177 lines
19 KiB
XML
177 lines
19 KiB
XML
<document id="805DC094A0C3295076BBD3439ED7A7EA" ID-DOI="10.11646/zootaxa.3640.3.2" ID-GBIF-Dataset="44e24dab-b3fb-4719-a84d-fa667ab6c4ac" ID-ISSN="1175-5326" ID-Zenodo-Dep="223281" ID-ZooBank="0101E600-90AB-4EDE-9F3C-5C9075BE6066" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460593625392" checkinUser="plazi" docAuthor="Sartori, André F., Gargominy, Olivier & Fontaine, Benoît" docDate="2013" docId="03DD87A3FF88D45655AC7ACCFBD6D5A1" docLanguage="en" docName="zt03640p372.pdf" docOrigin="Zootaxa 3640 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Minidonta bieleri Sartori, Gargominy & Fontaine, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="365" masterDocId="FFE4FFDBFF9CD440553B7C11FFB9D269" masterDocTitle="Anthropogenic extinction of Pacific land snails: A case study of Rurutu, French Polynesia, with description of eight new species of endodontids (Pulmonata)" masterLastPageNumber="372" masterPageNumber="343" pageNumber="363" updateTime="1698326412613" updateUser="plazi">
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<mods:mods id="632FFDB9CC7EF22377345AA91A492680" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="430E9D4A761375AE02C075A4D3C7C2F1">
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<mods:title id="83F9E263D5FB3FB5152F2A61433E3643">Anthropogenic extinction of Pacific land snails: A case study of Rurutu, French Polynesia, with description of eight new species of endodontids (Pulmonata)</mods:title>
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</mods:titleInfo>
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<mods:name id="5E9970E75976A13292DACED50FCB48F8" type="personal">
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<mods:role id="8880B0B3AD76A8A3D41FD1E87672ED23">
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<mods:roleTerm id="45EC70FEF310C12372DF7383DAE0E95D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="795CD2497A57C40B99F244ED83D5B861">Sartori, André F.</mods:namePart>
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</mods:name>
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<mods:name id="B698357253D95AEB5BD847236BE7374C" type="personal">
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<mods:role id="B8066EB4241AAA00431AC1289EBF3AC5">
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<mods:roleTerm id="D663C3AEE7C25D9264117470D2112993">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4C394BB7C25C5ED1B511A9EDC8D24960">Gargominy, Olivier</mods:namePart>
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</mods:name>
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<mods:name id="1FB3222784D9A718573DAE5C968580A2" type="personal">
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<mods:role id="86A27A4A5447E2BAC762B8CD079AEF01">
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<mods:roleTerm id="79CB0D25BC52D6B4A3B4E2964B35E3A7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F5E8895D8489C6F1D04ECC2E79DD6745">Fontaine, Benoît</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="19DCA5C3D2926CF649BDC685BBE8C5D5">text</mods:typeOfResource>
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<mods:titleInfo id="09E6344303297ACF79F3E7D1B67664CD">
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<mods:title id="0A09DBB81AE63633057BE42B77D62214">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7278569BF5B9607BCFB8296A9F407125">
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<mods:date id="A3EC007CE56E95A7FFAFB9AA1EE1C275">2013</mods:date>
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<mods:detail id="0436344A88E05D5354C0D61A46D2321D" type="volume">
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<mods:number id="D71AE6A6C6295E1B6EB934C9489B3CC9">3640</mods:number>
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</mods:detail>
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<mods:detail id="1E3E90735805F17DBAB1B191B5BB3574" type="issue">
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<mods:number id="B301EC7B4DDFF45C4B441814A53BB82A">3</mods:number>
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<mods:start id="DC160FF8EDE175CE1BB09101EFD4F762">343</mods:start>
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<mods:end id="080B634C6F9873D2B3C9442B8E87568D">372</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="6D2805F59F1C8F3765BA54AB6D957577">journal article</mods:classification>
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<mods:identifier id="AF65E8FF30B06413318C5811A731DCA4" type="DOI">10.11646/zootaxa.3640.3.2</mods:identifier>
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<mods:identifier id="13089F05BDA5E0F32BC9A293F4BB01DD" type="GBIF-Dataset">44e24dab-b3fb-4719-a84d-fa667ab6c4ac</mods:identifier>
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<mods:identifier id="54C0533C176F0073A3387C0507E23A64" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9F8273291DBA118582C0B666D0300D4A" type="Zenodo-Dep">223281</mods:identifier>
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<mods:identifier id="3E3BB808F4193EC8151BD2566A33FA51" type="ZooBank">0101E600-90AB-4EDE-9F3C-5C9075BE6066</mods:identifier>
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</mods:mods>
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<treatment id="03DD87A3FF88D45655AC7ACCFBD6D5A1" ID-DOI="http://doi.org/10.5281/zenodo.5615966" ID-GBIF-Taxon="119566233" ID-Zenodo-Dep="5615966" LSID="urn:lsid:plazi:treatment:03DD87A3FF88D45655AC7ACCFBD6D5A1" httpUri="http://treatment.plazi.org/id/03DD87A3FF88D45655AC7ACCFBD6D5A1" lastPageId="22" lastPageNumber="365" pageId="20" pageNumber="363">
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<subSubSection id="C36E653EFF88D45455AC7ACCFE7BD571" pageId="20" pageNumber="363" type="nomenclature">
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<paragraph id="8BCB36B5FF88D45455AC7ACCFE74D49E" blockId="20.[151,461,1757,1816]" box="[151,461,1757,1783]" pageId="20" pageNumber="363">
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<heading id="D08381D9FF88D45455AC7ACCFE74D49E" bold="true" box="[151,461,1757,1783]" fontSize="11" level="1" pageId="20" pageNumber="363" reason="1">
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<emphasis id="B900EAA7FF88D45455AC7ACCFE74D49E" bold="true" box="[151,461,1757,1783]" pageId="20" pageNumber="363">
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<taxonomicName id="4C744D36FF88D45455AC7ACCFED0D49E" ID-CoL="7S5SY" box="[151,361,1757,1783]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="bieleri" status="sp. nov.">
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<emphasis id="B900EAA7FF88D45455AC7ACCFED0D49E" bold="true" box="[151,361,1757,1783]" italics="true" pageId="20" pageNumber="363">Minidonta bieleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23357DCFF88D454544B7ACCFE74D49E" box="[368,461,1757,1783]" pageId="20" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCB36B5FF88D45455AC7B11FE7BD571" blockId="20.[151,461,1757,1816]" box="[151,450,1792,1816]" pageId="20" pageNumber="363">
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<figureCitation id="134F2A30FF88D45455AC7B11FEB6D571" box="[151,271,1792,1816]" captionStart="FIGURE 10" captionStartId="19.[151,250,1773,1796]" captionTargetBox="[151,1435,519,1751]" captionTargetId="figure@19.[151,1436,519,1751]" captionTargetPageId="19" captionText="FIGURE 10. Holotypes of: A. Australdonta teaae sp. nov.; B. Minidonta boucheti sp. nov.; C. M. bieleri sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/223290/files/figure.png" pageId="20" pageNumber="363">Figures 10</figureCitation>
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C, 13, 14H, 15F
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E653EFF88D45455AC7B59FE50D5ED" pageId="20" pageNumber="363" type="materials_examined">
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<paragraph id="8BCB36B5FF88D45455AC7B59FE50D5ED" blockId="20.[151,1436,1864,2032]" pageId="20" pageNumber="363">
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<emphasis id="B900EAA7FF88D45455AC7B59FE39D508" bold="true" box="[151,384,1864,1889]" pageId="20" pageNumber="363">Examined material</emphasis>
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(
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<specimenCount id="9D72FD3CFF88D45454AE7B59FD90D509" box="[405,553,1864,1888]" pageId="20" pageNumber="363" type="generic">4 specimens</specimenCount>
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).
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<emphasis id="B900EAA7FF88D45457047B58FD15D508" bold="true" box="[575,684,1865,1889]" pageId="20" pageNumber="363">
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<typeStatus id="54CF8817FF88D45457047B58FD15D508" box="[575,684,1865,1889]" pageId="20" pageNumber="363" type="holotype">Holotype</typeStatus>
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</emphasis>
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: MNHN 25209, Rr8.
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<emphasis id="B900EAA7FF88D454568E7B58FB8FD508" bold="true" box="[949,1078,1865,1889]" pageId="20" pageNumber="363">
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<typeStatus id="54CF8817FF88D454568E7B58FB88D508" box="[949,1073,1865,1889]" pageId="20" pageNumber="363" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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MNHN 25210, 1 shell, Rr35; MNHN 25211, 2 shells, Rr66.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E653EFF88D45455FC7B81FE6CD599" pageId="20" pageNumber="363" type="diagnosis">
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<paragraph id="8BCB36B5FF88D45455FC7B81FE6CD599" blockId="20.[151,1436,1864,2032]" pageId="20" pageNumber="363">
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<emphasis id="B900EAA7FF88D45455FC7B81FE83D5C0" bold="true" box="[199,314,1936,1961]" pageId="20" pageNumber="363">Diagnosis</emphasis>
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: Shell with rounded periphery, coarse primary axial sculpture and prominent apertural barriers (four barriers and three traces on palatal wall; five barriers on parietal); umbilicus very narrow (width less than one tenth of the diameter of the shell).
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</paragraph>
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</subSubSection>
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<caption id="DF0B663DFF89D45555AC79D0FDC8D45D" httpUri="https://zenodo.org/record/223291/files/figure.png" pageId="21" pageNumber="364" targetBox="[151,1435,193,1451]" targetPageId="21">
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<paragraph id="8BCB36B5FF89D45555AC79D0FDC8D45D" blockId="21.[151,1436,1473,1588]" pageId="21" pageNumber="364">
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<emphasis id="B900EAA7FF89D45555AC79D0FEA6D7BE" bold="true" box="[151,287,1473,1496]" pageId="21" pageNumber="364">FIGURE 11.</emphasis>
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<taxonomicName id="4C744D36FF89D455541E79D3FE5ED7B1" box="[293,487,1474,1496]" class="Gastropoda" family="Endodontidae" genus="Australdonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="teaae" status="sp. nov.">
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<emphasis id="B900EAA7FF89D455541E79D3FE5ED7B1" box="[293,487,1474,1496]" italics="true" pageId="21" pageNumber="364">Australdonta teaae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23357DCFF89D45554D579D2FD84D7B1" box="[494,573,1475,1496]" pageId="21" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B900EAA7FF89D455577F79D3FDEFD7BE" bold="true" box="[580,598,1474,1495]" pageId="21" pageNumber="364">A</emphasis>
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. Holotype (MNHN 25205), apical view;
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<emphasis id="B900EAA7FF89D45556C779D3FBB4D7BE" bold="true" box="[1020,1037,1474,1495]" pageId="21" pageNumber="364">B</emphasis>
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and
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<emphasis id="B900EAA7FF89D455517B79D0FBEBD7B1" bold="true" box="[1088,1106,1473,1496]" pageId="21" pageNumber="364">C</emphasis>
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. Paratype 6 (MNHN 25206);
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<emphasis id="B900EAA7FF89D45550BF79D3FA2CD7BE" bold="true" box="[1412,1429,1474,1495]" pageId="21" pageNumber="364">B</emphasis>
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. Sculpture of the protoconch and early teleoconch;
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<emphasis id="B900EAA7FF89D45557AC79F1FD10D79E" bold="true" box="[663,681,1504,1527]" pageId="21" pageNumber="364">C</emphasis>
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. Sculpture of the late teleoconch;
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<emphasis id="B900EAA7FF89D455513E79F0FBAED79F" bold="true" box="[1029,1047,1505,1526]" pageId="21" pageNumber="364">D</emphasis>
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. Paratype 2 (MNHN 25206), detail of the peristome;
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<emphasis id="B900EAA7FF89D45554167A11FE87D47C" bold="true" box="[301,318,1536,1557]" pageId="21" pageNumber="364">E</emphasis>
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. Paratype 1 (MNHN 25206), displaying a moderately raised spire;
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<emphasis id="B900EAA7FF89D45556D17A11FC4ED47C" bold="true" box="[1002,1015,1536,1557]" pageId="21" pageNumber="364">F</emphasis>
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. Paratype 2, displaying a flat spire. Scale bars: A = 1 mm; B–D = 400 µm; E–F = 2 mm.
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</paragraph>
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</caption>
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<subSubSection id="C36E653EFF89D45655FC7A71FAA9D3EE" lastPageId="22" lastPageNumber="365" pageId="21" pageNumber="364" type="description">
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<paragraph id="8BCB36B5FF89D45655FC7A71FAA9D3EE" blockId="21.[151,1437,1632,2017]" lastBlockId="22.[151,1437,151,391]" lastPageId="22" lastPageNumber="365" pageId="21" pageNumber="364">
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<emphasis id="B900EAA7FF89D45555FC7A71FEE8D410" bold="true" box="[199,337,1632,1657]" pageId="21" pageNumber="364">Description</emphasis>
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: Shell depressed, white, with dark reddish-brown, scattered flammulations on the apical surface and shell base. Shell thin, opaque; periostracum adherent, matt. Apex barely raised, spire elevated, later whorls descending more rapidly. Apical and umbilical sutures impressed; whorls and periphery rounded. Transition between protoconch and teleoconch indistinct. Protoconch sculptured by very fine spiral and axial lirae, forming a reticulated pattern; spiral lirae fading on the transitional zone between protoconch and teleoconch. Axial lirae progressively transitioning into coarse ribs, with interspaces two to four times their width, comprising the primary sculpture of the teleoconch. Secondary sculpture of the teleoconch formed by crowded, wavy axial riblets, with interspaces approximately equal to their width, overlaying the primary sculpture. Umbilicus very narrow, deep. Peristome crescent-shaped; columellar lip reflected towards the shell axis, constricting the umbilicus in part. Palatal wall with four barriers and three traces, all recessed, extending approximately one-eighth whorl, with gradual anterior and posterior descension; all traces deeply recessed; first trace columellar in position, second and third traces infraperipheral and peripheral in position; barriers regularly spaced, similar in prominence, progressively transitioning from very shallowly recessed (first) to deeply recessed (fourth barrier); first barrier positioned at the confluence of basal and columellar lips; second and third barriers mid-basal in position; fourth barrier positioned at the confluence of basal and peripheral walls, as deeply recessed as traces. Parietal barriers five in number, not recessed within aperture, extending approximately one-quarter whorl, regularly spaced, with gradual anterior and somewhat abrupt posterior descension; barriers one to four similar in morphology; fifth barrier lower than the remainder. Other shell features that can be expressed numerically are shown in
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<tableCitation id="C6F6030EFF8AD456518C7D7EFAB4D3EE" box="[1207,1293,367,391]" captionStart="TABLE 2" captionStartId="8.[151,239,871,895]" captionTargetBox="[159,1432,1024,2009]" captionTargetPageId="8" captionText="TABLE 2. Dimensions (in mm) and ratios of type specimens. See Figure 2 for the placement of measurements. Tildes (~) and dashes (–) indicate approximate values and data missing due to poor preservation, respectively. Abbreviations: ah, aperture height; aw, aperture width; d, shell diameter; h, shell height; nr, number of ribs on body whorl; sp, spire protrusion; u, diameter of umbilicus; wn, number of whorls." httpUri="http://table.plazi.org/id/DF0B663DFF94D44855AC7F76FCA1D1B6" pageId="22" pageNumber="365" tableUuid="DF0B663DFF94D44855AC7F76FCA1D1B6">Table 2</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF0B663DFF8AD45655AC7A8EFC5CD49B" httpUri="https://zenodo.org/record/223292/files/figure.png" pageId="22" pageNumber="365" targetBox="[151,1435,445,1673]" targetPageId="22">
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<paragraph id="8BCB36B5FF8AD45655AC7A8EFC5CD49B" blockId="22.[151,1436,1695,1779]" pageId="22" pageNumber="365">
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<emphasis id="B900EAA7FF8AD45655AC7A8EFE8AD4DC" bold="true" box="[151,307,1695,1718]" pageId="22" pageNumber="365">FIGURE 12.</emphasis>
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<taxonomicName id="4C744D36FF8AD45654027A8EFDB3D4DC" box="[313,522,1695,1717]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="365" phylum="Mollusca" rank="species" species="boucheti" status="sp. nov.">
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<emphasis id="B900EAA7FF8AD45654027A8EFDB3D4DC" box="[313,522,1695,1717]" italics="true" pageId="22" pageNumber="365">Minidonta boucheti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23357DCFF8AD45657237AB1FDD4D4DC" box="[536,621,1696,1717]" pageId="22" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B900EAA7FF8AD456574E7AB1FD31D4DC" bold="true" box="[629,648,1696,1717]" pageId="22" pageNumber="365">A</emphasis>
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and
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<emphasis id="B900EAA7FF8AD45657F17A8EFD65D4DF" bold="true" box="[714,732,1695,1718]" pageId="22" pageNumber="365">C</emphasis>
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. Holotype (MNHN 25207), apical views;
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<emphasis id="B900EAA7FF8AD456518F7A8EFB7FD4DD" bold="true" box="[1204,1222,1695,1716]" pageId="22" pageNumber="365">A</emphasis>
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. General view;
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<emphasis id="B900EAA7FF8AD45650BF7AB1FA2CD4DC" bold="true" box="[1412,1429,1696,1717]" pageId="22" pageNumber="365">B</emphasis>
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. Sculpture of the protoconch and early teleoconch (specimen from Rr42);
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<emphasis id="B900EAA7FF8AD45656AE7AACFC1ED4BD" bold="true" box="[917,935,1725,1748]" pageId="22" pageNumber="365">C</emphasis>
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. Sculpture of the late teleoconch;
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<emphasis id="B900EAA7FF8AD45650367AAFFAA6D4BA" bold="true" box="[1293,1311,1726,1747]" pageId="22" pageNumber="365">D</emphasis>
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. Paratype 8 (MNHN 25208), detail of the peristome. Scale bars: A = 800
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<emphasis id="B900EAA7FF8AD45656287ACFFC99D49A" box="[787,800,1758,1779]" italics="true" pageId="22" pageNumber="365">µ</emphasis>
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m; B–D = 200
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<emphasis id="B900EAA7FF8AD45656817ACFFC70D49A" box="[954,969,1758,1779]" italics="true" pageId="22" pageNumber="365">µ</emphasis>
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m.
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</paragraph>
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</caption>
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<subSubSection id="C36E653EFF8AD45655FC7B31FAB0D5CD" pageId="22" pageNumber="365" type="discussion">
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<paragraph id="8BCB36B5FF8AD45655FC7B31FAB0D5CD" blockId="22.[151,1436,1823,1992]" pageId="22" pageNumber="365">
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<emphasis id="B900EAA7FF8AD45655FC7B31FE8DD551" bold="true" box="[199,308,1824,1848]" pageId="22" pageNumber="365">Remarks</emphasis>
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: In overall shape, sculpture and coloration this species closely resembles
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<taxonomicName id="4C744D36FF8AD45651547B30FB4DD551" box="[1135,1268,1824,1848]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="365" phylum="Mollusca" rank="species" species="boucheti">
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<emphasis id="B900EAA7FF8AD45651547B30FB4DD551" box="[1135,1268,1824,1848]" italics="true" pageId="22" pageNumber="365">M. boucheti</emphasis>
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</taxonomicName>
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, from which it is easily distinguished by its very narrow umbilicus. A relatively higher shell and a larger number of apertural barriers in
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<taxonomicName id="4C744D36FF8AD456542B7B78FEC6D5E9" box="[272,383,1896,1920]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="365" phylum="Mollusca" rank="species" species="bieleri">
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<emphasis id="B900EAA7FF8AD456542B7B78FEC6D5E9" box="[272,383,1896,1920]" italics="true" pageId="22" pageNumber="365">M. bieleri</emphasis>
|
||
</taxonomicName>
|
||
appear to provide the most useful additional criteria for separation of these seemingly closely related species. For a comparison with
|
||
<taxonomicName id="4C744D36FF8AD45657777B9DFD78D5CD" box="[588,705,1932,1956]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="365" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B900EAA7FF8AD45657777B9DFD78D5CD" box="[588,705,1932,1956]" italics="true" pageId="22" pageNumber="365">Minidonta</emphasis>
|
||
</taxonomicName>
|
||
from Raivavae, refer to remarks under
|
||
<taxonomicName id="4C744D36FF8AD45651477B9CFABBD5CD" box="[1148,1282,1932,1956]" class="Gastropoda" family="Endodontidae" genus="Minidonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="365" phylum="Mollusca" rank="species" species="boucheti">
|
||
<emphasis id="B900EAA7FF8AD45651477B9CFABBD5CD" box="[1148,1282,1932,1956]" italics="true" pageId="22" pageNumber="365">M. boucheti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653EFF8AD45655FC7BA1FBD6D5A1" box="[199,1135,1968,1992]" pageId="22" pageNumber="365" type="etymology">
|
||
<paragraph id="8BCB36B5FF8AD45655FC7BA1FBD6D5A1" blockId="22.[151,1436,1823,1992]" box="[199,1135,1968,1992]" pageId="22" pageNumber="365">
|
||
<emphasis id="B900EAA7FF8AD45655FC7BA1FEF1D5A1" bold="true" box="[199,328,1968,1992]" pageId="22" pageNumber="365">Etymology</emphasis>
|
||
: The species is dedicated to Rüdiger Bieler, distinguished malacologist.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |