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<document id="E08C5F426F98687B4E54AF4A7948D6FE" ID-CLB-Dataset="25871" ID-DOI="10.11646/zootaxa.4647.1.17" ID-GBIF-Dataset="dac99f74-3514-47c5-a5fe-e33ef247dde0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353053" ID-ZooBank="6B225596-A393-40E9-A386-8758A5ABC5D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564249432578" checkinUser="plazi" docAuthor="Norton, Roy A. &amp; Ermilov, Sergey G." docDate="2019" docId="03DD87AFFFDC966B70DE2216FEEBFAE6" docLanguage="en" docName="zootaxa.4647.1.17.pdf" docOrigin="Zootaxa 4647 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anderemaeus dentatus Norton &amp; Ermilov 2019, n. sp." docType="treatment" docVersion="12" lastPageNumber="264" masterDocId="FFE4FFD7FFD3967C70492523FFB2FFA2" masterDocTitle="Anderemaeus (Acari, Oribatida) - overview, three new species from South America and reassessment of Anderemaeidae supported by ontogeny" masterLastPageNumber="289" masterPageNumber="241" pageNumber="256" updateTime="1698747558919" updateUser="ExternalLinkService">
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<mods:title id="F7C8B18A12512FDACCF4002D3F002132">Anderemaeus (Acari, Oribatida) - overview, three new species from South America and reassessment of Anderemaeidae supported by ontogeny</mods:title>
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<mods:namePart id="57769E4C13957BECD233F987634729F5">Norton, Roy A.</mods:namePart>
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<mods:namePart id="01C28682E20734A64541BCCEAE03D2A9">Ermilov, Sergey G.</mods:namePart>
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<treatment id="03DD87AFFFDC966B70DE2216FEEBFAE6" ID-DOI="http://doi.org/10.5281/zenodo.5629715" ID-GBIF-Taxon="159305164" ID-Zenodo-Dep="5629715" LSID="urn:lsid:plazi:treatment:03DD87AFFFDC966B70DE2216FEEBFAE6" httpUri="http://treatment.plazi.org/id/03DD87AFFFDC966B70DE2216FEEBFAE6" lastPageId="23" lastPageNumber="264" pageId="15" pageNumber="256">
<subSubSection id="C36E6532FFDC967370DE2216FE47F8ED" box="[151,501,1845,1872]" pageId="15" pageNumber="256" type="nomenclature">
<paragraph id="8BCB36B9FFDC967370DE2216FE47F8ED" blockId="15.[151,501,1845,1908]" box="[151,501,1845,1872]" pageId="15" pageNumber="256">
<heading id="D08381D5FFDC967370DE2216FE47F8ED" bold="true" box="[151,501,1845,1872]" fontSize="11" level="1" pageId="15" pageNumber="256" reason="1">
<emphasis id="B900EAABFFDC967370DE2216FE47F8ED" bold="true" box="[151,501,1845,1872]" pageId="15" pageNumber="256">
<taxonomicName id="4C744D3AFFDC967370DE2216FE1EF8F2" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[151,428,1845,1872]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="256" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFDC967370DE2216FE1EF8F2" bold="true" box="[151,428,1845,1872]" italics="true" pageId="15" pageNumber="256">Anderemaeus dentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357D0FFDC967371FA2216FE47F8ED" box="[435,501,1845,1871]" pageId="15" pageNumber="256" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36E6532FFDC966B70DE227AFEEBFAE6" lastPageId="23" lastPageNumber="264" pageId="15" pageNumber="256" type="description">
<paragraph id="8BCB36B9FFDC967370DE227AFE95F8D6" blockId="15.[151,501,1845,1908]" box="[151,295,1881,1908]" pageId="15" pageNumber="256">
(
<figureCitation id="134F2A3CFFDC967370D6227AFEACF8D6" box="[159,286,1881,1908]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStart-6="FIGURE 16" captionStartId-0="16.[151,250,1628,1653]" captionStartId-1="17.[151,250,1792,1817]" captionStartId-2="18.[151,250,1843,1868]" captionStartId-3="19.[151,250,1427,1452]" captionStartId-4="20.[151,250,1688,1713]" captionStartId-5="21.[151,250,1974,1999]" captionStartId-6="22.[151,250,1795,1820]" captionTargetBox-0="[151,1436,366,1551]" captionTargetBox-1="[152,1436,252,1752]" captionTargetBox-2="[164,1423,179,1816]" captionTargetBox-3="[151,1435,179,1399]" captionTargetBox-4="[160,1426,394,1630]" captionTargetBox-5="[155,1430,282,1933]" captionTargetBox-6="[151,1435,288,1767]" captionTargetId-0="figure@16.[151,1436,342,1600]" captionTargetId-1="figure@17.[151,1436,270,1764]" captionTargetId-2="figure@18.[164,1423,179,1816]" captionTargetId-3="figure@19.[151,1435,179,1399]" captionTargetId-4="figure@20.[151,1436,376,1661]" captionTargetId-5="figure@21.[151,1436,270,1946]" captionTargetId-6="figure@22.[151,1435,306,1767]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionText-0="FIGURE 10. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm." captionText-1="FIGURE 11. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, lateral view; B, posterior view. Scale bar 100 μm." captionText-2="FIGURE 12. Anderemaeus dentatus sp. nov., adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta h 1; F, notogastral seta p 1; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others)." captionText-3="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." captionText-4="FIGURE 14. Anderemaeus dentatus sp. nov., adult gnathosoma: A, subcapitulum, ventral view (palp omitted); B, left adoral lip; C, right palp, adaxial view; D, left chelicera, adaxial view. Scale bars 20 μm (A, C to same scale; D), 10 μm (B)." captionText-5="FIGURE 15. Anderemaeus dentatus sp. nov., adult legs (abaxial view): A, right leg I; B, trochanter, femur and genu of right leg II; C, trochanter, femur and genu of left leg III; D, left leg IV. Scale bar 50 μm." captionText-6="FIGURE 16. Anderemaeus dentatus sp. nov., nymphs: A, tritonymph, dorsal view (larger hysterosomal setae broken except lp, h 1); B, same, lateral view (inserts: close-up of seta le in upper right, bothridial region in lower left); C, as in A, close-up of bothridial seta; D, as in A, close-up of stalk bearing opening to opisthonotal gland; E, deutonymph, leg I, adaxial view; F, tritonymph, prodorsum, dorsal view. Photomicrographs in transmitted light, DIC. Scale bars:100 μm (A, B); 20 μm (C, E, F); 5 μm (D)." httpUri-0="https://zenodo.org/record/3353073/files/figure.png" httpUri-1="https://zenodo.org/record/3353075/files/figure.png" httpUri-2="https://zenodo.org/record/3353077/files/figure.png" httpUri-3="https://zenodo.org/record/3353079/files/figure.png" httpUri-4="https://zenodo.org/record/3353081/files/figure.png" httpUri-5="https://zenodo.org/record/3353083/files/figure.png" httpUri-6="https://zenodo.org/record/3353085/files/figure.png" pageId="15" pageNumber="256">Figs 1016</figureCitation>
)
</paragraph>
<paragraph id="8BCB36B9FFDC967370DE22B8FEBBF816" blockId="15.[151,1437,1947,2045]" box="[151,265,1947,1972]" pageId="15" pageNumber="256">
<emphasis id="B900EAABFFDC967370DE22B8FEBBF816" box="[151,265,1947,1972]" italics="true" pageId="15" pageNumber="256">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BCB36B9FFDC966C70DE229DFDDBFEE1" blockId="15.[151,1437,1947,2045]" lastBlockId="16.[151,1437,153,323]" lastPageId="16" lastPageNumber="257" pageId="15" pageNumber="256">
<taxonomicName id="4C744D3AFFDC967370DE229DFE9CF87A" authorityName="Hammer" authorityYear="1958" box="[151,302,1982,2008]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAABFFDC967370DE229DFE9CF87A" box="[151,302,1982,2008]" italics="true" pageId="15" pageNumber="256">Anderemaeus</emphasis>
</taxonomicName>
species with adults 614680. Rostrum with terminal mucro. Lamella with strong triangular tooth projecting anterolaterally, just proximal to small cusp; lamella continuing anteriorly as distinct prolamella; pair strongly convergent, with mutual distance of setae
<emphasis id="B900EAABFFC3966C723925BAFD36FF11" box="[624,644,153,179]" italics="true" pageId="16" pageNumber="257">le</emphasis>
distinctly less than that of pair
<emphasis id="B900EAABFFC3966C73AE25B9FC4DFF11" box="[999,1023,154,179]" italics="true" pageId="16" pageNumber="257">ro</emphasis>
. With distinct transverse prelamellar ridge. Seta
<emphasis id="B900EAABFFC3966C7150259DFE9DFF75" box="[281,303,190,215]" italics="true" pageId="16" pageNumber="257">in</emphasis>
long, inserted well anterior to notogastral margin, without supporting tubercle; with isolated subtriangular tubercle posterior to each seta. Bothridial seta long, baculiform. Notogastral setae erect, heavily barbed, relatively long. Femora III, IV with simple ventral keel (III) or large blade (IV); trochanters III and IV with large dorsodistal spine, smaller proximal tooth.
</paragraph>
<caption id="DF0B6631FFC3966C70DE237FFF5EF93B" httpUri="https://zenodo.org/record/3353073/files/figure.png" pageId="16" pageNumber="257" startId="16.[151,250,1628,1653]" targetBox="[151,1436,366,1551]" targetPageId="16">
<paragraph id="8BCB36B9FFC3966C70DE237FFF5EF93B" blockId="16.[151,1436,1628,1689]" pageId="16" pageNumber="257">
<emphasis id="B900EAABFFC3966C70DE237FFE92F9D7" bold="true" box="[151,288,1628,1653]" pageId="16" pageNumber="257">FIGURE 10.</emphasis>
<taxonomicName id="4C744D3AFFC3966C716E237EFDBFF9D7" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[295,525,1629,1653]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC3966C716E237EFDBFF9D7" box="[295,525,1629,1653]" italics="true" pageId="16" pageNumber="257">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC3966C725D237EFDD7F9D7" bold="true" box="[532,613,1629,1653]" pageId="16" pageNumber="257">
<taxonomicNameLabel id="A23357D0FFC3966C725D237EFDD7F9D7" box="[532,613,1629,1653]" pageId="16" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm.
</paragraph>
</caption>
<paragraph id="8BCB36B9FFC3966C708E23C8FC70F8EF" blockId="16.[151,1436,1771,1870]" pageId="16" pageNumber="257">
This is the only known
<taxonomicName id="4C744D3AFFC3966C719823C8FDDAF8A7" authorityName="Hammer" authorityYear="1958" box="[465,616,1771,1797]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAABFFC3966C719823C8FDDAF8A7" box="[465,616,1771,1797]" italics="true" pageId="16" pageNumber="257">Anderemaeus</emphasis>
</taxonomicName>
species with a strong, conspicuous tooth on the lamella, just posterior to the lamellar seta. Among neotropical species, only
<taxonomicName id="4C744D3AFFC3966C72932233FCEFF88B" authorityName="Hammer" authorityYear="1962" box="[730,861,1807,1834]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B900EAABFFC3966C72932233FCEFF88B" box="[730,861,1807,1834]" italics="true" pageId="16" pageNumber="257">A. chilensis</emphasis>
</taxonomicName>
shares the anteriorly displaced seta
<emphasis id="B900EAABFFC3966C74B12233FABCF88B" box="[1272,1294,1808,1833]" italics="true" pageId="16" pageNumber="257">in</emphasis>
, inserted far from the sejugal groove and distant from the pair of triangular tubercles.
</paragraph>
<paragraph id="8BCB36B9FFC3966C70DE22BBFF66F810" blockId="16.[151,1436,1944,2043]" box="[151,212,1944,1970]" pageId="16" pageNumber="257">
<emphasis id="B900EAABFFC3966C70DE22BBFF66F810" box="[151,212,1944,1970]" italics="true" pageId="16" pageNumber="257">Adult</emphasis>
</paragraph>
<paragraph id="8BCB36B9FFC3966C70DE229EFA9DF859" blockId="16.[151,1436,1944,2043]" pageId="16" pageNumber="257">
<emphasis id="B900EAABFFC3966C70DE229EFE8BF874" box="[151,313,1981,2006]" italics="true" pageId="16" pageNumber="257">Measurements</emphasis>
.
<typeStatus id="54CF881BFFC3966C7101229FFE02F874" box="[328,432,1980,2006]" pageId="16" pageNumber="257" type="holotype">Holotype</typeStatus>
(female) length 680, maximum notogastral width 431; range (including
<specimenCount id="9D72FD30FFC3966C74A9229EFAF7F874" box="[1248,1349,1980,2007]" pageId="16" pageNumber="257" type="female">4 female</specimenCount>
,
<specimenCount id="9D72FD30FFC3966C7519229EFA2EF874" box="[1360,1436,1980,2007]" pageId="16" pageNumber="257" type="male">3 male</specimenCount>
<typeStatus id="54CF881BFFC3966C70DE22C2FEB6F859" box="[151,260,2017,2043]" pageId="16" pageNumber="257" type="paratype">paratypes</typeStatus>
) 614680 × 398448. Females larger than males: 614640 × 415431 vs. 547597 × 365398.
</paragraph>
<paragraph id="8BCB36B9FFC2966D708E25B9FB01FF59" blockId="17.[151,1436,153,252]" pageId="17" pageNumber="258">
<emphasis id="B900EAABFFC2966D708E25B9FEF7FF11" box="[199,325,154,179]" italics="true" pageId="17" pageNumber="258">Integument</emphasis>
. Cuticle shiny in reflected light or dulled with organic and mineral debris attached to cerotegument (
<figureCitation id="134F2A3CFFC2966D70E9259EFE98FF75" box="[160,298,189,215]" captionStart="FIGURE 12" captionStartId="18.[151,250,1843,1868]" captionTargetBox="[164,1423,179,1816]" captionTargetId="figure@18.[164,1423,179,1816]" captionTargetPageId="18" captionText="FIGURE 12. Anderemaeus dentatus sp. nov., adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta h 1; F, notogastral seta p 1; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others)." httpUri="https://zenodo.org/record/3353077/files/figure.png" pageId="17" pageNumber="258">Fig. 12A, B</figureCitation>
). Cerotegument (
<figureCitation id="134F2A3CFFC2966D71B9259EFDF0FF75" box="[496,578,189,215]" captionStart="FIGURE 13" captionStartId="19.[151,250,1427,1452]" captionTargetBox="[151,1435,179,1399]" captionTargetId="figure@19.[151,1435,179,1399]" captionTargetPageId="19" captionText="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="17" pageNumber="258">Fig. 13</figureCitation>
) with larger excrescences columnar to phallus-shaped, covered in dust-like granules (
<figureCitation id="134F2A3CFFC2966D714C25C2FED7FF59" box="[261,357,225,251]" captionStart="FIGURE 13" captionStartId="19.[151,250,1427,1452]" captionTargetBox="[151,1435,179,1399]" captionTargetId="figure@19.[151,1435,179,1399]" captionTargetPageId="19" captionText="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="17" pageNumber="258">Fig. 13F</figureCitation>
insert); smaller excrescences amorphous to irregularly granular (
<figureCitation id="134F2A3CFFC2966D747625C2FB14FF59" box="[1087,1190,225,251]" captionStart="FIGURE 13" captionStartId="19.[151,250,1427,1452]" captionTargetBox="[151,1435,179,1399]" captionTargetId="figure@19.[151,1435,179,1399]" captionTargetPageId="19" captionText="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="17" pageNumber="258">Fig. 13B</figureCitation>
).
</paragraph>
<caption id="DF0B6631FFC2966D70DE2223FEA1F89F" httpUri="https://zenodo.org/record/3353075/files/figure.png" pageId="17" pageNumber="258" startId="17.[151,250,1792,1817]" targetBox="[152,1436,252,1752]" targetPageId="17">
<paragraph id="8BCB36B9FFC2966D70DE2223FEA1F89F" blockId="17.[151,1436,1792,1853]" pageId="17" pageNumber="258">
<emphasis id="B900EAABFFC2966D70DE2223FE92F8BB" bold="true" box="[151,288,1792,1817]" pageId="17" pageNumber="258">FIGURE 11.</emphasis>
<taxonomicName id="4C744D3AFFC2966D71612222FDBDF8BB" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[296,527,1793,1817]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="258" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC2966D71612222FDBDF8BB" box="[296,527,1793,1817]" italics="true" pageId="17" pageNumber="258">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC2966D725E2222FDDBF8BB" bold="true" box="[535,617,1793,1817]" pageId="17" pageNumber="258">
<taxonomicNameLabel id="A23357D0FFC2966D725E2222FDDBF8BB" box="[535,617,1793,1817]" pageId="17" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult (gnathosoma and legs omitted): A, lateral view; B, posterior view. Scale bar 100 μm.
</paragraph>
</caption>
<caption id="DF0B6631FFC1966E70DE2210FD4CF87E" httpUri="https://zenodo.org/record/3353077/files/figure.png" pageId="18" pageNumber="259" startId="18.[151,250,1843,1868]" targetBox="[164,1423,179,1816]" targetPageId="18">
<paragraph id="8BCB36B9FFC1966E70DE2210FD4CF87E" blockId="18.[151,1436,1843,2013]" pageId="18" pageNumber="259">
<emphasis id="B900EAABFFC1966E70DE2210FE93F8EE" bold="true" box="[151,289,1843,1868]" pageId="18" pageNumber="259">FIGURE 12.</emphasis>
<taxonomicName id="4C744D3AFFC1966E71612217FDBCF8EE" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[296,526,1844,1868]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="259" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC1966E71612217FDBCF8EE" box="[296,526,1844,1868]" italics="true" pageId="18" pageNumber="259">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC1966E725C2217FDD4F8EE" bold="true" box="[533,614,1844,1868]" pageId="18" pageNumber="259">
<taxonomicNameLabel id="A23357D0FFC1966E725C2217FDD4F8EE" box="[533,614,1844,1868]" pageId="18" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta
<emphasis id="B900EAABFFC1966E70A7225FFF49F836" box="[238,251,1916,1940]" italics="true" pageId="18" pageNumber="259">h</emphasis>
<subScript id="17F034FCFFC1966E70B222A8FEB0F83B" attach="left" box="[251,258,1931,1945]" fontSize="6" pageId="18" pageNumber="259">1</subScript>
; F, notogastral seta
<emphasis id="B900EAABFFC1966E7186225EFE6EF836" box="[463,476,1917,1940]" italics="true" pageId="18" pageNumber="259">p</emphasis>
<subScript id="17F034FCFFC1966E719222A8FE50F83B" attach="left" box="[475,482,1931,1945]" fontSize="6" pageId="18" pageNumber="259">1</subScript>
; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others).
</paragraph>
</caption>
<caption id="DF0B6631FFC0966F70DE20B0FDEEF99E" httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="19" pageNumber="260" startId="19.[151,250,1427,1452]" targetBox="[151,1435,179,1399]" targetPageId="19">
<paragraph id="8BCB36B9FFC0966F70DE20B0FDEEF99E" blockId="19.[151,1437,1427,1596]" pageId="19" pageNumber="260">
<emphasis id="B900EAABFFC0966F70DE20B0FE93FA0E" bold="true" box="[151,289,1427,1452]" pageId="19" pageNumber="260">FIGURE 13.</emphasis>
<taxonomicName id="4C744D3AFFC0966F716020B7FDBDFA0E" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[297,527,1428,1452]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="260" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC0966F716020B7FDBDFA0E" box="[297,527,1428,1452]" italics="true" pageId="19" pageNumber="260">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC0966F725E20B7FDDBFA0E" bold="true" box="[535,617,1428,1452]" pageId="19" pageNumber="260">
<taxonomicNameLabel id="A23357D0FFC0966F725E20B7FDDBFA0E" box="[535,617,1428,1452]" pageId="19" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B).
</paragraph>
</caption>
<paragraph id="8BCB36B9FFC0966F708E23ADFDCAF84E" blockId="19.[151,1437,1678,2029]" pageId="19" pageNumber="260">
<emphasis id="B900EAABFFC0966F708E23ADFEF6F90A" box="[199,324,1678,1704]" italics="true" pageId="19" pageNumber="260">Prodorsum</emphasis>
(
<figureCitation id="134F2A3CFFC0966F711F23ADFE77F90A" box="[342,453,1678,1704]" captionStart="FIGURE 10" captionStartId="16.[151,250,1628,1653]" captionTargetBox="[151,1436,366,1551]" captionTargetId="figure@16.[151,1436,342,1600]" captionTargetPageId="16" captionText="FIGURE 10. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353073/files/figure.png" pageId="19" pageNumber="260">Figs 10A</figureCitation>
,
<figureCitation id="134F2A3CFFC0966F719A23ADFDB4F90A" box="[467,518,1678,1704]" captionStart="FIGURE 11" captionStartId="17.[151,250,1792,1817]" captionTargetBox="[152,1436,252,1752]" captionTargetId="figure@17.[151,1436,270,1764]" captionTargetPageId="17" captionText="FIGURE 11. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, lateral view; B, posterior view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353075/files/figure.png" pageId="19" pageNumber="260">11A</figureCitation>
). Rostrum with terminal mucro, rostral bulge strongly developed. Lamella with complex form: broadened at mid-length, with additional strong triangular tooth projecting anterolaterally, just proximal to small cusp; continuing anteriorly as distinct prolamella. Space between lamellae distinctly foveate anteriorly; pair strongly convergent, prolamellae nearly touching anteriorly: mutual distance of setae
<emphasis id="B900EAABFFC0966F74FD23D9FB7AF8B6" box="[1204,1224,1786,1812]" italics="true" pageId="19" pageNumber="260">le</emphasis>
distinctly less than that of pair
<emphasis id="B900EAABFFC0966F715C223CFE9FF89A" box="[277,301,1823,1848]" italics="true" pageId="19" pageNumber="260">ro</emphasis>
. With distinct transverse prelamellar ridge (
<emphasis id="B900EAABFFC0966F735B223DFC81F89A" box="[786,819,1822,1848]" italics="true" pageId="19" pageNumber="260">plr</emphasis>
), length similar to
<emphasis id="B900EAABFFC0966F744A223CFBA9F89A" box="[1027,1051,1823,1848]" italics="true" pageId="19" pageNumber="260">ro</emphasis>
mutual distance. Setae
<emphasis id="B900EAABFFC0966F7556223CFA85F89A" box="[1311,1335,1823,1848]" italics="true" pageId="19" pageNumber="260">ro</emphasis>
(5361),
<emphasis id="B900EAABFFC0966F70DE2261FF19F8FE" box="[151,171,1858,1884]" italics="true" pageId="19" pageNumber="260">le</emphasis>
(7386) and
<emphasis id="B900EAABFFC0966F71092260FEEBF8FE" box="[320,345,1859,1884]" italics="true" pageId="19" pageNumber="260">ex</emphasis>
(3236) thin, attenuate, weakly barbed,
<emphasis id="B900EAABFFC0966F73552261FC82F8FE" box="[796,816,1858,1884]" italics="true" pageId="19" pageNumber="260">le</emphasis>
usually with accentuated bend. Seta
<emphasis id="B900EAABFFC0966F74842260FB51F8FE" box="[1229,1251,1859,1884]" italics="true" pageId="19" pageNumber="260">in</emphasis>
long (118127), relatively thick, acicular, barbed; inserted without distinct basal tubercle, well anterior of notogastral margin. With isolated, subtriangular interbothridial tubercle located posterior to but distant from each seta
<emphasis id="B900EAABFFC0966F748A22A8FB6BF806" box="[1219,1241,1931,1956]" italics="true" pageId="19" pageNumber="260">in</emphasis>
. Bothridial setae (118127) shorter than mutual distance of pair; baculiform or slightly tapered distally, with sparse, inconspicuous barbs, directed laterally or posterolaterally.
</paragraph>
<paragraph id="8BCB36B9FFC79668708E25B9FDFAFEC5" blockId="20.[151,1437,153,364]" pageId="20" pageNumber="261">
<emphasis id="B900EAABFFC79668708E25B9FEF1FF11" box="[199,323,154,179]" italics="true" pageId="20" pageNumber="261">Notogaster</emphasis>
(
<figureCitation id="134F2A3CFFC79668711D25BAFE73FF11" box="[340,449,153,179]" captionStart="FIGURE 10" captionStartId="16.[151,250,1628,1653]" captionTargetBox="[151,1436,366,1551]" captionTargetId="figure@16.[151,1436,342,1600]" captionTargetPageId="16" captionText="FIGURE 10. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353073/files/figure.png" pageId="20" pageNumber="261">Figs 10A</figureCitation>
,
<figureCitation id="134F2A3CFFC79668718425BAFE58FF11" box="[461,490,153,179]" captionStart="FIGURE 11" captionStartId="17.[151,250,1792,1817]" captionTargetBox="[152,1436,252,1752]" captionTargetId="figure@17.[151,1436,270,1764]" captionTargetPageId="17" captionText="FIGURE 11. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, lateral view; B, posterior view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353075/files/figure.png" pageId="20" pageNumber="261">11</figureCitation>
). With uniform, shallow foveolation visible in reflected light (
<figureCitation id="134F2A3CFFC7966874E725BAFAA0FF11" box="[1198,1298,153,179]" captionStart="FIGURE 12" captionStartId="18.[151,250,1843,1868]" captionTargetBox="[164,1423,179,1816]" captionTargetId="figure@18.[164,1423,179,1816]" captionTargetPageId="18" captionText="FIGURE 12. Anderemaeus dentatus sp. nov., adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta h 1; F, notogastral seta p 1; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others)." httpUri="https://zenodo.org/record/3353077/files/figure.png" pageId="20" pageNumber="261">Fig. 12B</figureCitation>
insert). Humeral process (
<emphasis id="B900EAABFFC79668710B259EFEDBFF75" box="[322,361,189,215]" italics="true" pageId="20" pageNumber="261">hpr</emphasis>
;
<figureCitation id="134F2A3CFFC79668713E259EFE67FF75" box="[375,469,189,215]" captionStart="FIGURE 12" captionStartId="18.[151,250,1843,1868]" captionTargetBox="[164,1423,179,1816]" captionTargetId="figure@18.[164,1423,179,1816]" captionTargetPageId="18" captionText="FIGURE 12. Anderemaeus dentatus sp. nov., adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta h 1; F, notogastral seta p 1; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others)." httpUri="https://zenodo.org/record/3353077/files/figure.png" pageId="20" pageNumber="261">Fig 12H</figureCitation>
) strongly developed. Notogastral setae of medium length (
<emphasis id="B900EAABFFC79668742E259EFBC7FF75" box="[1127,1141,189,215]" italics="true" pageId="20" pageNumber="261">h</emphasis>
<subScript id="17F034FCFFC79668743C25EDFBCFFF7E" attach="left" box="[1141,1149,206,220]" fontSize="6" pageId="20" pageNumber="261">1</subScript>
, 6165;
<emphasis id="B900EAABFFC796687496259DFB5FFF75" box="[1247,1261,190,215]" italics="true" pageId="20" pageNumber="261">p</emphasis>
<subScript id="17F034FCFFC7966874A425EDFB47FF7E" attach="left" box="[1261,1269,206,220]" fontSize="6" pageId="20" pageNumber="261">1</subScript>
, 5357; others 7786). Seta
<emphasis id="B900EAABFFC79668717A25C1FE8DFF59" box="[307,319,226,251]" italics="true" pageId="20" pageNumber="261">c</emphasis>
attenuate, barbed, curving posterolaterally, extending past lyrifissure
<emphasis id="B900EAABFFC79668742025C1FBCDFF59" box="[1129,1151,226,251]" italics="true" pageId="20" pageNumber="261">ia</emphasis>
; others thickened, erect, heavily barbed (
<figureCitation id="134F2A3CFFC7966871032426FE78FEBD" box="[330,458,261,287]" captionStart="FIGURE 13" captionStartId="19.[151,250,1427,1452]" captionTargetBox="[151,1435,179,1399]" captionTargetId="figure@19.[151,1435,179,1399]" captionTargetPageId="19" captionText="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="20" pageNumber="261">Fig. 13E, F</figureCitation>
), slightly tapered or sometimes slightly dilated distally, inserted on small tubercles. Seta
<emphasis id="B900EAABFFC7966870DE240AFF01FEE1" box="[151,179,297,323]" italics="true" pageId="20" pageNumber="261">lm</emphasis>
distinctly anterior to level of
<emphasis id="B900EAABFFC796687249240AFDA4FEE1" box="[512,534,297,323]" italics="true" pageId="20" pageNumber="261">la</emphasis>
, about equidistant between
<emphasis id="B900EAABFFC796687319240AFCD4FEE1" box="[848,870,297,323]" italics="true" pageId="20" pageNumber="261">la</emphasis>
,
<emphasis id="B900EAABFFC79668733D2409FC32FEE1" box="[884,896,298,323]" italics="true" pageId="20" pageNumber="261">c</emphasis>
; mutual distance of pair
<emphasis id="B900EAABFFC7966874D1240AFB06FEE1" box="[1176,1204,297,323]" italics="true" pageId="20" pageNumber="261">lm</emphasis>
only slightly greater than setal length;
<emphasis id="B900EAABFFC796687114246EFED9FEC5" box="[349,363,333,359]" italics="true" pageId="20" pageNumber="261">h</emphasis>
<subScript id="17F034FCFFC796687122247DFEC1FECE" attach="left" box="[363,371,350,364]" fontSize="6" pageId="20" pageNumber="261">2</subScript>
aligned with
<emphasis id="B900EAABFFC796687243246EFDAAFEC5" box="[522,536,333,359]" italics="true" pageId="20" pageNumber="261">h</emphasis>
<subScript id="17F034FCFFC796687251247DFD92FECE" attach="left" box="[536,544,350,364]" fontSize="6" pageId="20" pageNumber="261">1</subScript>
,
<emphasis id="B900EAABFFC796687262246EFD8BFEC5" box="[555,569,333,359]" italics="true" pageId="20" pageNumber="261">h</emphasis>
<subScript id="17F034FCFFC796687270247DFDF3FECE" attach="left" box="[569,577,350,364]" fontSize="6" pageId="20" pageNumber="261">3</subScript>
.
</paragraph>
<caption id="DF0B6631FFC7966870DE23BBFA81F977" httpUri="https://zenodo.org/record/3353081/files/figure.png" pageId="20" pageNumber="261" startId="20.[151,250,1688,1713]" targetBox="[160,1426,394,1630]" targetPageId="20">
<paragraph id="8BCB36B9FFC7966870DE23BBFA81F977" blockId="20.[151,1436,1688,1749]" pageId="20" pageNumber="261">
<emphasis id="B900EAABFFC7966870DE23BBFE93F913" bold="true" box="[151,289,1688,1713]" pageId="20" pageNumber="261">FIGURE 14.</emphasis>
<taxonomicName id="4C744D3AFFC79668716123BAFDBDF913" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[296,527,1689,1713]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="261" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC79668716123BAFDBDF913" box="[296,527,1689,1713]" italics="true" pageId="20" pageNumber="261">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC79668725F23BAFDDAF913" bold="true" box="[534,616,1689,1713]" pageId="20" pageNumber="261">
<taxonomicNameLabel id="A23357D0FFC79668725F23BAFDDAF913" box="[534,616,1689,1713]" pageId="20" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult gnathosoma: A, subcapitulum, ventral view (palp omitted); B, left adoral lip; C, right palp, adaxial view; D, left chelicera, adaxial view. Scale bars 20 μm (A, C to same scale; D), 10 μm (B).
</paragraph>
</caption>
<paragraph id="8BCB36B9FFC79668708E220BFC6DF857" blockId="20.[151,1437,1831,2038]" pageId="20" pageNumber="261">
<emphasis id="B900EAABFFC79668708E220BFEE4F8E0" box="[199,342,1832,1858]" italics="true" pageId="20" pageNumber="261">Coxisternum</emphasis>
(
<figureCitation id="134F2A3CFFC7966871212204FE63F8E3" box="[360,465,1831,1857]" captionStart="FIGURE 10" captionStartId="16.[151,250,1628,1653]" captionTargetBox="[151,1436,366,1551]" captionTargetId="figure@16.[151,1436,342,1600]" captionTargetPageId="16" captionText="FIGURE 10. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353073/files/figure.png" pageId="20" pageNumber="261">Fig. 10B</figureCitation>
). Epimere I with small area of foveation in anterolateral corner and on mentotectum (
<figureCitation id="134F2A3CFFC7966870D62268FEB6F8C7" box="[159,260,1867,1893]" captionStart="FIGURE 12" captionStartId="18.[151,250,1843,1868]" captionTargetBox="[164,1423,179,1816]" captionTargetId="figure@18.[164,1423,179,1816]" captionTargetPageId="18" captionText="FIGURE 12. Anderemaeus dentatus sp. nov., adult: A, dorsal view of dark, dirty specimen; B, dorsal view of lighter, cleaner specimen (insert: enlargement from central notogaster, showing shallow foveolae); C, ventral view; D, prodorsum; E, notogastral seta h 1; F, notogastral seta p 1; G, region of pedotectum I and epimere I, ventral view; H, left sejugal region, dorsal view; I, sagittal section of sejugal region (inset: closeup of transverse incision). Photomicrographs in transmitted light, DIC, except AC reflected light. Scale bars 100 μm (AC); 20 μm (all others)." httpUri="https://zenodo.org/record/3353077/files/figure.png" pageId="20" pageNumber="261">Fig 12G</figureCitation>
). Enantiophysis
<emphasis id="B900EAABFFC79668718B226FFE6EF8C4" box="[450,476,1868,1894]" italics="true" pageId="20" pageNumber="261">e3</emphasis>
absent, or represented by vaguely-defined, small posterior tubercle only. Tubercles bearing setae
<emphasis id="B900EAABFFC796687166224CFEF9F82B" box="[303,331,1903,1929]" italics="true" pageId="20" pageNumber="261">3b</emphasis>
,
<emphasis id="B900EAABFFC7966871112253FEC0F828" box="[344,370,1904,1930]" italics="true" pageId="20" pageNumber="261">3c</emphasis>
strong, isolated, that of
<emphasis id="B900EAABFFC796687234224CFD2BF82B" box="[637,665,1903,1929]" italics="true" pageId="20" pageNumber="261">3b</emphasis>
sometimes expressed as low mound extending across sejugal groove. Aggenital enantiophysis (
<emphasis id="B900EAABFFC7966871F322B7FE67F80C" box="[442,469,1940,1966]" italics="true" pageId="20" pageNumber="261">e4</emphasis>
) well developed across border
<emphasis id="B900EAABFFC79668737222B0FCDEF80F" box="[827,876,1939,1965]" italics="true" pageId="20" pageNumber="261">bo.4</emphasis>
, its posterior tubercle about twice as large as anterior; latter, bearing seta
<emphasis id="B900EAABFFC7966871F52294FE6BF873" box="[444,473,1975,2001]" italics="true" pageId="20" pageNumber="261">4b</emphasis>
, with low ridge running anteriorly across epimere IV. Epimeral setae thin, attenuate, weakly and sparsely barbed;
<emphasis id="B900EAABFFC79668719522FFFE44F854" box="[476,502,2012,2038]" italics="true" pageId="20" pageNumber="261">3c</emphasis>
and
<emphasis id="B900EAABFFC79668726422FFFDF5F854" box="[557,583,2012,2038]" italics="true" pageId="20" pageNumber="261">4c</emphasis>
(3641) longer than others (2024).
</paragraph>
<paragraph id="8BCB36B9FFC69669708E25BAFD02FF59" blockId="21.[151,1437,153,251]" pageId="21" pageNumber="262">
<emphasis id="B900EAABFFC69669708E25BAFE27FF11" box="[199,405,153,179]" italics="true" pageId="21" pageNumber="262">Anogenital region</emphasis>
(
<figureCitation id="134F2A3CFFC6966971E125BAFDA4FF11" box="[424,534,153,179]" captionStart="FIGURE 10" captionStartId="16.[151,250,1628,1653]" captionTargetBox="[151,1436,366,1551]" captionTargetId="figure@16.[151,1436,342,1600]" captionTargetPageId="16" captionText="FIGURE 10. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, dorsal view; B, ventral view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353073/files/figure.png" pageId="21" pageNumber="262">Figs 10B</figureCitation>
,
<figureCitation id="134F2A3CFFC69669726C25BAFDEAFF11" box="[549,600,153,179]" captionStart="FIGURE 11" captionStartId="17.[151,250,1792,1817]" captionTargetBox="[152,1436,252,1752]" captionTargetId="figure@17.[151,1436,270,1764]" captionTargetPageId="17" captionText="FIGURE 11. Anderemaeus dentatus sp. nov., adult (gnathosoma and legs omitted): A, lateral view; B, posterior view. Scale bar 100 μm." httpUri="https://zenodo.org/record/3353075/files/figure.png" pageId="21" pageNumber="262">11A</figureCitation>
). Setae thin, attenuate, smooth or sparsely barbed. Anterior genital seta (2024) longer than others (1216); aggenital (2024), anal (1216) and adanal (2024) setae in typical positions. Preanal organ only slightly expanded internally.
</paragraph>
<caption id="DF0B6631FFC6966970DE2295FC05F851" httpUri="https://zenodo.org/record/3353083/files/figure.png" pageId="21" pageNumber="262" startId="21.[151,250,1974,1999]" targetBox="[155,1430,282,1933]" targetPageId="21">
<paragraph id="8BCB36B9FFC6966970DE2295FC05F851" blockId="21.[151,1436,1974,2035]" pageId="21" pageNumber="262">
<emphasis id="B900EAABFFC6966970DE2295FEADF86D" bold="true" box="[151,287,1974,1999]" pageId="21" pageNumber="262">FIGURE 15.</emphasis>
<taxonomicName id="4C744D3AFFC69669716C2294FDBBF86D" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[293,521,1975,1999]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="262" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC69669716C2294FDBBF86D" box="[293,521,1975,1999]" italics="true" pageId="21" pageNumber="262">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC6966972462294FDEDF86D" bold="true" box="[527,607,1975,1999]" pageId="21" pageNumber="262">
<taxonomicNameLabel id="A23357D0FFC6966972462294FDEDF86D" box="[527,607,1975,1999]" pageId="21" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult legs (abaxial view): A, right leg I; B, trochanter, femur and genu of right leg II; C, trochanter, femur and genu of left leg III; D, left leg IV. Scale bar 50 μm.
</paragraph>
</caption>
<paragraph id="8BCB36B9FFC5966A708E25BAFD24FEBD" blockId="22.[151,1437,152,288]" pageId="22" pageNumber="263">
<emphasis id="B900EAABFFC5966A708E25BAFEE4FF11" box="[199,342,153,179]" italics="true" pageId="22" pageNumber="263">Gnathosoma</emphasis>
(
<figureCitation id="134F2A3CFFC5966A712E25BAFE09FF11" box="[359,443,153,179]" captionStart="FIGURE 14" captionStartId="20.[151,250,1688,1713]" captionTargetBox="[160,1426,394,1630]" captionTargetId="figure@20.[151,1436,376,1661]" captionTargetPageId="20" captionText="FIGURE 14. Anderemaeus dentatus sp. nov., adult gnathosoma: A, subcapitulum, ventral view (palp omitted); B, left adoral lip; C, right palp, adaxial view; D, left chelicera, adaxial view. Scale bars 20 μm (A, C to same scale; D), 10 μm (B)." httpUri="https://zenodo.org/record/3353081/files/figure.png" pageId="22" pageNumber="263">Fig. 14</figureCitation>
). Subcapitulum longer than wide (123135 × 90102). Subcapitular setae attenuate, inconspicuously roughened;
<emphasis id="B900EAABFFC5966A718F259EFE66FF75" box="[454,468,189,215]" italics="true" pageId="22" pageNumber="263">h</emphasis>
(2832) longer than
<emphasis id="B900EAABFFC5966A728E259DFD67FF75" box="[711,725,190,215]" italics="true" pageId="22" pageNumber="263">a</emphasis>
and
<emphasis id="B900EAABFFC5966A7347259DFC90FF75" box="[782,802,190,215]" italics="true" pageId="22" pageNumber="263">m</emphasis>
(2024). Adoral setae (1216) attenuate, smooth. Palps (9094), postpalpal seta (8), and chelicera (123135) typical of genus; cheliceral setae acuminate to attenuate, barbed,
<emphasis id="B900EAABFFC5966A70B82426FEABFEBD" box="[241,281,261,287]" italics="true" pageId="22" pageNumber="263">cha</emphasis>
(3641) longer than
<emphasis id="B900EAABFFC5966A724F2426FD9CFEBD" box="[518,558,261,287]" italics="true" pageId="22" pageNumber="263">chb</emphasis>
(2428).
</paragraph>
<caption id="DF0B6631FFC5966A70DE2220FAC8F82A" httpUri="https://zenodo.org/record/3353085/files/figure.png" pageId="22" pageNumber="263" startId="22.[151,250,1795,1820]" targetBox="[151,1435,288,1767]" targetPageId="22">
<paragraph id="8BCB36B9FFC5966A70DE2220FAC8F82A" blockId="22.[151,1437,1795,1928]" pageId="22" pageNumber="263">
<emphasis id="B900EAABFFC5966A70DE2220FEADF8BE" bold="true" box="[151,287,1795,1820]" pageId="22" pageNumber="263">FIGURE 16.</emphasis>
<taxonomicName id="4C744D3AFFC5966A716C2227FDB8F8BE" authority="Norton &amp; Ermilov, 2019" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[293,522,1796,1820]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="263" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="B900EAABFFC5966A716C2227FDB8F8BE" box="[293,522,1796,1820]" italics="true" pageId="22" pageNumber="263">Anderemaeus dentatus</emphasis>
</taxonomicName>
<emphasis id="B900EAABFFC5966A72592227FDD2F8BE" bold="true" box="[528,608,1796,1820]" pageId="22" pageNumber="263">
<taxonomicNameLabel id="A23357D0FFC5966A72592227FDD2F8BE" box="[528,608,1796,1820]" pageId="22" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymphs: A, tritonymph, dorsal view (larger hysterosomal setae broken except
<emphasis id="B900EAABFFC5966A75CB2227FA24F8BE" box="[1410,1430,1796,1820]" italics="true" pageId="22" pageNumber="263">lp</emphasis>
,
<emphasis id="B900EAABFFC5966A70DE220BFF16F8E2" box="[151,164,1832,1856]" italics="true" pageId="22" pageNumber="263">h</emphasis>
<subScript id="17F034FCFFC5966A70ED2215FF19F8E6" attach="left" box="[164,171,1846,1860]" fontSize="6" pageId="22" pageNumber="263">1</subScript>
); B, same, lateral view (inserts: close-up of seta
<emphasis id="B900EAABFFC5966A72DC220BFD15F8E2" box="[661,679,1832,1856]" italics="true" pageId="22" pageNumber="263">le</emphasis>
in upper right, bothridial region in lower left); C, as in A, close-up of bothridial seta; D, as in A, close-up of stalk bearing opening to opisthonotal gland; E, deutonymph, leg I, adaxial view; F, tritonymph, prodorsum, dorsal view. Photomicrographs in transmitted light, DIC. Scale bars:100 μm (A, B); 20 μm (C, E, F); 5 μm (D).
</paragraph>
</caption>
<paragraph id="8BCB36B9FFC4966B708E25B9FDD1FE29" blockId="23.[151,1437,152,395]" pageId="23" pageNumber="264">
<emphasis id="B900EAABFFC4966B708E25B9FEB6FF11" box="[199,260,154,179]" italics="true" pageId="23" pageNumber="264">Legs.</emphasis>
Form and proportions of segments shown in
<figureCitation id="134F2A3CFFC4966B734D25BAFCE5FF11" box="[772,855,152,179]" captionStart="FIGURE 15" captionStartId="21.[151,250,1974,1999]" captionTargetBox="[155,1430,282,1933]" captionTargetId="figure@21.[151,1436,270,1946]" captionTargetPageId="21" captionText="FIGURE 15. Anderemaeus dentatus sp. nov., adult legs (abaxial view): A, right leg I; B, trochanter, femur and genu of right leg II; C, trochanter, femur and genu of left leg III; D, left leg IV. Scale bar 50 μm." httpUri="https://zenodo.org/record/3353083/files/figure.png" pageId="23" pageNumber="264">Fig. 15</figureCitation>
. Tarsi significantly (I, II) to slightly (III, IV) longer than respective tibiae. Genua I, II notably longer than III, IV. Femora III, IV only slightly longer than wide, ventral keel moderate on III, large, blade-like on IV (
<figureCitation id="134F2A3CFFC4966B72DB25C2FD45FF59" box="[658,759,225,251]" captionStart="FIGURE 13" captionStartId="19.[151,250,1427,1452]" captionTargetBox="[151,1435,179,1399]" captionTargetId="figure@19.[151,1435,179,1399]" captionTargetPageId="19" captionText="FIGURE 13. Anderemaeus dentatus sp. nov., adult, cerotegument (separated from cuticle in B, partly dislodged in C, D, F): A, region of circumpedal carina, lateral view; B, lateral podosomal region; C, femur IV; D, humeral tubercle, lateral view; E, bothridium, lateral view; F, partial trochanter and femur III (insert: enlargement of dislodged cerotegument); G, subcapitulum, dorsal view, near base of right palp; H, trochanter and base of femur IV, paraxial view. Photomicrographs in transmitted light, DIC. Scale bars: 10 μm (A, CH); 5 μm (B)." httpUri="https://zenodo.org/record/3353079/files/figure.png" pageId="23" pageNumber="264">Fig. 13C</figureCitation>
), neither with tooth or spine. Trochanters III, IV with prominent dorsodistal spine; III with smaller proximal spine, IV with small proximal tooth. Setation typical of genus; form and locations of setae and solenidia shown in
<figureCitation id="134F2A3CFFC4966B72ED240AFD4AFEE1" box="[676,760,296,323]" captionStart="FIGURE 15" captionStartId="21.[151,250,1974,1999]" captionTargetBox="[155,1430,282,1933]" captionTargetId="figure@21.[151,1436,270,1946]" captionTargetPageId="21" captionText="FIGURE 15. Anderemaeus dentatus sp. nov., adult legs (abaxial view): A, right leg I; B, trochanter, femur and genu of right leg II; C, trochanter, femur and genu of left leg III; D, left leg IV. Scale bar 50 μm." httpUri="https://zenodo.org/record/3353083/files/figure.png" pageId="23" pageNumber="264">Fig. 15</figureCitation>
. Solenidion
<emphasis id="B900EAABFFC4966B73CF2409FC26FEE1" box="[902,916,298,323]" italics="true" pageId="23" pageNumber="264">σ</emphasis>
of genu I subflagellate, distinctly longer than segment; that of other genua shorter than segment (II) or of similar length (III);
<emphasis id="B900EAABFFC4966B7441246DFBA5FEC5" box="[1032,1047,334,359]" italics="true" pageId="23" pageNumber="264">φ</emphasis>
of tibiae IIIV subflagellate, equal or only slightly less than segment length.
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE24FEFE72FE5A" blockId="23.[151,448,477,534]" box="[151,448,477,504]" pageId="23" pageNumber="264">
<emphasis id="B900EAABFFC4966B70DE24FEFF4DFE55" box="[151,255,477,503]" italics="true" pageId="23" pageNumber="264">Juveniles</emphasis>
(larva unknown)
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE24DEFF42FDB4" blockId="23.[151,448,477,534]" box="[151,240,509,534]" pageId="23" pageNumber="264">
(
<figureCitation id="134F2A3CFFC4966B70D624DEFF5AFDB4" box="[159,232,509,534]" captionStart="FIGURE 16" captionStartId="22.[151,250,1795,1820]" captionTargetBox="[151,1435,288,1767]" captionTargetId="figure@22.[151,1435,306,1767]" captionTargetPageId="22" captionText="FIGURE 16. Anderemaeus dentatus sp. nov., nymphs: A, tritonymph, dorsal view (larger hysterosomal setae broken except lp, h 1); B, same, lateral view (inserts: close-up of seta le in upper right, bothridial region in lower left); C, as in A, close-up of bothridial seta; D, as in A, close-up of stalk bearing opening to opisthonotal gland; E, deutonymph, leg I, adaxial view; F, tritonymph, prodorsum, dorsal view. Photomicrographs in transmitted light, DIC. Scale bars:100 μm (A, B); 20 μm (C, E, F); 5 μm (D)." httpUri="https://zenodo.org/record/3353085/files/figure.png" pageId="23" pageNumber="264">Fig. 16</figureCitation>
)
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE2767FB1EFD20" blockId="23.[151,1436,579,642]" pageId="23" pageNumber="264">
Length and width of protonymph 310359 × 199 (n =2), deutonymph 426 × 245 (n = 1), tritonymph 582 × 348 (n=1). Bothridial seta distinctly tapered and consistently, strongly curved anteriad (
<figureCitation id="134F2A3CFFC4966B7471274BFB2DFD20" box="[1080,1183,615,642]" captionStart="FIGURE 16" captionStartId="22.[151,250,1795,1820]" captionTargetBox="[151,1435,288,1767]" captionTargetId="figure@22.[151,1435,306,1767]" captionTargetPageId="22" captionText="FIGURE 16. Anderemaeus dentatus sp. nov., nymphs: A, tritonymph, dorsal view (larger hysterosomal setae broken except lp, h 1); B, same, lateral view (inserts: close-up of seta le in upper right, bothridial region in lower left); C, as in A, close-up of bothridial seta; D, as in A, close-up of stalk bearing opening to opisthonotal gland; E, deutonymph, leg I, adaxial view; F, tritonymph, prodorsum, dorsal view. Photomicrographs in transmitted light, DIC. Scale bars:100 μm (A, B); 20 μm (C, E, F); 5 μm (D)." httpUri="https://zenodo.org/record/3353085/files/figure.png" pageId="23" pageNumber="264">Fig. 16C</figureCitation>
).
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE27F7FEBDFD4C" blockId="23.[151,1437,724,822]" box="[151,271,724,750]" pageId="23" pageNumber="264">
<emphasis id="B900EAABFFC4966B70DE27F7FEBDFD4C" box="[151,271,724,750]" italics="true" pageId="23" pageNumber="264">Etymology</emphasis>
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE27DBFF6FFC94" blockId="23.[151,1437,724,822]" pageId="23" pageNumber="264">
The specific name
<taxonomicName id="4C744D3AFFC4966B712627DBFE7CFCB0" authorityName="Norton &amp; Ermilov" authorityYear="2019" box="[367,462,760,786]" class="Arachnida" family="Caleremaeidae" genus="Anderemaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="264" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B900EAABFFC4966B712627DBFE7CFCB0" box="[367,462,760,786]" italics="true" pageId="23" pageNumber="264">dentatus</emphasis>
</taxonomicName>
(toothed) is a Latin adjective that refers to the conspicuous tooth on the prodorsal lamella.
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE26ABFED9FC00" blockId="23.[151,1436,904,1147]" box="[151,363,904,930]" pageId="23" pageNumber="264">
<emphasis id="B900EAABFFC4966B70DE26ABFED9FC00" box="[151,363,904,930]" italics="true" pageId="23" pageNumber="264">Material examined</emphasis>
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE268FFDB4FBD9" blockId="23.[151,1436,904,1147]" pageId="23" pageNumber="264">
<materialsCitation id="3B1C3CE4FFC4966B70DE268FFBD4FC48" ID-GBIF-Occurrence="2366524998" collectingDate="1982-12-26" collectorName="A. Newton &amp; M. Thayer &amp; Valdivian" country="Chile" county="Osorno Province" elevation="430" location="Anticura" municipality="Parque Nacional Puyehue" pageId="23" pageNumber="264" specimenCount="1" stateProvince="Los Lagos" typeStatus="holotype">
<typeStatus id="54CF881BFFC4966B70DE268FFF4DFC64" box="[151,255,940,966]" pageId="23" pageNumber="264" type="holotype">Holotype</typeStatus>
(female):
<collectingCountry id="F3637629FFC4966B713A268FFE00FC64" box="[371,434,940,966]" name="Chile" pageId="23" pageNumber="264">Chile</collectingCountry>
,
<collectingRegion id="49B0F85BFFC4966B71F7268FFD3FFC64" box="[446,653,940,966]" country="Chile" name="Los Lagos" pageId="23" pageNumber="264">Los Lagos Region</collectingRegion>
,
<collectingCounty id="62AA4E35FFC4966B72D1268FFCE5FC64" box="[664,855,940,966]" pageId="23" pageNumber="264">Osorno Province</collectingCounty>
,
<collectingMunicipality id="6BAFACC3FFC4966B732B268FFB37FC64" box="[866,1157,940,966]" pageId="23" pageNumber="264">Parque Nacional Puyehue</collectingMunicipality>
,
<quantity id="4C8C9B5CFFC4966B74D8268FFB6CFC64" box="[1169,1246,940,966]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="23" pageNumber="264" unit="km" value="4.1">4.1 km</quantity>
east of
<location id="8EAB6062FFC4966B757A268FFA2AFC64" LSID="urn:lsid:plazi:treatment:03DD87AFFFDC966B70DE2216FEEBFAE6:8EAB6062FFC4966B757A268FFA2AFC64" box="[1331,1432,940,966]" country="Chile" county="Osorno Province" municipality="Parque Nacional Puyehue" name="Anticura" pageId="23" pageNumber="264" stateProvince="Los Lagos">Anticura</location>
,
<elevation id="0059D18AFFC4966B70DE26F3FEADFC48" box="[151,287,976,1002]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="23" pageNumber="264" unit="m" value="430.0">
<quantity id="4C8C9B5CFFC4966B70DE26F3FF52FC48" box="[151,224,976,1002]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="23" pageNumber="264" unit="m" value="430.0">430 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCA1079FFC4966B716426ECFE0FFC4B" box="[301,445,975,1001]" pageId="23" pageNumber="264" value="1982-12-26">
<collectingDate id="EF8EE991FFC4966B716426ECFE0FFC4B" box="[301,445,975,1001]" pageId="23" pageNumber="264" value="1982-12-26">26-XII-1982</collectingDate>
</date>
,
<collectorName id="2681536FFFC4966B718026F3FDF4FC48" box="[457,582,976,1002]" pageId="23" pageNumber="264">A. Newton</collectorName>
and
<collectorName id="2681536FFFC4966B72C926F3FD48FC48" box="[640,762,976,1002]" pageId="23" pageNumber="264">M. Thayer</collectorName>
, from
<collectorName id="2681536FFFC4966B730F26F3FC03FC48" box="[838,945,976,1002]" pageId="23" pageNumber="264">Valdivian</collectorName>
rainforest litter
</materialsCitation>
.
<materialsCitation id="3B1C3CE4FFC4966B743B26F3FCD3FB90" ID-GBIF-Occurrence="2366524997" collectingDate="1985-01-03" collectorName="N. Platnick &amp; O. Francke" country="Chile" county="Osorno Province" elevation="425" location="Valdivian forest" municipality="Parque Nacional Puyehue" pageId="23" pageNumber="264" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Los Lagos" typeStatus="paratype">
<typeStatus id="54CF881BFFC4966B743B26F3FB51FC48" box="[1138,1251,976,1002]" pageId="23" pageNumber="264" type="paratype">Paratypes</typeStatus>
: 7 (
<specimenCount id="9D72FD30FFC4966B754626F3FA33FC48" box="[1295,1409,976,1002]" pageId="23" pageNumber="264" type="female">4 females</specimenCount>
,
<specimenCount id="9D72FD30FFC4966B75C726F3FF68FBAD" pageId="23" pageNumber="264" type="male">3 males</specimenCount>
) with same data as holotype; 12 (unsexed) from Parque Nacional Puyehue, Aguas Caliente,
<elevation id="0059D18AFFC4966B754426D0FA25FBAC" box="[1293,1431,1011,1038]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="23" pageNumber="264" unit="m" value="425.0">
<quantity id="4C8C9B5CFFC4966B754426D0FAE5FBAC" box="[1293,1367,1011,1038]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="23" pageNumber="264" unit="m" value="425.0">425 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCA1079FFC4966B70DE2134FF4EFB90" box="[151,252,1047,1074]" pageId="23" pageNumber="264" value="1985-01-03">
<collectingDate id="EF8EE991FFC4966B70DE2134FF4EFB90" box="[151,252,1047,1074]" pageId="23" pageNumber="264" value="1985-01-03">3-I-1985</collectingDate>
</date>
,
<collectorName id="2681536FFFC4966B7143213BFE3CFB90" box="[266,398,1048,1074]" pageId="23" pageNumber="264">N. Platnick</collectorName>
,
<collectorName id="2681536FFFC4966B71D5213BFDACFB90" box="[412,542,1048,1074]" pageId="23" pageNumber="264">O. Francke</collectorName>
, from
<location id="8EAB6062FFC4966B7222213BFC92FB90" LSID="urn:lsid:plazi:treatment:03DD87AFFFDC966B70DE2216FEEBFAE6:8EAB6062FFC4966B7222213BFC92FB90" box="[619,800,1048,1074]" name="Valdivian forest" pageId="23" pageNumber="264">Valdivian forest</location>
litter
</materialsCitation>
. Other material:
<collectingRegion id="49B0F85BFFC4966B746C213BFB58FB90" box="[1061,1258,1048,1074]" country="Chile" name="Los Rios" pageId="23" pageNumber="264">Los Ríos Region</collectingRegion>
, Valdivia Province, Rt. 39, 30 km east of
<collectingRegion id="49B0F85BFFC4966B7181211FFDF2FBF4" box="[456,576,1084,1110]" country="Chile" name="Los Lagos" pageId="23" pageNumber="264">Los Lagos</collectingRegion>
, near Panguilulli,
<date id="FFCA1079FFC4966B73442118FC23FBF4" box="[781,913,1083,1110]" pageId="23" pageNumber="264" value="2005-03-11">11-III-2005</date>
, E. Toledo, col. (
<specimenCount id="9D72FD30FFC4966B741B211FFB0EFBF4" box="[1106,1212,1084,1110]" pageId="23" pageNumber="264" type="adult">13 adults</specimenCount>
, 1 trito-, 1 deuto-, 2 protonymphs; habitat unknown).
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE219CFEFAFB7A" blockId="23.[151,1436,1214,1348]" box="[151,328,1214,1240]" pageId="23" pageNumber="264">
<emphasis id="B900EAABFFC4966B70DE219CFEFAFB7A" box="[151,328,1214,1240]" italics="true" pageId="23" pageNumber="264">Type deposition</emphasis>
</paragraph>
<paragraph id="8BCB36B9FFC4966B70DE21C1FEEBFAE6" blockId="23.[151,1436,1214,1348]" pageId="23" pageNumber="264">
The
<typeStatus id="54CF881BFFC4966B708321C1FE9EFB5E" box="[202,300,1250,1276]" pageId="23" pageNumber="264" type="holotype">holotype</typeStatus>
is deposited in the SMNH; five
<typeStatus id="54CF881BFFC4966B72D021C0FCB6FB5F" box="[665,772,1251,1277]" pageId="23" pageNumber="264" type="paratype">paratypes</typeStatus>
are deposited in the TSUMZ, one in the UCMZ, preserved in ethanol with a drop of glycerol. The remainder of
<typeStatus id="54CF881BFFC4966B72932024FCF7FA83" box="[730,837,1287,1313]" pageId="23" pageNumber="264" type="paratype">paratypes</typeStatus>
are in the personal collection of RAN, three on slides, others in ethanol.
</paragraph>
</subSubSection>
</treatment>
</document>