280 lines
30 KiB
XML
280 lines
30 KiB
XML
<document id="1306477F73B2C2823A04808F3AE1E2C2" ID-DOI="10.5281/zenodo.190015" ID-GBIF-Dataset="a4c97e03-e184-4220-bbc2-ba1f955c580c" ID-ISSN="1175-5326" ID-Zenodo-Dep="190015" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461107024962" checkinUser="plazi" docAuthor="Kohlmann, Bert & Solis, Angel" docDate="2009" docId="03DD8799FFF48529FF66F92BFE0CFC1C" docLanguage="en" docName="zt02219p037.pdf" docOrigin="Zootaxa 2219" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ateuchus earthorum Kohlmann & Solis, sp. nov." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFE4FFE1FFF4852AFFF1FF8CFFE7FFA6" masterDocTitle="New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica" masterLastPageNumber="37" masterPageNumber="31" pageNumber="31" updateTime="1698593446454" updateUser="plazi">
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<mods:mods id="48842B2096186A6B58752081BBA6A99E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FE7A245C5195C3DCDEE8395FD84A246B">
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<mods:title id="A7B7825CA9663F22F53587BC320CD57E">New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica</mods:title>
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</mods:titleInfo>
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<mods:name id="536849C6049E9AFBFABE4271CA4B9872" type="personal">
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<mods:role id="27230F23660FB17003D082703A015031">
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<mods:roleTerm id="57C311C1427A616115CE10E501B95829">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="12D3B86837C719E8C474314D66611F1A">Kohlmann, Bert</mods:namePart>
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</mods:name>
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<mods:name id="95639B8FC530410D11AB6B22C08609FB" type="personal">
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<mods:role id="A07C5AF069AFAB7658D14D3A7CE53DD8">
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<mods:roleTerm id="969496933B0D9F109D68A3014E5737AA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D0AE85C71D6966D4CC859A2D3A84298B">Solis, Angel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="251AD096401B52285F81E5923BECECCD">text</mods:typeOfResource>
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<mods:relatedItem id="2CEC829F6A39168355BC002B2D24F8D0" type="host">
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<mods:titleInfo id="90A8287CEE35EE54AA77518F92603698">
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<mods:title id="87BF6B0BD1467C48430C1B619FA2B98D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C1CCC64DD0A96E7BD120C56A8B345DE0">
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<mods:date id="23BD4BA573DF19304B7A5471D52C0D79">2009</mods:date>
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<mods:detail id="1E4EB9B2486FC50E6FAC9606AE39F46E" type="volume">
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<mods:number id="E6819613B8E4391335BAD1D010EF3E6C">2219</mods:number>
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</mods:detail>
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<mods:extent id="84015FC6ADF91CAC3E5E9F0E72460E89" unit="page">
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<mods:start id="7C9562F34E3A2AA00BA4D35C271B436E">31</mods:start>
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<mods:end id="DF0D3159BD8CD1CF42F55DF6BDC841CE">37</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="935A44AAB352CCE64ED44903943B6F93">journal article</mods:classification>
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<mods:identifier id="20ACDA8E1B386E8075D1089DE41D4248" type="DOI">10.5281/zenodo.190015</mods:identifier>
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<mods:identifier id="8DD8AC17B9CADB6DE38DB5A157DEC391" type="GBIF-Dataset">a4c97e03-e184-4220-bbc2-ba1f955c580c</mods:identifier>
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<mods:identifier id="9B55BCE2FB6231042FC4D08FFCCF36CD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="598B202BA91DA1C3A6BEAEEC0F951B22" type="Zenodo-Dep">190015</mods:identifier>
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</mods:mods>
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<treatment id="03DD8799FFF48529FF66F92BFE0CFC1C" ID-DOI="http://doi.org/10.5281/zenodo.6216041" ID-GBIF-Taxon="119623535" ID-Zenodo-Dep="6216041" LSID="urn:lsid:plazi:treatment:03DD8799FFF48529FF66F92BFE0CFC1C" httpUri="http://treatment.plazi.org/id/03DD8799FFF48529FF66F92BFE0CFC1C" lastPageId="3" lastPageNumber="34" pageId="0" pageNumber="31">
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<subSubSection id="C36E6504FFF4852AFF66F92BFED2F942" pageId="0" pageNumber="31" type="nomenclature">
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||
<paragraph id="8BCB368FFFF4852AFF66F92BFD0CF967" blockId="0.[151,747,1703,1764]" box="[151,747,1703,1729]" pageId="0" pageNumber="31">
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<heading id="D08381E3FFF4852AFF66F92BFD0CF967" bold="true" box="[151,747,1703,1729]" fontSize="11" level="1" pageId="0" pageNumber="31" reason="1">
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<emphasis id="B900EA9DFFF4852AFF66F92BFD0CF967" bold="true" box="[151,747,1703,1729]" pageId="0" pageNumber="31">
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<taxonomicName id="4C744D0CFFF4852AFF66F92BFD66F967" ID-CoL="9X5PQ" authority="Kohlmann & Solis" authorityName="Kohlmann & Solis" box="[151,641,1703,1729]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="earthorum" status="sp. nov.">
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<emphasis id="B900EA9DFFF4852AFF66F92BFE76F967" bold="true" box="[151,401,1703,1729]" italics="true" pageId="0" pageNumber="31">Ateuchus earthorum</emphasis>
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Kohlmann & Solís
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23357E6FFF4852AFD7FF92BFD0CF967" box="[654,747,1703,1729]" pageId="0" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCB368FFFF4852AFF66F946FED2F942" blockId="0.[151,747,1703,1764]" box="[151,309,1738,1764]" pageId="0" pageNumber="31">
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||
(
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<figureCitation id="134F2A0AFFF4852AFF6EF946FEF7F942" box="[159,272,1738,1764]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,255,1552,1576]" captionStartId-1="2.[151,255,1401,1425]" captionText-0="FIGURE 1. Dorsal view of a male Ateuchus earthorum." captionText-1="FIGURE 2. Internal sac of the aedeagus of Ateuchus earthorum. Line = 1 mm" pageId="0" pageNumber="31">Figs. 1–2</figureCitation>
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,
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<figureCitation id="134F2A0AFFF4852AFEEDF946FECBF942" box="[284,300,1738,1764]" captionStart="FIGURE 4" captionStartId="5.[151,255,1308,1332]" captionTargetBox="[226,1360,194,1284]" captionTargetId="figure@5.[226,1360,194,1285]" captionTargetPageId="5" captionText="FIGURE 4. Known distribution of the new species in Costa Rica: A. earthorum (․) and C. darwini (˔)." httpUri="https://zenodo.org/record/190016/files/figure.png" pageId="0" pageNumber="31">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6504FFF4852AFF66F89BFCC9F855" pageId="0" pageNumber="31" type="diagnosis">
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<paragraph id="8BCB368FFFF4852AFF66F89BFCC9F855" blockId="0.[151,1436,1815,2035]" pageId="0" pageNumber="31">
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<emphasis id="B900EA9DFFF4852AFF66F89BFEFAF897" bold="true" box="[151,285,1815,1841]" pageId="0" pageNumber="31">Diagnosis:</emphasis>
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This species is distinguished from other Costa Rican species by the following combination of characters: head disc finely punctured with coarse punctures at anterior margin; pronotum finely punctured with coarse punctures at the base of the pronotal midline; anterior pronotal margin incomplete; eyes viewed from above 3 times longer than wide; head and pronotum with coppery red reflections; body oval; profemur coarsely punctured; proepimeron without punctures; elytra not shagreened; pygidium very convex; last abdominal segment broad; internal sac with three hooks.
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</paragraph>
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</subSubSection>
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<caption id="DF0B6607FFF5852BFF66F99CFD15F981" box="[151,754,1552,1576]" pageId="1" pageNumber="32">
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<paragraph id="8BCB368FFFF5852BFF66F99CFD15F981" blockId="1.[151,754,1552,1576]" box="[151,754,1552,1576]" pageId="1" pageNumber="32">
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<emphasis id="B900EA9DFFF5852BFF66F99CFEFEF98E" bold="true" box="[151,281,1552,1576]" pageId="1" pageNumber="32">FIGURE 1.</emphasis>
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Dorsal view of a male
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<taxonomicName id="4C744D0CFFF5852BFDE3F99CFD0DF981" box="[530,746,1552,1575]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="earthorum">
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<emphasis id="B900EA9DFFF5852BFDE3F99CFD0DF981" box="[530,746,1552,1575]" italics="true" pageId="1" pageNumber="32">Ateuchus earthorum</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<subSubSection id="C36E6504FFF58528FF34F9D9FEC0F8B6" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32" type="materials_examined">
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<paragraph id="8BCB368FFFF5852BFF34F9D9FA7BF930" blockId="1.[151,1437,1621,2034]" pageId="1" pageNumber="32">
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<emphasis id="B900EA9DFFF5852BFF34F9D9FE33F9C9" bold="true" box="[197,468,1621,1647]" pageId="1" pageNumber="32">
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<typeStatus id="54CF882DFFF5852BFF34F9D9FED0F9C9" box="[197,311,1621,1647]" pageId="1" pageNumber="32" type="holotype">Holotype</typeStatus>
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description:
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</emphasis>
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Male (
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<figureCitation id="134F2A0AFFF5852BFDD6F9D9FD7EF9C9" box="[551,665,1621,1647]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,255,1552,1576]" captionStartId-1="2.[151,255,1401,1425]" captionText-0="FIGURE 1. Dorsal view of a male Ateuchus earthorum." captionText-1="FIGURE 2. Internal sac of the aedeagus of Ateuchus earthorum. Line = 1 mm" pageId="1" pageNumber="32">Figs. 1–2</figureCitation>
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). Total length
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<quantity id="4C8C9B6AFFF5852BFCB0F9D9FC78F9C9" box="[833,927,1621,1647]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="32" unit="mm" value="6.7">6.7 mm</quantity>
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. Elytral width
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<quantity id="4C8C9B6AFFF5852BFBBAF9D9FB4EF9C9" box="[1099,1193,1621,1647]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="32" unit="mm" value="4.4">4.4 mm</quantity>
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. Body slightly ovoid and convex, dorsum dark brown, head and pronotum with strong cupreous red reflections; venter dark brown.
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</paragraph>
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<paragraph id="8BCB368FFFF5852BFF37F92FFDD2F8AC" blockId="1.[151,1437,1621,2034]" pageId="1" pageNumber="32">Clypeal margin coarsely punctate and broadly V-shaped, tooth on each side rounded, lateral margin arcuate, dorsal surface of head finely punctate and granulate, frons and vertex feebly tumid, eye dorsally small (ten times the interocular distance).</paragraph>
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<paragraph id="8BCB368FFFF5852BFF37F89BFDF4F8D8" blockId="1.[151,1437,1621,2034]" pageId="1" pageNumber="32">Pronotum finely punctate and granulate, moderately punctate at posterior end of midline, midline impressed only one-third pronotal length, anterior pronotal margin incomplete. Proepisternum finely wrinkled, proepimeron granular.</paragraph>
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<paragraph id="8BCB368FFFF5852BFF37F807FE6FF854" blockId="1.[151,1437,1621,2034]" pageId="1" pageNumber="32">Elytral surface smooth and shiny; striae slightly impressed, more strongly so anteriorly; striae feebly punctate, intervals slightly convex. Pygidium very convex, surface slightly granulate and minutely punctate, completely grooved.</paragraph>
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<caption id="DF0B6607FFF68528FF66FAF5FC3BFA36" box="[151,988,1401,1425]" pageId="2" pageNumber="33">
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||
<paragraph id="8BCB368FFFF68528FF66FAF5FC3BFA36" blockId="2.[151,988,1401,1425]" box="[151,988,1401,1425]" pageId="2" pageNumber="33">
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||
<emphasis id="B900EA9DFFF68528FF66FAF5FEFEFA37" bold="true" box="[151,281,1401,1425]" pageId="2" pageNumber="33">FIGURE 2.</emphasis>
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Internal sac of the aedeagus of
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||
<taxonomicName id="4C744D0CFFF68528FD9AFAF5FCA3FA36" box="[619,836,1401,1424]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="earthorum">
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||
<emphasis id="B900EA9DFFF68528FD9AFAF5FCA3FA36" box="[619,836,1401,1424]" italics="true" pageId="2" pageNumber="33">Ateuchus earthorum</emphasis>
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</taxonomicName>
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. Line = 1 mm
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</paragraph>
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</caption>
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<paragraph id="8BCB368FFFF68528FF37FA4DFC15F98E" blockId="2.[151,1437,1473,2002]" pageId="2" pageNumber="33">Protibia quadridentate, basal tooth small, protibial spur oval; apical one-half of profemur ventrally coarsely punctate, punctures extending along posterior margin to base of femur, punctate area black; mesofemur and metafemur short, thick, with minute punctures near apex.</paragraph>
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<paragraph id="8BCB368FFFF68528FF37F9B9FE1FF9D0" blockId="2.[151,1437,1473,2002]" pageId="2" pageNumber="33">
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Internal sac of the aedeagus (
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<figureCitation id="134F2A0AFFF68528FDD2F9B9FD8CF9E9" box="[547,619,1589,1615]" captionStart="FIGURE 2" captionStartId="2.[151,255,1401,1425]" captionText="FIGURE 2. Internal sac of the aedeagus of Ateuchus earthorum. Line = 1 mm" pageId="2" pageNumber="33">Fig. 2</figureCitation>
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) with three hooks, one small, two large; three apical lamellae; and a well-developed, spiny fascies.
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</paragraph>
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<paragraph id="8BCB368FFFF68528FF34F90EFEC0F8B6" blockId="2.[151,1437,1473,2002]" pageId="2" pageNumber="33">
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<emphasis id="B900EA9DFFF68528FF34F90EFEA6F93A" bold="true" box="[197,321,1666,1692]" pageId="2" pageNumber="33">Allotype:</emphasis>
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Female. Total length:
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<quantity id="4C8C9B6AFFF68528FD93F90EFD20F93A" box="[610,711,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="33" unit="mm" value="7.2">7.2 mm</quantity>
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. Elytral width:
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<quantity id="4C8C9B6AFFF68528FC7CF90EFC16F93A" box="[909,1009,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="33" unit="mm" value="4.9">4.9 mm</quantity>
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. Differs from the
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<typeStatus id="54CF882DFFF68528FB27F90EFAA3F93A" box="[1238,1348,1666,1692]" pageId="2" pageNumber="33" type="holotype">holotype</typeStatus>
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by the following characters: Clypeal margin anteriorly moderately V-shaped, anterior clypeal border moderately punctate, protibia with acute, slender spur slightly bent apically; last abdominal segment broader, pygidium less convex.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6504FFF68528FF34F891FDC6F8F8" pageId="2" pageNumber="33" type="description">
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<paragraph id="8BCB368FFFF68528FF34F891FDC6F8F8" blockId="2.[151,1437,1473,2002]" pageId="2" pageNumber="33">
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<emphasis id="B900EA9DFFF68528FF34F891FEACF891" bold="true" box="[197,331,1821,1847]" pageId="2" pageNumber="33">Variation:</emphasis>
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Total length:
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||
<quantity id="4C8C9B6AFFF68528FDF1F891FD7CF891" box="[512,667,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.2" metricValueMin="6.3" pageId="2" pageNumber="33" unit="mm" value="6.75" valueMax="7.2" valueMin="6.3">6.3–7.2 mm</quantity>
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||
. Elytral width:
|
||
<quantity id="4C8C9B6AFFF68528FCAEF891FC1DF891" box="[863,1018,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="2" pageNumber="33" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.1–4.9 mm</quantity>
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. The color intensity varies of the cupreous-red head and pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6504FFF68528FF34F8E6FA94F877" pageId="2" pageNumber="33" type="materials_examined">
|
||
<paragraph id="8BCB368FFFF68528FF34F8E6FA94F877" blockId="2.[151,1437,1473,2002]" pageId="2" pageNumber="33">
|
||
<emphasis id="B900EA9DFFF68528FF34F8E6FE58F822" bold="true" box="[197,447,1898,1924]" pageId="2" pageNumber="33">Material Examined</emphasis>
|
||
(
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||
<specimenCount id="9D72FD06FFF68528FE21F8E6FD92F822" box="[464,629,1898,1924]" pageId="2" pageNumber="33" type="generic">11 specimens</specimenCount>
|
||
):
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||
<emphasis id="B900EA9DFFF68528FD7EF8E6FCEBF822" bold="true" box="[655,780,1898,1924]" pageId="2" pageNumber="33">
|
||
<typeStatus id="54CF882DFFF68528FD7EF8E6FCE2F822" box="[655,773,1898,1924]" pageId="2" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
,
|
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</emphasis>
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||
male:
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<collectingCountry id="F363761FFFF68528FCAEF8E6FBF4F822" box="[863,1043,1898,1924]" name="Costa Rica" pageId="2" pageNumber="33">COSTA RICA</collectingCountry>
|
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: Est. Cacao,
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<quantity id="4C8C9B6AFFF68528FB41F8E6FB0AF822" box="[1200,1261,1898,1924]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="33" unit="km" value="2.0">2 km</quantity>
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||
SW del Cerro Cacao, Prov. Guanacaste,
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<quantity id="4C8C9B6AFFF68528FE3AF81DFDC1F80D" box="[459,550,1937,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="33" unit="m" value="1100.0">1100 m</quantity>
|
||
,
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||
<date id="FFCA104FFFF68528FDC3F81DFD0AF80D" box="[562,749,1937,1963]" pageId="2" pageNumber="33" value="1995-09-12" valueMax="1995-09-14" valueMin="1995-09-12">12–14SET1995</date>
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, C.
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<taxonomicName id="4C744D0CFFF68528FCEDF81DFC5EF80D" box="[796,953,1937,1963]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
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, caca de caballo. L_N _323100_375800, #6292.
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<emphasis id="B900EA9DFFF68528FF1EF83BFE87F877" bold="true" box="[239,352,1975,2001]" pageId="2" pageNumber="33">Allotype,</emphasis>
|
||
female:
|
||
<emphasis id="B900EA9DFFF68528FE37F834FDF2F877" box="[454,533,1976,2001]" italics="true" pageId="2" pageNumber="33">ibidem</emphasis>
|
||
.
|
||
<emphasis id="B900EA9DFFF68528FDD4F83BFD4EF877" bold="true" box="[549,681,1975,2001]" pageId="2" pageNumber="33">
|
||
<typeStatus id="54CF882DFFF68528FDD4F83BFD43F877" box="[549,676,1975,2001]" pageId="2" pageNumber="33" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B900EA9DFFF68528FD40F834FCEFF877" box="[689,776,1976,2001]" italics="true" pageId="2" pageNumber="33">ibidem,</emphasis>
|
||
<specimenCount id="9D72FD06FFF68528FCFEF834FC8BF874" box="[783,876,1976,2002]" pageId="2" pageNumber="33" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D72FD06FFF68528FC89F834FC0AF874" box="[888,1005,1976,2002]" pageId="2" pageNumber="33" type="female">3 females</specimenCount>
|
||
;
|
||
<emphasis id="B900EA9DFFF68528FC08F834FBAFF877" box="[1017,1096,1976,2001]" italics="true" pageId="2" pageNumber="33">ibidem</emphasis>
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, caca de mono,
|
||
<specimenCount id="9D72FD06FFF68528FAF6F834FA8AF874" box="[1287,1389,1976,2002]" pageId="2" pageNumber="33" type="female">1 female</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF34FF1BFD6FFDFD" pageId="3" pageNumber="34" type="discussion">
|
||
<paragraph id="8BCB368FFFF78529FF34FF1BFA74FDA8" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FF1BFEA4FF17" bold="true" box="[197,323,151,177]" pageId="3" pageNumber="34">Remarks:</emphasis>
|
||
This species will key to
|
||
<taxonomicName id="4C744D0CFFF78529FD86FF14FD3AFF17" box="[631,733,152,177]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ginae">
|
||
<emphasis id="B900EA9DFFF78529FD86FF14FD3AFF17" box="[631,733,152,177]" italics="true" pageId="3" pageNumber="34">A. ginae</emphasis>
|
||
</taxonomicName>
|
||
in Kohlmann’s (1997) key. These species are cryptic and cannot be separated on the grounds of external morphology; only the internal sac differences will distinguish them. In both species there are three hooks, two are long and similar and the third is long and like a simple bar in
|
||
<taxonomicName id="4C744D0CFFF78529FF49FE80FEF8FE83" box="[184,287,268,293]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ginae">
|
||
<emphasis id="B900EA9DFFF78529FF49FE80FEF8FE83" box="[184,287,268,293]" italics="true" pageId="3" pageNumber="34">A. ginae</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is short and spine-like in
|
||
<taxonomicName id="4C744D0CFFF78529FD07FE80FC7CFE83" box="[758,923,268,293]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="earthorum">
|
||
<emphasis id="B900EA9DFFF78529FD07FE80FC7CFE83" box="[758,923,268,293]" italics="true" pageId="3" pageNumber="34">A. earthorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134F2A0AFFF78529FC5CFE80FC1CFE80" box="[941,1019,268,294]" captionStart="FIGURE 2" captionStartId="2.[151,255,1401,1425]" captionText="FIGURE 2. Internal sac of the aedeagus of Ateuchus earthorum. Line = 1 mm" pageId="3" pageNumber="34">Fig. 2</figureCitation>
|
||
). These sac-hooks morphological differences are not only consistent (external morphology can be very variable and misleading) and geographically circumscribed to Cacao volcano, but also typical for separating
|
||
<taxonomicName id="4C744D0CFFF78529FBB5FED5FB57FED4" box="[1092,1200,345,370]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B900EA9DFFF78529FBB5FED5FB57FED4" box="[1092,1200,345,370]" italics="true" pageId="3" pageNumber="34">Ateuchus</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFE54B7EFFF78529FAECFED5FF33FE3C" author="Kohlmann" pageId="3" pageNumber="37" refString="Kohlmann, B. (1984) Biosistematica de las especies norteamericanas del genero Ateuchus (Coleoptera: Scarabaeidae: Scarabaeinae). Folia Entomologica Mexicana, 60, 3 - 81." type="journal article" year="1984">Kohlmann 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B7EFFF78529FF11FE0CFEFAFE3C" author="Kohlmann" box="[224,285,384,410]" pageId="3" pageNumber="37" refString="Kohlmann, B. (1997) The Costa Rican species of Ateuchus (Coleoptera: Scarabaeidae). Revista de Biologia Tropical, 44 (3) / 45 (1), 177 - 192." type="journal article" year="1997">1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B7EFFF78529FEDBFE0CFE8DFE3C" author="Kohlmann" box="[298,362,384,410]" pageId="3" pageNumber="37" refString="Kohlmann, B. (2000) New species and distribution records of Mesoamerican Ateuchus (Coleoptera: Scarabaeidae). Revista de Biologia Tropical, 48, 233 - 244." type="journal article" year="2000">2000</bibRefCitation>
|
||
). Moreover, the ecology and geology are also critical; the Guanacaste volcanoes are a wellknown species generating area for small-sized dung beetle taxa like
|
||
<taxonomicName id="4C744D0CFFF78529FC49FE2BFBC3FE66" box="[952,1060,423,448]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B900EA9DFFF78529FC49FE2BFBC3FE66" box="[952,1060,423,448]" italics="true" pageId="3" pageNumber="34">Ateuchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D0CFFF78529FBC2FE2BFB5AFE66" box="[1075,1213,423,448]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B900EA9DFFF78529FBC2FE2BFB5AFE66" box="[1075,1213,423,448]" italics="true" pageId="3" pageNumber="34">Canthidium</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C744D0CFFF78529FB0FFE2BFA7BFE66" box="[1278,1436,423,448]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B900EA9DFFF78529FB0FFE2BFA7BFE66" box="[1278,1436,423,448]" italics="true" pageId="3" pageNumber="34">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE54B7EFFF78529FF6EFE41FE80FE41" author="Kohlmann" box="[159,359,461,487]" pageId="3" pageNumber="37" refString="Kohlmann, B. (1997) The Costa Rican species of Ateuchus (Coleoptera: Scarabaeidae). Revista de Biologia Tropical, 44 (3) / 45 (1), 177 - 192." type="journal article" year="1997">Kohlmann 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B7EFFF78529FE84FE41FD47FE41" author="Kohlmann" box="[373,672,461,487]" pageId="3" pageNumber="37" refString="Kohlmann, B. & Solis, A. (2001) The genus Onthophagus (Coleoptera: Scarabaeidae) in Costa Rica. Giornale Italiano di Entomologia, 9, 159 - 261." type="journal article" year="2001">Kohlmann & Solís 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B7EFFF78529FD5CFE41FBFFFE41" author="Kohlmann" box="[685,1048,461,487]" pageId="3" pageNumber="37" refString="Kohlmann, B. & Wilkinson, J. (2007) The Tarcoles line: biogeographic effects of the Talamanca Range in lower Central America. Giornale Italiano di Entomologia, 12, 1 - 30." type="journal article" year="2007">Kohlmann & Wilkinson 2007</bibRefCitation>
|
||
, Kohlmann
|
||
<emphasis id="B900EA9DFFF78529FB5CFE41FB04FE40" box="[1197,1251,461,486]" italics="true" pageId="3" pageNumber="34">et al</emphasis>
|
||
. 2007,
|
||
<bibRefCitation id="EFE54B7EFFF78529FAB1FE41FEB1FDA8" author="Solis" pageId="3" pageNumber="37" refString="Solis, A. & Kohlmann, B. (2004) El genero Canthidium (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 52 (11), 1 - 73." type="journal article" year="2004">Solís & Kohlmann 2004</bibRefCitation>
|
||
) and for plants (
|
||
<taxonomicName id="4C744D0CFFF78529FDE7FE78FD9CFDA8" box="[534,635,500,526]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="34" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D0CFFF78529FD77FE78FCE4FDA8" box="[646,771,500,526]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="34" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C744D0CFFF78529FCB1FE78FC04FDA8" box="[832,995,500,526]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="3" pageNumber="34" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
|
||
) as well (Kohlmann
|
||
<emphasis id="B900EA9DFFF78529FB28FE78FAEBFDAB" box="[1241,1292,500,525]" italics="true" pageId="3" pageNumber="34">et al</emphasis>
|
||
. accepted).
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF34FD96FD6FFDFD" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FD96FEC8FD92" bold="true" box="[197,303,538,564]" pageId="3" pageNumber="34">Habitat:</emphasis>
|
||
The new species lives in mountain tropical forest at
|
||
<quantity id="4C8C9B6AFFF78529FC64FD96FC0BFD92" box="[917,1004,538,564]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="34" unit="m" value="1100.0">1100 m</quantity>
|
||
and has been collected in September in horse manure and from monkey’s dung.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF34FDEBFC35FCE5" pageId="3" pageNumber="34" type="description">
|
||
<paragraph id="8BCB368FFFF78529FF34FDEBFD5EFD0E" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FDEBFD6EFD27" bold="true" box="[197,649,615,641]" pageId="3" pageNumber="34">
|
||
Geographical distribution (
|
||
<figureCitation id="134F2A0AFFF78529FDD8FDEBFD90FD27" box="[553,631,615,641]" captionStart="FIGURE 4" captionStartId="5.[151,255,1308,1332]" captionTargetBox="[226,1360,194,1284]" captionTargetId="figure@5.[226,1360,194,1285]" captionTargetPageId="5" captionText="FIGURE 4. Known distribution of the new species in Costa Rica: A. earthorum (․) and C. darwini (˔)." httpUri="https://zenodo.org/record/190016/files/figure.png" pageId="3" pageNumber="34">Fig. 4</figureCitation>
|
||
):
|
||
</emphasis>
|
||
The new species is so far only known from the Pacific slope of Cacao volcano, in the province of Guanacaste.
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF34FD39FBD7FD50" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FD39FE04FD69" bold="true" box="[197,483,693,719]" pageId="3" pageNumber="34">Chorological affinities:</emphasis>
|
||
The new species is found at the same altitude, in the Guanacaste Cordillera, as its sister species,
|
||
<taxonomicName id="4C744D0CFFF78529FECEFD50FDC0FD50" authority="Kohlmann" authorityName="Kohlmann" box="[319,551,732,758]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ginae">
|
||
<emphasis id="B900EA9DFFF78529FECEFD50FE47FD53" box="[319,416,732,757]" italics="true" pageId="3" pageNumber="34">A. ginae</emphasis>
|
||
Kohlmann
|
||
</taxonomicName>
|
||
, in the Central Cordillera (
|
||
<bibRefCitation id="EFE54B7EFFF78529FCAEFD50FBC4FD50" author="Kohlmann" box="[863,1059,732,758]" pageId="3" pageNumber="37" refString="Kohlmann, B. (1997) The Costa Rican species of Ateuchus (Coleoptera: Scarabaeidae). Revista de Biologia Tropical, 44 (3) / 45 (1), 177 - 192." type="journal article" year="1997">Kohlmann 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF34FC8EFC35FCE5" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FC8EFDF4FCBA" bold="true" box="[197,531,770,796]" pageId="3" pageNumber="34">Taxonomic relationships:</emphasis>
|
||
<taxonomicName id="4C744D0CFFF78529FDEDFC8EFCFDFCBD" box="[540,794,770,795]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="earthorum">
|
||
<emphasis id="B900EA9DFFF78529FDEDFC8EFCFDFCBD" box="[540,794,770,795]" italics="true" pageId="3" pageNumber="34">Ateuchus earthorum</emphasis>
|
||
</taxonomicName>
|
||
is hypothesized to be the sister species to
|
||
<taxonomicName id="4C744D0CFFF78529FAC5FC8EFEF4FCE5" authority="Kohlmann" authorityName="Kohlmann" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ginae">
|
||
<emphasis id="B900EA9DFFF78529FAC5FC8EFA7AFCBD" box="[1332,1437,770,795]" italics="true" pageId="3" pageNumber="34">A. ginae</emphasis>
|
||
Kohlmann
|
||
</taxonomicName>
|
||
based on shared morphological characters discussed above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF34FCC3FE0CFC1C" pageId="3" pageNumber="34" type="etymology">
|
||
<paragraph id="8BCB368FFFF78529FF34FCC3FE0CFC1C" blockId="3.[151,1437,151,954]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FCC3FEBCFCCF" bold="true" box="[197,347,847,873]" pageId="3" pageNumber="34">Etymology:</emphasis>
|
||
The name is a Latinized noun in the genitive case. This species is dedicated to EARTH University in
|
||
<collectingCountry id="F363761FFFF78529FECFFCFAFE22FC36" box="[318,453,886,912]" name="Costa Rica" pageId="3" pageNumber="34">Costa Rica</collectingCountry>
|
||
, an institution committed to the sustainable development of the humid tropics, to celebrate its 20th anniversary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |