083,1172,1514,1538]" italics="true" pageId="11" pageNumber="40">plicaria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA13C26C8CA48C3201ADF8CD" blockId="11.[151,1437,151,2027]" pageId="11" pageNumber="40">
|
||
The second lineage is composed of two clades, the first comprising all specimens of
|
||
<taxonomicName id="4C744D28FA13C26C88158C310523F9F1" box="[1140,1351,1549,1573]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C88158C3104ECF9F1" box="[1140,1160,1550,1573]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88C18C3204D5F9F1" box="[1184,1201,1549,1573]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88A98C310523F9F1" box="[1224,1351,1550,1573]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
(posterior probability value of 100), and the seconded all specimens of
|
||
<taxonomicName id="4C744D28FA13C26C8F088C0E0466F99C" box="[873,1026,1584,1608]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F088C0E0319F99C" box="[873,893,1585,1608]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8FF58C0F03C1F99C" box="[916,933,1584,1608]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FDD8C0F0466F99C" box="[956,1026,1584,1608]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C88698C0F0422F99C" bold="true" box="[1032,1094,1584,1608]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C88698C0F0422F99C" box="[1032,1094,1584,1608]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(posterior probability value of 97). Mean p-genetic distances between species included in this study range from 0.018 between
|
||
<taxonomicName id="4C744D28FA13C26C88A88C6B0509F9BF" box="[1225,1389,1619,1643]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="inutilis" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C88A88C6B04B9F9BF" box="[1225,1245,1620,1643]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88958C6B056CF9BF" box="[1268,1288,1620,1643]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C897E8C6C0509F9BF" box="[1311,1389,1619,1643]" italics="true" pageId="11" pageNumber="40">inutilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C8CF68C48010BF95A" box="[151,367,1654,1678]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="pouchadan" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8CF68C4800CFF95A" box="[151,171,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8CA48C4800BDF95A" box="[197,217,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8C928C49010BF95A" box="[243,367,1654,1678]" italics="true" pageId="11" pageNumber="40">pouchadan</emphasis>
|
||
</taxonomicName>
|
||
and 0.145 between
|
||
<taxonomicName id="4C744D28FA13C26C8E3B8C48036AF95A" box="[602,782,1654,1678]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="plicaria" subGenus="H.">
|
||
<emphasis id="B900EAB9FA13C26C8E3B8C48020AF95A" box="[602,622,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8EE98C4802F8F95A" box="[648,668,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8ED78C49036AF95A" box="[694,782,1654,1678]" italics="true" pageId="11" pageNumber="40">plicaria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C8F2B8C4804AAF95A" box="[842,1230,1654,1678]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA13C26C8F2B8C48033AF95A" box="[842,862,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F198C4803E8F95A" box="[888,908,1655,1678]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FC78C4904AAF95A" box="[934,1230,1654,1678]" italics="true" pageId="11" pageNumber="40">quadricincta quadricincta</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<tableCitation id="C6F60310FA13C26C896F8C49053CF95A" box="[1294,1368,1654,1678]" captionStart="TABLE 6" captionStartId="13.[151,235,472,493]" captionText="TABLE 6. Mean genetic p-distances between species within the genus Hemicycla based on an alignment of 661 bp fragments of the COI gene. Calculations were conducted in MEGA 4 4.0. 2 (Tamura et al., 2007). Taxon 1 2 3 4 5 6 7 8 9 10 H. (H.) plicaria (1) -" pageId="11" pageNumber="40">Tab. 6</tableCitation>
|
||
). The mean p-genetic distance between
|
||
<taxonomicName id="4C744D28FA13C26C8E728CA5028CF965" box="[531,744,1689,1713]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8E728CA50243F965" box="[531,551,1690,1713]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8E5E8CA60234F965" box="[575,592,1689,1713]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8E098CA5028CF965" box="[616,744,1690,1713]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D28FA13C26C8F7E8CA503DEF965" box="[799,954,1689,1713]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F7E8CA50357F965" box="[799,819,1690,1713]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8F2A8CA60338F965" box="[843,860,1689,1713]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8F158CA603DEF965" box="[884,954,1689,1713]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8FA38CA60465F965" bold="true" box="[962,1025,1689,1713]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8FA38CA60465F965" box="[962,1025,1689,1713]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is 0.045, whereas within this group, mean p-genetic distances for
|
||
<taxonomicName id="4C744D28FA13C26C8D868C8202A4F900" box="[487,704,1724,1748]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8D868C82019FF900" box="[487,507,1725,1748]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8E748C830242F900" box="[533,550,1724,1748]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8E218C8202A4F900" box="[576,704,1725,1748]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
are 0.003, and 0.004 for
|
||
<taxonomicName id="4C744D28FA13C26C8F878C8204E1F900" box="[998,1157,1724,1748]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F878C82039EF900" box="[998,1018,1725,1748]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88758C830441F900" box="[1044,1061,1724,1748]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C885E8C8304E1F900" box="[1087,1157,1724,1748]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C88EF8C8304AAF900" bold="true" box="[1166,1230,1724,1748]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C88EF8C8304AAF900" box="[1166,1230,1724,1748]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The gap between mean p-distances within groups and mean distances among groups for these two taxa strongly corroborates the validity of the new species.
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA13C26C8CA68D1B02E3F83F" blockId="11.[151,1437,151,2027]" pageId="11" pageNumber="40">
|
||
<emphasis id="B900EAB9FA13C26C8CA68D1B018DF8E9" bold="true" box="[199,489,1828,1853]" pageId="11" pageNumber="40">Morphometric analysis.</emphasis>
|
||
The preliminary comparison of some of the shell measurements in
|
||
<figureCitation id="134F2A2EFA13C26C89768D1B053AF8E8" box="[1303,1374,1828,1852]" captionStart="FIGURE 7" captionStartId="14.[151,250,726,749]" captionTargetBox="[187,1400,209,701]" captionTargetId="figure@14.[183,1403,195,705]" captionTargetPageId="14" captionText="FIGURE 7. Scatter plots of some linear shell measurements (see Fig. 1) that are frequently used to compare gastropod species. Black dots: Hemicycla (Adiverticula) diegoi n. sp.; gray dots: H. (A.) mascaensis." httpUri="https://zenodo.org/record/276756/files/figure.png" pageId="11" pageNumber="40">Fig. 7</figureCitation>
|
||
A–F already shows the value of the maximum shell diameter (Dmax), shell height (SH), spire height (SpH), body whorl height (BWH) and maximum aperture width (AWmax) as discriminant characters. Descriptive statistics and results of Welch’s t-test are shown in
|
||
<tableCitation id="C6F60310FA13C26C8D928DB2025EF871" box="[499,570,1933,1957]" captionStart="TABLE 3" captionStartId="7.[151,235,404,425]" captionTargetBox="[151,1437,514,1405]" captionTargetPageId="7" captionText="TABLE 3. Shell measurements (in mm) and univariate statistics for Hemicycla (Adiverticula) mascaensis and H. (A.) diegoi n." httpUri="http://table.plazi.org/id/DF0B6623FA1FC2608CF68BAB05F8FE7D" pageId="11" pageNumber="40" tableUuid="DF0B6623FA1FC2608CF68BAB05F8FE7D">Tab. 3</tableCitation>
|
||
for all linear shell measurements. Welch’s t-test results in significant differences in means (p <0.001) between shells belonging either to
|
||
<taxonomicName id="4C744D28FA13C26C8F378D8E0395F81C" box="[854,1009,1968,1992]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C8F378D8E030EF81C" box="[854,874,1969,1992]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C8FE28D8F03F0F81C" box="[899,916,1968,1992]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C8FCA8D8F0395F81C" box="[939,1009,1968,1992]" italics="true" pageId="11" pageNumber="40">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA13C26C8F988D8E045CF81D" bold="true" box="[1017,1080,1969,1993]" pageId="11" pageNumber="40">
|
||
<taxonomicNameLabel id="A23357C2FA13C26C8F988D8E045CF81D" box="[1017,1080,1969,1993]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
or to
|
||
<taxonomicName id="4C744D28FA13C26C881D8D8E0535F81C" box="[1148,1361,1968,1992]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="40" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA13C26C881D8D8E04F4F81C" box="[1148,1168,1969,1992]" italics="true" pageId="11" pageNumber="40">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA13C26C88C98D8F04DDF81C" box="[1192,1209,1968,1992]" italics="true" pageId="11" pageNumber="40">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA13C26C88B08D8E0535F81C" box="[1233,1361,1969,1992]" italics="true" pageId="11" pageNumber="40">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
for all measurements, except aperture height (AH).
|
||
</paragraph>
|
||
<caption id="DF0B6623FA14C26B8CF68CB20555F92B" httpUri="https://zenodo.org/record/276754/files/figure.png" pageId="12" pageNumber="41" targetBox="[164,1411,176,1651]" targetPageId="12">
|
||
<paragraph id="8BCB36ABFA14C26B8CF68CB20555F92B" blockId="12.[151,1436,1677,1792]" pageId="12" pageNumber="41">
|
||
<emphasis id="B900EAB9FA14C26B8CF68CB20176F977" bold="true" box="[151,274,1677,1700]" pageId="12" pageNumber="41">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C744D28FA14C26B8D798CB202F6F977" box="[280,658,1677,1699]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="41" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA14C26B8D798CB201E0F977" box="[280,388,1677,1699]" italics="true" pageId="12" pageNumber="41">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA14C26B8DF28CB20274F977" box="[403,528,1677,1699]" italics="true" pageId="12" pageNumber="41">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA14C26B8E7F8CB102F6F977" box="[542,658,1678,1699]" italics="true" pageId="12" pageNumber="41">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B900EAB9FA14C26B8EFC8CB202D2F976" bold="true" box="[669,694,1677,1698]" pageId="12" pageNumber="41">A.</emphasis>
|
||
Living animal from the type locality (scale bar = 10 mm).
|
||
<emphasis id="B900EAB9FA14C26B89648CB10578F977" bold="true" box="[1285,1308,1678,1699]" pageId="12" pageNumber="41">B.</emphasis>
|
||
Genital system (scale bar = 5 mm).
|
||
<emphasis id="B900EAB9FA14C26B8DF98C9401D4F916" bold="true" box="[408,432,1707,1730]" pageId="12" pageNumber="41">C.</emphasis>
|
||
Internal anatomy of the penis (scale bar = 1 mm) with proximal penial papilla (pp), distal penial papilla (dp), contact organ (co) and distal part of epiphallus (ep).
|
||
<emphasis id="B900EAB9FA14C26B8F538CF4032FF934" bold="true" box="[818,843,1739,1760]" pageId="12" pageNumber="41">D.</emphasis>
|
||
Dart (scale bar = 1 mm).
|
||
<emphasis id="B900EAB9FA14C26B88358CF4040FF934" bold="true" box="[1108,1131,1739,1760]" pageId="12" pageNumber="41">E.</emphasis>
|
||
Mandible (scale bar = 1mm).
|
||
<emphasis id="B900EAB9FA14C26B8CF68CD500AAF92B" bold="true" box="[151,206,1770,1791]" pageId="12" pageNumber="41">F–H.</emphasis>
|
||
Radula teeth (scale bar = 10 μm): marginal teeth (
|
||
<emphasis id="B900EAB9FA14C26B8EAD8CD502BFF92B" bold="true" box="[716,731,1770,1791]" pageId="12" pageNumber="41">F</emphasis>
|
||
), outer lateral teeth (
|
||
<emphasis id="B900EAB9FA14C26B8FD18CD603A0F8D4" bold="true" box="[944,964,1769,1792]" pageId="12" pageNumber="41">G</emphasis>
|
||
), central and inner lateral teeth (
|
||
<emphasis id="B900EAB9FA14C26B896E8CD50547F92B" bold="true" box="[1295,1315,1770,1791]" pageId="12" pageNumber="41">H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA14C26B8CA48D18024BF827" blockId="12.[151,1436,1831,2035]" pageId="12" pageNumber="41">
|
||
Results from the discriminant function analysis for the log-transformed data are shown in
|
||
<tableCitation id="C6F60310FA14C26B88D98D180564F8EB" box="[1208,1280,1831,1855]" captionStart="TABLE 5" captionStartId="13.[151,235,152,173]" captionTargetBox="[151,1436,304,408]" captionTargetPageId="13" captionText="TABLE 5. Non-standardised and standardised structure coefficients of the discriminant function for all log-transformed linear shell measurements of Hemicycla (Adiverticula) diegoi n. sp. and Hemicycla (Adiverticula) mascaensis as defined in Fig. 1. Highest loadings in the standardised case are indicated in bold. For the meaning of the abbreviations see. Fig. 1." httpUri="http://table.plazi.org/id/DF0B6623FA15C26A8CF68AA70560FF3E" pageId="12" pageNumber="41" tableUuid="DF0B6623FA15C26A8CF68AA70560FF3E">Tab. 5</tableCitation>
|
||
. Standardised structure coefficients of the discriminant function show that shell height (SH), maximum shell diameter (Dmax), spire height (SpH), minimum aperture width (ABmin) and callus width (CW) have the strongest influence on the discriminant variable.
|
||
<taxonomicName id="4C744D28FA14C26B8DF38DAC027DF87F" box="[402,537,1939,1963]" class="Gastropoda" family="Helicidae" genus="Adiverticula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="41" phylum="Mollusca" rank="species" species="posteriori">
|
||
<emphasis id="B900EAB9FA14C26B8DF38DAC027DF87F" box="[402,537,1939,1963]" italics="true" pageId="12" pageNumber="41">A posteriori</emphasis>
|
||
</taxonomicName>
|
||
indentification of specimens results in 94.6% correct placements, and Wilks’-Λ statistics strongly supports the hypothesis, that
|
||
<taxonomicName id="4C744D28FA14C26B8ECC8D87032EF81B" box="[685,842,1975,1999]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="41" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA14C26B8ECC8D8702A5F81B" box="[685,705,1976,1999]" italics="true" pageId="12" pageNumber="41">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA14C26B8EBB8D88028FF81B" box="[730,747,1975,1999]" italics="true" pageId="12" pageNumber="41">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA14C26B8F658D88032EF81B" box="[772,842,1975,1999]" italics="true" pageId="12" pageNumber="41">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA14C26B8F338D8803F6F81B" bold="true" box="[850,914,1975,1999]" pageId="12" pageNumber="41">
|
||
<taxonomicNameLabel id="A23357C2FA14C26B8F338D8803F6F81B" box="[850,914,1975,1999]" pageId="12" pageNumber="41" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C744D28FA14C26B8FAA8D8704C4F81B" box="[971,1184,1975,1999]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="41" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA14C26B8FAA8D8703BBF81B" box="[971,991,1976,1999]" italics="true" pageId="12" pageNumber="41">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA14C26B8F968D88046CF81B" box="[1015,1032,1975,1999]" italics="true" pageId="12" pageNumber="41">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA14C26B88408D8704C4F81B" box="[1057,1184,1976,1999]" italics="true" pageId="12" pageNumber="41">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
represent two distinct taxa (Λ = 0.078, df = 10, p <0.001).
|
||
</paragraph>
|
||
<caption id="DF0B6623FA15C26A8CF68AA70560FF3E" ID-Table-UUID="DF0B6623FA15C26A8CF68AA70560FF3E" httpUri="http://table.plazi.org/id/DF0B6623FA15C26A8CF68AA70560FF3E" pageId="13" pageNumber="42" targetBox="[151,1436,304,408]" targetIsTable="true" targetPageId="13">
|
||
<paragraph id="8BCB36ABFA15C26A8CF68AA70560FF3E" blockId="13.[151,1436,152,284]" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8CF68AA70160FF79" bold="true" box="[151,260,152,173]" pageId="13" pageNumber="42">TABLE 5.</emphasis>
|
||
Non-standardised and standardised structure coefficients of the discriminant function for all log-transformed linear shell measurements of
|
||
<taxonomicName id="4C744D28FA15C26A8DE48A8802ABFF19" box="[389,719,183,205]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA15C26A8DE48A880195FF19" box="[389,497,183,205]" italics="true" pageId="13" pageNumber="42">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA15C26A8E638A88021BFF19" box="[514,639,183,205]" italics="true" pageId="13" pageNumber="42">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA15C26A8EEE8A8802ABFF19" box="[655,719,183,205]" italics="true" pageId="13" pageNumber="42">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA15C26A8EB78A880375FF18" bold="true" box="[726,785,183,204]" pageId="13" pageNumber="42">
|
||
<taxonomicNameLabel id="A23357C2FA15C26A8EB78A880375FF18" box="[726,785,183,204]" pageId="13" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C744D28FA15C26A8F278A8804A7FF19" box="[838,1219,183,205]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA15C26A8F278A8803D6FF19" box="[838,946,183,205]" italics="true" pageId="13" pageNumber="42">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B900EAB9FA15C26A8FA28A880424FF19" box="[963,1088,183,205]" italics="true" pageId="13" pageNumber="42">Adiverticula</emphasis>
|
||
)
|
||
<emphasis id="B900EAB9FA15C26A88318A8704A7FF19" box="[1104,1219,184,205]" italics="true" pageId="13" pageNumber="42">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
as defined in Fig. 1. Highest loadings in the standardised case are indicated in bold. For the meaning of the abbreviations see. Fig. 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA15C26A8DC68B39025BFEC8" blockId="13.[151,1436,152,284]" box="[423,575,262,284]" pageId="13" pageNumber="42">Shell character</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE8B0F0516FE4C" pageId="13" pageNumber="42">
|
||
<table id="F974C40BFA153D988CF68B0F05F8FE4C" box="[151,1436,304,408]" gridcols="9" gridrows="3" pageId="13" pageNumber="42">
|
||
<tr id="354434E9FA153D988CF68B0F05F8FE92" box="[151,1436,304,326]" gridrow="0" pageId="13" pageNumber="42">
|
||
<th id="76955D95FA153D988CF68B0F01E5FE92" box="[151,385,304,326]" gridcol="0" gridrow="0" pageId="13" pageNumber="42">Canonical coefficients</th>
|
||
<th id="76955D95FA153D988DC68B0F0181FE92" box="[423,485,304,326]" gridcol="1" gridrow="0" pageId="13" pageNumber="42">Dmax</th>
|
||
<th id="76955D95FA153D988E6C8B0F022CFE92" box="[525,584,304,326]" gridcol="2" gridrow="0" pageId="13" pageNumber="42">Dmin</th>
|
||
<th id="76955D95FA153D988E158B0F02D1FE92" box="[628,693,304,326]" gridcol="3" gridrow="0" pageId="13" pageNumber="42">SH</th>
|
||
<th id="76955D95FA153D988EB88B0F0362FE92" box="[729,774,304,326]" gridcol="4" gridrow="0" pageId="13" pageNumber="42">SpH</th>
|
||
<th id="76955D95FA153D988F5E8B0F031EFE92" box="[831,890,304,326]" gridcol="5" gridrow="0" pageId="13" pageNumber="42">BWH</th>
|
||
<th id="76955D95FA153D988FC48B0F03BEFE92" box="[933,986,304,326]" gridcol="6" gridrow="0" pageId="13" pageNumber="42">AW</th>
|
||
<th id="76955D95FA153D98886D8B0F045DFE92" box="[1036,1081,304,326]" gridcol="7" gridrow="0" pageId="13" pageNumber="42">AH</th>
|
||
<th id="76955D95FA153D9888108B0F05F8FE92" box="[1137,1436,304,326]" gridcol="8" gridrow="0" pageId="13" pageNumber="42">ABmin ABmax CW</th>
|
||
</tr>
|
||
<tr id="354434E9FA153D988CF68B6605F8FEBB" box="[151,1436,345,367]" gridrow="1" pageId="13" pageNumber="42">
|
||
<th id="76955D95FA153D988CF68B6601E5FEBB" box="[151,385,345,367]" gridcol="0" gridrow="1" pageId="13" pageNumber="42">non-standardised</th>
|
||
<td id="76955D95FA153D988DC68B660181FEBB" box="[423,485,345,367]" gridcol="1" gridrow="1" pageId="13" pageNumber="42">18.71</td>
|
||
<td id="76955D95FA153D988E6C8B66022CFEBB" box="[525,584,345,367]" gridcol="2" gridrow="1" pageId="13" pageNumber="42">0.90</td>
|
||
<td id="76955D95FA153D988E158B6602D1FEBB" box="[628,693,345,367]" gridcol="3" gridrow="1" pageId="13" pageNumber="42">-19.83</td>
|
||
<td id="76955D95FA153D988EB88B660362FEBB" box="[729,774,345,367]" gridcol="4" gridrow="1" pageId="13" pageNumber="42">2.72</td>
|
||
<td id="76955D95FA153D988F5E8B66031EFEBB" box="[831,890,345,367]" gridcol="5" gridrow="1" pageId="13" pageNumber="42">-0.12</td>
|
||
<td id="76955D95FA153D988FC48B6603BEFEBB" box="[933,986,345,367]" gridcol="6" gridrow="1" pageId="13" pageNumber="42">-4.48</td>
|
||
<td id="76955D95FA153D98886D8B66045DFEBB" box="[1036,1081,345,367]" gridcol="7" gridrow="1" pageId="13" pageNumber="42">1.73</td>
|
||
<td id="76955D95FA153D9888108B6605F8FEBB" box="[1137,1436,345,367]" gridcol="8" gridrow="1" pageId="13" pageNumber="42">-6.02 -2.37 -7.71</td>
|
||
</tr>
|
||
<tr id="354434E9FA153D988CF68BBD05F8FE4C" box="[151,1436,386,408]" gridrow="2" pageId="13" pageNumber="42">
|
||
<th id="76955D95FA153D988CF68BBD01E5FE4C" box="[151,385,386,408]" gridcol="0" gridrow="2" pageId="13" pageNumber="42">standardised</th>
|
||
<td id="76955D95FA153D988DC68BBD0181FE4C" box="[423,485,386,408]" gridcol="1" gridrow="2" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8DC68BBC01B7FE4C" bold="true" box="[423,467,387,408]" pageId="13" pageNumber="42">0.87</emphasis>
|
||
</td>
|
||
<td id="76955D95FA153D988E6C8BBD022CFE4C" box="[525,584,386,408]" gridcol="2" gridrow="2" pageId="13" pageNumber="42">0.04</td>
|
||
<td id="76955D95FA153D988E158BBD02D1FE4C" box="[628,693,386,408]" gridcol="3" gridrow="2" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8E158BBC02CCFE4C" bold="true" box="[628,680,387,408]" pageId="13" pageNumber="42">-1.23</emphasis>
|
||
</td>
|
||
<td id="76955D95FA153D988EB88BBD0362FE4C" box="[729,774,386,408]" gridcol="4" gridrow="2" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8EB88BBC0361FE4C" bold="true" box="[729,773,387,408]" pageId="13" pageNumber="42">0.61</emphasis>
|
||
</td>
|
||
<td id="76955D95FA153D988F5E8BBD031EFE4C" box="[831,890,386,408]" gridcol="5" gridrow="2" pageId="13" pageNumber="42">-0.01</td>
|
||
<td id="76955D95FA153D988FC48BBD03BEFE4C" box="[933,986,386,408]" gridcol="6" gridrow="2" pageId="13" pageNumber="42">-0.30</td>
|
||
<td id="76955D95FA153D98886D8BBD045DFE4C" box="[1036,1081,386,408]" gridcol="7" gridrow="2" pageId="13" pageNumber="42">0.22</td>
|
||
<td id="76955D95FA153D9888108BBD05F8FE4C" box="[1137,1436,386,408]" gridcol="8" gridrow="2" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A88108BBC04C2FE4C" bold="true" box="[1137,1190,387,408]" pageId="13" pageNumber="42">-0.55</emphasis>
|
||
-0.16 -0.47
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF0B6623FA15C26A8CF68BE70246FDB3" pageId="13" pageNumber="42">
|
||
<paragraph id="8BCB36ABFA15C26A8CF68BE70246FDB3" blockId="13.[151,1436,472,987]" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8CF68BE70160FE39" bold="true" box="[151,260,472,493]" pageId="13" pageNumber="42">TABLE 6.</emphasis>
|
||
Mean genetic p-distances between species within the genus
|
||
<taxonomicName id="4C744D28FA15C26A8F0E8BE703BFFE3A" box="[879,987,472,494]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAB9FA15C26A8F0E8BE703BFFE3A" box="[879,987,472,494]" italics="true" pageId="13" pageNumber="42">Hemicycla</emphasis>
|
||
</taxonomicName>
|
||
based on an alignment of 661 bp fragments of the COI gene. Calculations were conducted in MEGA 4 4.0.2 (Tamura
|
||
<emphasis id="B900EAB9FA15C26A8FE28BC703D4FDD9" box="[899,944,503,525]" italics="true" pageId="13" pageNumber="42">et al</emphasis>
|
||
., 2007). Taxon 1 2 3 4 5 6 7 8 9 10
|
||
<taxonomicName id="4C744D28FA15C26A8CFE886D0159FDB3" box="[159,317,593,615]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="plicaria" subGenus="H.">
|
||
<emphasis id="B900EAB9FA15C26A8CFE886D00D5FDB3" box="[159,177,594,615]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7886D00BCFDB3" box="[198,216,594,615]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C886E0159FDB3" box="[237,317,593,615]" italics="true" pageId="13" pageNumber="42">plicaria</emphasis>
|
||
</taxonomicName>
|
||
(1) -
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE88440219FD44" blockId="13.[151,1436,472,987]" box="[159,637,634,657]" pageId="13" pageNumber="42">
|
||
<taxonomicName id="4C744D28FA15C26A8CFE8844010FFD44" box="[159,363,634,656]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="bidentalis" subGenus="H." subSpecies="bidentalis">
|
||
<emphasis id="B900EAB9FA15C26A8CFE884400D5FD44" box="[159,177,635,656]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7884400BCFD44" box="[198,216,635,656]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C8845009EFD44" box="[237,250,634,656]" italics="true" pageId="13" pageNumber="42">b</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA15C26A8D668845010FFD44" box="[263,363,634,656]" italics="true" pageId="13" pageNumber="42">bidentalis</emphasis>
|
||
</taxonomicName>
|
||
(2) 0.139 -
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE889A02B3FD6E" blockId="13.[151,1436,472,987]" box="[159,727,676,698]" pageId="13" pageNumber="42">
|
||
<taxonomicName id="4C744D28FA15C26A8CFE889A013AFD6E" box="[159,350,676,698]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="pouchadan" subGenus="H.">
|
||
<emphasis id="B900EAB9FA15C26A8CFE889A00D5FD6E" box="[159,177,677,698]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7889A00BCFD6E" box="[198,216,677,698]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C889B013AFD6E" box="[237,350,676,698]" italics="true" pageId="13" pageNumber="42">pouchadan</emphasis>
|
||
</taxonomicName>
|
||
(3) 0.138 0.083 -
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE88F10356FD37" blockId="13.[151,1436,472,987]" box="[159,818,717,739]" pageId="13" pageNumber="42">
|
||
<taxonomicName id="4C744D28FA15C26A8CFE88F10157FD37" box="[159,307,717,739]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="inutilis" subGenus="H.">
|
||
<emphasis id="B900EAB9FA15C26A8CFE88F100D5FD37" box="[159,177,718,739]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA788F100BCFD37" box="[198,216,718,739]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C88F20157FD37" box="[237,307,717,739]" italics="true" pageId="13" pageNumber="42">inutilis</emphasis>
|
||
</taxonomicName>
|
||
(4) 0.132 0.080 0.018 -
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE88C703E8FCD8" blockId="13.[151,1436,472,987]" box="[159,908,758,781]" pageId="13" pageNumber="42">
|
||
<taxonomicName id="4C744D28FA15C26A8CFE88C701F1FCD9" box="[159,405,759,781]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="quadricincta" subGenus="H." subSpecies="quadricincta">
|
||
<emphasis id="B900EAB9FA15C26A8CFE88C700D5FCD9" box="[159,177,760,781]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA788C700BCFCD9" box="[198,216,760,781]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C88C70162FCD9" box="[237,262,760,781]" italics="true" pageId="13" pageNumber="42">qu</emphasis>
|
||
.
|
||
<emphasis id="B900EAB9FA15C26A8D7588C801F1FCD9" box="[276,405,759,781]" italics="true" pageId="13" pageNumber="42">quadricincta</emphasis>
|
||
</taxonomicName>
|
||
(5) 0.145 0.118 0.124 0.120 -
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE891E0382FCE2" blockId="13.[151,1436,472,987]" box="[159,998,800,822]" pageId="13" pageNumber="42">
|
||
<emphasis id="B900EAB9FA15C26A8CFE891E00D5FCE2" box="[159,177,801,822]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7891E00BCFCE2" box="[198,216,801,822]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.) cf.
|
||
<emphasis id="B900EAB9FA15C26A8D6F891E01E3FCE2" box="[270,391,801,822]" italics="true" pageId="13" pageNumber="42">paivanopsis</emphasis>
|
||
(6) 0.139 0.113 0.128 0.126 0.092 -
|
||
</paragraph>
|
||
<paragraph id="8BCB36ABFA15C26A8CFE89750535FC0F" blockId="13.[151,1436,472,987]" pageId="13" pageNumber="42">
|
||
<taxonomicName id="4C744D28FA15C26A8CFE89750122FC8B" box="[159,326,841,863]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="berkeleii" subGenus="H.">
|
||
<emphasis id="B900EAB9FA15C26A8CFE897500D5FC8B" box="[159,177,842,863]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7897500BCFC8B" box="[198,216,842,863]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C89760122FC8B" box="[237,326,841,863]" italics="true" pageId="13" pageNumber="42">berkeleii</emphasis>
|
||
</taxonomicName>
|
||
(7) 0.124 0.106 0.104 0.098 0.125 0.121 -
|
||
<taxonomicName id="4C744D28FA15C26A8CFE894C013DFC5C" box="[159,345,883,904]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="saponacea" subGenus="H.">
|
||
<emphasis id="B900EAB9FA15C26A8CFE894C00D5FC5C" box="[159,177,883,904]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA7894C00BCFC5C" box="[198,216,883,904]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8C894C013DFC5C" box="[237,345,883,904]" italics="true" pageId="13" pageNumber="42">saponacea</emphasis>
|
||
</taxonomicName>
|
||
(8) 0.132 0.117 0.118 0.116 0.126 0.126 0.051 -
|
||
<taxonomicName id="4C744D28FA15C26A8CFE89A2013BFC66" box="[159,351,924,946]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="mascaensis" subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA15C26A8CFE89A200D5FC66" box="[159,177,925,946]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA789A300B1FC65" box="[198,213,924,945]" italics="true" pageId="13" pageNumber="42">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8A89A2013BFC66" box="[235,351,925,946]" italics="true" pageId="13" pageNumber="42">mascaensis</emphasis>
|
||
</taxonomicName>
|
||
(9) 0.131 0.102 0.108 0.104 0.129 0.129 0.105 0.107 -
|
||
<taxonomicName id="4C744D28FA15C26A8CFE89F9014FFC0F" box="[159,299,965,987]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="42" phylum="Mollusca" rank="species" species="diegoi" status="sp. nov." subGenus="Adiverticula">
|
||
<emphasis id="B900EAB9FA15C26A8CFE89F900D5FC0F" box="[159,177,966,987]" italics="true" pageId="13" pageNumber="42">H</emphasis>
|
||
. (
|
||
<emphasis id="B900EAB9FA15C26A8CA789F900B1FC0F" box="[198,213,966,987]" italics="true" pageId="13" pageNumber="42">A</emphasis>
|
||
.)
|
||
<emphasis id="B900EAB9FA15C26A8C8A89FA014FFC0F" box="[235,299,965,987]" italics="true" pageId="13" pageNumber="42">diegoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAB9FA15C26A8D5189FA010DFC0E" bold="true" box="[304,361,965,986]" pageId="13" pageNumber="42">
|
||
<taxonomicNameLabel id="A23357C2FA15C26A8D5189FA010DFC0E" box="[304,361,965,986]" pageId="13" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(10) 0.135 0.106 0.111 0.109 0.132 0.132 0.112 0.116 0.045 -
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |