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<document id="AD1C39B9A881988CAA92F0FE9072201E" ID-DOI="10.11646/zootaxa.4743.4.1" ID-GBIF-Dataset="44ef21b3-550b-4f82-950c-3d7db7e04de0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3690502" ID-ZooBank="4F90419E-E06F-4990-84F7-3D0B1BA9AF0D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582874459487" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2020" docId="03DD878B5879CC6FD3824C21FCDEFC11" docLanguage="en" docName="zootaxa.4743.4.1.pdf" docOrigin="Zootaxa 4743 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maera sagamiensis Ariyama 2020, sp. nov." docType="treatment" docUuid="C19E1F68-574F-4B8D-8296-98A13F215795" docUuidSource="ZooBank" docVersion="7" lastPageNumber="460" masterDocId="FFE4FFF35871CC66D3154D6CFFD7FFDD" masterDocTitle="Species of the Maera - clade collected from Japan. Part 3: genera Maera Leach, 1814, Meximaera Barnard, 1969 and Orientomaera Ariyama, 2018 (addendum), with a key to Japanese species of the clade (Crustacea: Amphipoda: Maeridae)" masterLastPageNumber="479" masterPageNumber="451" pageNumber="459" updateTime="1698723202199" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A1B48F344673F7A2022B5AC7CC1800E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0400E731208943411D10E930ED9B6AA5">
<mods:title id="E6E381C235D27922055F45590AEF5265">Species of the Maera - clade collected from Japan. Part 3: genera Maera Leach, 1814, Meximaera Barnard, 1969 and Orientomaera Ariyama, 2018 (addendum), with a key to Japanese species of the clade (Crustacea: Amphipoda: Maeridae)</mods:title>
</mods:titleInfo>
<mods:name id="7C671A59A97618C15BAB7E592984B615" type="personal">
<mods:role id="08C35F3CA56595A576836A4A342FBEDA">
<mods:roleTerm id="B97011F82724A7DA0AC9099E75623A80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09A62F1157476A88F2EDCB1283AB292A">Ariyama, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="61085D3DE13884E6A25154C077ECEC80">text</mods:typeOfResource>
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<mods:title id="4921491696C271FF8CB19EFCF665B606">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9DE2BDBE26BA2594C85A44F2B8A5E85F">
<mods:date id="A469DDB01DF8124FC9D9FA8973DC5F17">2020</mods:date>
<mods:detail id="673DEBCC0D12653FD852DCE128B87645" type="pubDate">
<mods:number id="1141DC5FB776541A2F1759E894DA03C2">2020-02-27</mods:number>
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<mods:number id="DF87B2D38AF002A8C07F2B86CC448BCA">4743</mods:number>
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<mods:classification id="3A49F85EC8D0CDD213EB292FC8BCCB82">journal article</mods:classification>
<mods:identifier id="23B5314FAD53FD9058C98F5702A4D972" type="DOI">10.11646/zootaxa.4743.4.1</mods:identifier>
<mods:identifier id="62A7CA9DA45F8DC382616B8E30502127" type="GBIF-Dataset">44ef21b3-550b-4f82-950c-3d7db7e04de0</mods:identifier>
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<treatment id="03DD878B5879CC6FD3824C21FCDEFC11" ID-DOI="http://doi.org/10.5281/zenodo.3691396" ID-GBIF-Taxon="162361441" ID-Zenodo-Dep="3691396" LSID="urn:lsid:zoobank.org:act:C19E1F68-574F-4B8D-8296-98A13F215795" httpUri="http://treatment.plazi.org/id/03DD878B5879CC6FD3824C21FCDEFC11" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
<subSubSection id="C36E65165879CC6ED3824C21FD70FE56" pageId="8" pageNumber="459" type="nomenclature">
<paragraph id="8BCB369D5879CC6ED3824C21FE31FEBA" blockId="8.[151,679,333,432]" box="[151,486,333,359]" pageId="8" pageNumber="459">
<heading id="D08381F15879CC6ED3824C21FE31FEBA" bold="true" box="[151,486,333,359]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
<emphasis id="B900EA8F5879CC6ED3824C21FE31FEBA" bold="true" box="[151,486,333,359]" pageId="8" pageNumber="459">
<taxonomicName id="4C744D1E5879CC6ED3824C21FE56FEBA" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[151,385,333,359]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sagamiensis" status="sp. nov.">
<emphasis id="B900EA8F5879CC6ED3824C21FE56FEBA" bold="true" box="[151,385,333,359]" italics="true" pageId="8" pageNumber="459">Maera sagamiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357F45879CC6ED29D4C21FE31FEBA" box="[392,486,333,359]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB369D5879CC6ED3824C1DFD70FE56" blockId="8.[151,679,333,432]" box="[151,679,368,395]" pageId="8" pageNumber="459">[Japanese name: Sagami-sunnariyokoebi, new]</paragraph>
</subSubSection>
<subSubSection id="C36E65165879CC6ED3824CF9FECEFE6D" box="[151,281,405,432]" pageId="8" pageNumber="459" type="description">
<paragraph id="8BCB369D5879CC6ED3824CF9FECEFE6D" blockId="8.[151,679,333,432]" box="[151,281,405,432]" pageId="8" pageNumber="459">
(
<figureCitation id="134F2A185879CC6ED38A4CF9FEC6FE6D" box="[159,273,405,432]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="9.[151,250,1980,2005]" captionStartId-1="10.[151,250,1998,2023]" captionStartId-2="11.[151,250,1993,2018]" captionStartId-3="12.[151,250,1467,1492]" captionStartId-4="13.[151,250,1541,1566]" captionTargetBox-0="[157,1429,1000,1949]" captionTargetBox-1="[176,1414,192,1961]" captionTargetBox-2="[186,1401,189,1953]" captionTargetBox-3="[157,1430,190,1438]" captionTargetBox-4="[151,1435,417,1515]" captionTargetId-0="figure@9.[151,1436,993,1955]" captionTargetId-1="figure@10.[163,1424,181,1973]" captionTargetId-2="figure@11.[174,1413,181,1967]" captionTargetId-3="figure@12.[151,1436,181,1442]" captionTargetId-4="figure@13.[151,1435,417,1515]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 6. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Habitus." captionText-1="FIGURE 7. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.15 mm." captionText-2="FIGURE 8. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.1 mm." captionText-3="FIGURE 9. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.2 mm." captionText-4="FIGURE 10. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Live specimen, photograph by H. Kohtsuka." figureDoi-0="http://doi.org/10.5281/zenodo.3690514" figureDoi-1="http://doi.org/10.5281/zenodo.3690516" figureDoi-2="http://doi.org/10.5281/zenodo.3690518" figureDoi-3="http://doi.org/10.5281/zenodo.3690520" figureDoi-4="http://doi.org/10.5281/zenodo.3690522" httpUri-0="https://zenodo.org/record/3690514/files/figure.png" httpUri-1="https://zenodo.org/record/3690516/files/figure.png" httpUri-2="https://zenodo.org/record/3690518/files/figure.png" httpUri-3="https://zenodo.org/record/3690520/files/figure.png" httpUri-4="https://zenodo.org/record/3690522/files/figure.png" pageId="8" pageNumber="459">Figs 610</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E65165879CC6ED3824CB1FB0CFDBE" pageId="8" pageNumber="459" type="materials_examined">
<paragraph id="8BCB369D5879CC6ED3824CB1FEFBFDE2" blockId="8.[151,1437,476,1979]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3824CB1FE90FE2A" bold="true" box="[151,327,476,503]" pageId="8" pageNumber="459">Type material.</emphasis>
<materialsCitation id="3B1C3CC05879CC6ED2454CB1FEFBFDE2" ID-GBIF-Occurrence="2572153715" collectingDate="2015-02-12" collectorName="H. Kohtsuka." elevation="204" latitude="35.141945" location="Jogashima Island in Miura City" longLatPrecision="19" longitude="139.54112" municipality="Sagami Bay" pageId="8" pageNumber="459" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
<typeStatus id="54CF883F5879CC6ED2454CB1FE6AFE2A" box="[336,445,477,503]" pageId="8" pageNumber="459" type="holotype">Holotype</typeStatus>
: male,
<quantity id="4C8C9B785879CC6ED11A4CB1FDA1FE2A" box="[527,630,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="459" unit="mm" value="10.1">10.1 mm</quantity>
(OMNH-Ar-10625),
<collectingMunicipality id="6BAFACE75879CC6ED0794CB1FC20FE2A" box="[876,1015,477,503]" pageId="8" pageNumber="459">Sagami Bay</collectingMunicipality>
, off
<location id="8EAB60465879CC6ED73B4CB1FA4FFE2A" LSID="urn:lsid:plazi:treatment:03DD878B5879CC6FD3824C21FCDEFC11:8EAB60465879CC6ED73B4CB1FA4FFE2A" box="[1070,1432,477,503]" latitude="35.141945" longLatPrecision="19" longitude="139.54112" municipality="Sagami Bay" name="Jogashima Island in Miura City" pageId="8" pageNumber="459" stateProvince="Kanagawa">Jôgashima Island in Miura City</location>
,
<collectingRegion id="49B0F87F5879CC6ED3824F6DFE5EFDC6" box="[151,393,513,539]" country="Japan" name="Kanagawa" pageId="8" pageNumber="459">Kanagawa Prefecture</collectingRegion>
,
<geoCoordinate id="EE40505A5879CC6ED2864F6CFDCBFDC6" box="[403,540,512,539]" degrees="35" direction="north" minutes="08" orientation="latitude" pageId="8" pageNumber="459" precision="15" seconds="31" value="35.141945">35°0831”N</geoCoordinate>
<geoCoordinate id="EE40505A5879CC6ED1374F6DFD61FDC6" box="[546,694,513,539]" degrees="139" direction="east" minutes="32" orientation="longitude" pageId="8" pageNumber="459" precision="15" seconds="28" value="139.54112">139°3228”E</geoCoordinate>
(
<figureCitation id="134F2A185879CC6ED1D14F6DFCCBFDC6" box="[708,796,513,539]" captionStart="FIGURE 1" captionStartId="2.[151,250,1627,1652]" captionTargetBox="[187,1400,182,1601]" captionTargetId="figure@2.[186,1400,181,1601]" captionTargetPageId="2" captionText="FIGURE 1. Map showing collecting sites of the examined specimens. A, Sagami Bay, off Jōgashima Island in Miura City, Kanagawa Prefecture; B, Tagurazaki coast, Wakayama City in Wakayama Prefecture; C, Sea of Japan, north of Masuda City in Yamaguchi Prefecture." figureDoi="http://doi.org/10.5281/zenodo.3690504" httpUri="https://zenodo.org/record/3690504/files/figure.png" pageId="8" pageNumber="459">Fig. 1A</figureCitation>
),
<elevation id="0059D1AE5879CC6ED0254F6DFC23FDC6" box="[816,1012,513,539]" metricMagnitude="2" metricUnit="m" metricValue="2.035" metricValueMax="2.7399999999999998" metricValueMin="1.33" pageId="8" pageNumber="459" unit="m" value="203.5" valueMax="274.0" valueMin="133.0">
<quantity id="4C8C9B785879CC6ED0254F6DFC79FDC6" box="[816,942,513,539]" metricMagnitude="2" metricUnit="m" metricValue="2.035" metricValueMax="2.7399999999999998" metricValueMin="1.33" pageId="8" pageNumber="459" unit="m" value="203.5" valueMax="274.0" valueMin="133.0">133274 m</quantity>
depth
</elevation>
, using dredge,
<date id="FFCA105D5879CC6ED78F4F6DFAB6FDC6" box="[1178,1377,512,539]" pageId="8" pageNumber="459" value="2015-02-12">
<collectingDate id="EF8EE9B55879CC6ED78F4F6DFAB6FDC6" box="[1178,1377,512,539]" pageId="8" pageNumber="459" value="2015-02-12">12 February 2015</collectingDate>
</date>
, coll.
<collectorName id="2681534B5879CC6ED3824F49FEFBFDE2" box="[151,300,549,575]" pageId="8" pageNumber="459">H. Kohtsuka.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BCB369D5879CC6ED3D24F25FB0CFDBE" blockId="8.[151,1437,476,1979]" box="[199,1243,584,611]" pageId="8" pageNumber="459">
<materialsCitation id="3B1C3CC05879CC6ED3D24F25FB01FDBE" ID-GBIF-Occurrence="2572153718" box="[199,1238,584,611]" country="Japan" location="Sagami Bay" municipality="Jogashima Island in Miura City" pageId="8" pageNumber="459" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
<emphasis id="B900EA8F5879CC6ED3D24F25FEB1FDBE" bold="true" box="[199,358,584,611]" pageId="8" pageNumber="459">
<typeStatus id="54CF883F5879CC6ED3D24F25FED5FDBE" box="[199,258,585,611]" pageId="8" pageNumber="459">Type</typeStatus>
locality.
</emphasis>
<location id="8EAB60465879CC6ED2794F25FE22FDBE" LSID="urn:lsid:plazi:treatment:03DD878B5879CC6FD3824C21FCDEFC11:8EAB60465879CC6ED2794F25FE22FDBE" box="[364,501,585,611]" country="Japan" municipality="Jogashima Island in Miura City" name="Sagami Bay" pageId="8" pageNumber="459" stateProvince="Kanagawa">Sagami Bay</location>
, off
<collectingMunicipality id="6BAFACE75879CC6ED13D4F25FC5DFDBE" box="[552,906,585,611]" pageId="8" pageNumber="459">Jōgashima Island in Miura City</collectingMunicipality>
,
<collectingRegion id="49B0F87F5879CC6ED0804F25FB5FFDBE" box="[917,1160,585,611]" country="Japan" name="Kanagawa" pageId="8" pageNumber="459">Kanagawa Prefecture</collectingRegion>
,
<collectingCountry id="F363760D5879CC6ED7864F25FB01FDBE" box="[1171,1238,585,611]" name="Japan" pageId="8" pageNumber="459">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165879CC6ED3D24F01FD49FD5A" box="[199,670,621,647]" pageId="8" pageNumber="459" type="etymology">
<paragraph id="8BCB369D5879CC6ED3D24F01FD49FD5A" blockId="8.[151,1437,476,1979]" box="[199,670,621,647]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D24F01FE99FD5A" bold="true" box="[199,334,621,647]" pageId="8" pageNumber="459">Etymology.</emphasis>
Referring to the
<typeStatus id="54CF883F5879CC6ED1184F01FDEAFD5A" box="[525,573,621,647]" pageId="8" pageNumber="459">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165879CC6ED3D24FFCFC83FC82" pageId="8" pageNumber="459" type="diagnosis">
<paragraph id="8BCB369D5879CC6ED3D24FFCFC83FC82" blockId="8.[151,1437,476,1979]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D24FFCFE95FD76" bold="true" box="[199,322,656,683]" pageId="8" pageNumber="459">Diagnosis.</emphasis>
<emphasis id="B900EA8F5879CC6ED25D4FFEFEAAFD76" box="[328,381,658,683]" italics="true" pageId="8" pageNumber="459">Eyes</emphasis>
present.
<emphasis id="B900EA8F5879CC6ED2F74FFDFD9AFD76" box="[482,589,657,683]" italics="true" pageId="8" pageNumber="459">Mandible</emphasis>
, palp article 1 with acute distal tooth.
<emphasis id="B900EA8F5879CC6ED0EC4FFDFB5BFD76" box="[1017,1164,657,683]" italics="true" pageId="8" pageNumber="459">Gnathopod 1</emphasis>
, coxa anteroventral corner acutely produced, posteroventral margin with 3 notches.
<emphasis id="B900EA8F5879CC6ED02F4FD9FBF2FD12" box="[826,1061,693,719]" italics="true" pageId="8" pageNumber="459">
Gnathopod
<quantity id="4C8C9B785879CC6ED0AA4FD9FC3EFD12" box="[959,1001,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="459" unit="in" value="2.0">2 in</quantity>
male
</emphasis>
, coxa posteroventral margin with 3 notches, palm strongly oblique, defined by acute tooth.
<emphasis id="B900EA8F5879CC6ED0304FB5FC4BFD2E" box="[805,924,729,755]" italics="true" pageId="8" pageNumber="459">Pereopods</emphasis>
dactyli simple; coxa 3 posteroventral margin bearing 2 notches posteroventrally, bases of pereopods 57 rectangular, posterodistal corners square.
<emphasis id="B900EA8F5879CC6ED7EA4F91FA81FCCA" box="[1279,1366,765,791]" italics="true" pageId="8" pageNumber="459">Pleonal</emphasis>
e
<emphasis id="B900EA8F5879CC6ED67C4F92FF39FCE6" italics="true" pageId="8" pageNumber="459">pimera 13</emphasis>
acutely projected posteroventrally.
<emphasis id="B900EA8F5879CC6ED1964E4DFD38FCE6" box="[643,751,801,827]" italics="true" pageId="8" pageNumber="459">Uropod 3</emphasis>
, outer ramus longer than inner ramus, rami 2.22.5 times as long as peduncle.
<emphasis id="B900EA8F5879CC6ED2744E29FE7EFC82" box="[353,425,837,863]" italics="true" pageId="8" pageNumber="459">Telson</emphasis>
longer than wide, distal ends pointed.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165879CC6FD3D24E04FE57FD2E" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="description">
<paragraph id="8BCB369D5879CC6ED3D24E04FC04FC5E" blockId="8.[151,1437,476,1979]" box="[199,979,872,899]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D24E04FE8EFC5E" bold="true" box="[199,345,872,899]" pageId="8" pageNumber="459">Description.</emphasis>
Based on
<typeStatus id="54CF883F5879CC6ED2DB4E05FDE4FC5E" box="[462,563,873,899]" pageId="8" pageNumber="459" type="holotype">holotype</typeStatus>
, male,
<quantity id="4C8C9B785879CC6ED1974E05FD31FC5E" box="[642,742,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="459" unit="mm" value="10.1">10.1 mm</quantity>
(OMNH-Ar-10625).
</paragraph>
<paragraph id="8BCB369D5879CC6ED3D24EE1FEB6FA42" blockId="8.[151,1437,476,1979]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D24EE1FED0FC7A" bold="true" box="[199,263,909,935]" pageId="8" pageNumber="459">Head</emphasis>
(
<figureCitation id="134F2A185879CC6ED2024EE1FEA8FC75" box="[279,383,909,936]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="9.[151,250,1980,2005]" captionStartId-1="10.[151,250,1998,2023]" captionTargetBox-0="[157,1429,1000,1949]" captionTargetBox-1="[176,1414,192,1961]" captionTargetId-0="figure@9.[151,1436,993,1955]" captionTargetId-1="figure@10.[163,1424,181,1973]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 6. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Habitus." captionText-1="FIGURE 7. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.15 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3690514" figureDoi-1="http://doi.org/10.5281/zenodo.3690516" httpUri-0="https://zenodo.org/record/3690514/files/figure.png" httpUri-1="https://zenodo.org/record/3690516/files/figure.png" pageId="8" pageNumber="459">Figs 67</figureCitation>
).
<emphasis id="B900EA8F5879CC6ED2804EE2FE1DFC7A" box="[405,458,910,935]" italics="true" pageId="8" pageNumber="459">Eyes</emphasis>
oval, about 0.2 times length of head.
<emphasis id="B900EA8F5879CC6ED0684EE1FC26FC7A" box="[893,1009,909,935]" italics="true" pageId="8" pageNumber="459">Antenna 1</emphasis>
poorly setose; peduncle with ratio of lengths of articles 13 1:1.15:0.3, article 1 with 4 small posteromedial robust setae; accessory flagellum with 7 articles, terminal article minute; primary flagellum damaged.
<emphasis id="B900EA8F5879CC6ED05F4EB9FC6BFC32" box="[842,956,981,1007]" italics="true" pageId="8" pageNumber="459">Antenna 2</emphasis>
about 0.35 times length of body; peduncle weakly setose posteriorly, with ratio of lengths of articles 35 1:3.7:2.5; flagellum setose, with 9 articles, terminal article minute.
<emphasis id="B900EA8F5879CC6ED2564972FE66FBEA" box="[323,433,1053,1079]" italics="true" pageId="8" pageNumber="459">Upper lip</emphasis>
, ventral margin rounded, weakly setose.
<emphasis id="B900EA8F5879CC6ED0924971FC25FBEA" box="[903,1010,1053,1079]" italics="true" pageId="8" pageNumber="459">Mandible</emphasis>
, incisor bearing 5 and 4 cusps in left and right, respectively, each lacinia mobilis 4-dentate, with 9 accessory setae; palp article 1 with acute distal tooth, articles 1
<emphasis id="B900EA8F5879CC6ED215490AFED9FBA2" box="[256,270,1126,1151]" italics="true" pageId="8" pageNumber="459"></emphasis>
3 length ratio (excluding distal tooth of article 1) 1:2.5:1.5, articles 2
<emphasis id="B900EA8F5879CC6ED724490AFBE8FBA2" box="[1073,1087,1126,1151]" italics="true" pageId="8" pageNumber="459"></emphasis>
3 setose.
<emphasis id="B900EA8F5879CC6ED7B8490AFACAFBA2" box="[1197,1309,1125,1151]" italics="true" pageId="8" pageNumber="459">Lower lip</emphasis>
with inner lobes, distal margins and ventral surfaces of both lobes weakly setose, mandibular processes small.
<emphasis id="B900EA8F5879CC6ED7FF49E5FA82FB7E" box="[1258,1365,1161,1187]" italics="true" pageId="8" pageNumber="459">Maxilla 1</emphasis>
, inner plate with 3 long and 1 short apical setae, medial and lateral margins and ventral surface bearing feeble setae, outer plate with 10 apical robust setae and medial and ventral feeble setae, palp article 1 with several laterodistal setae, article 2 bearing many apical setae.
<emphasis id="B900EA8F5879CC6ED13A4999FD4BFAD2" box="[559,668,1269,1295]" italics="true" pageId="8" pageNumber="459">Maxilla 2</emphasis>
, outer plate larger than inner plate; both plates bearing many apical setae, medial margin of inner plate with 1 normal and many feeble setae.
<emphasis id="B900EA8F5879CC6ED0CC4875FB85FAEE" box="[985,1106,1305,1331]" italics="true" pageId="8" pageNumber="459">Maxilliped</emphasis>
, inner plate truncate, distally setose, distomedial corner with 4 robust setae ventrally; outer plate with 23 long-to-short robust setae on distomedial margin; palp 4-articulate, article 2 setose medioventrally, article 3 with many distolateral setae, article 4 with large apical robust seta.
</paragraph>
<paragraph id="8BCB369D5879CC6ED3D248C5FC34F93E" blockId="8.[151,1437,476,1979]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D248C5FECDFA1E" bold="true" box="[199,282,1449,1475]" pageId="8" pageNumber="459">Pereon</emphasis>
(
<figureCitation id="134F2A185879CC6ED23C48C5FEBCFA1E" box="[297,363,1449,1475]" captionStart="FIGURE 8" captionStartId="11.[151,250,1993,2018]" captionTargetBox="[186,1401,189,1953]" captionTargetId="figure@11.[174,1413,181,1967]" captionTargetPageId="11" captionText="FIGURE 8. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3690518" httpUri="https://zenodo.org/record/3690518/files/figure.png" pageId="8" pageNumber="459">Fig. 8</figureCitation>
).
<emphasis id="B900EA8F5879CC6ED29448C5FDC4FA1E" box="[385,531,1449,1475]" italics="true" pageId="8" pageNumber="459">Gnathopod 1</emphasis>
, coxa anteroventral corner acutely produced, ventral margin with 3 small notches posteriorly; basis setose on posterior margin; carpus swollen posterodistally, posterior margin and medial surface heavily setose; propodus about 90% as long as carpus, anterior and posterior margins setose, palmar margin bearing many small robust setae medially; dactylus slender, almost straight, anterior margin with single seta.
<emphasis id="B900EA8F5879CC6ED60B4B79FF72F98E" italics="true" pageId="8" pageNumber="459">Gnathopod 2</emphasis>
, coxa subquadrate, posteroventral margin with 3 small notches; basis relatively narrow, about 2.9 times as long as width, posterior margin with several setae; merus with acute posterodistal spine; carpus swollen posterodistally, posterior margin setose; propodus slender, length about 1.95 times width, anteromedial surface and posterior margin setose; palm strongly oblique, defined by acute tooth, palmar margin with 5 small teeth, bearing 9 lateral and 13 medial robust setae; dactylus curved distally, anterior margin with 10 setae.
</paragraph>
<paragraph id="8BCB369D5879CC6ED3D24B81FAE9F866" blockId="8.[151,1437,476,1979]" pageId="8" pageNumber="459">
<emphasis id="B900EA8F5879CC6ED3D24B81FEA7F8DA" box="[199,368,1773,1799]" italics="true" pageId="8" pageNumber="459">Pereopods 34</emphasis>
, shapes subequal; coxae trapezoidal, coxa 3 with 2 small notches posteroventrally, gills large, subequal to bases lengths; bases rectilinear, anterior and posterior margins each with a few long setae proximally; dactylus of pereopod 4 slender, simple.
<emphasis id="B900EA8F5879CC6ED17C4A59FCC3F892" box="[617,788,1845,1871]" italics="true" pageId="8" pageNumber="459">Pereopods 57</emphasis>
, shapes subsimilar, pereopods 5, 7 about 1.2, 1.6 times length of pereopod 4, respectively; coxae 67, anteroventral margins each with single robust seta; bases rectangular, lengths about 2.9, 2.6, 2.2 times widths respectively, posteroproximal corners roundly projected, anterior margins with many short robust setae, posterior margins serrate, posterodistal corners square; dactyli short, simple.
</paragraph>
<paragraph id="8BCB369D5878CC6FD3D24DF5FE20FD76" blockId="9.[151,1437,153,972]" pageId="9" pageNumber="460">
<emphasis id="B900EA8F5878CC6FD3D24DF5FEDDFF6E" bold="true" box="[199,266,153,179]" pageId="9" pageNumber="460">Pleon</emphasis>
(
<figureCitation id="134F2A185878CC6FD20E4DF5FEB7FF6E" box="[283,352,153,179]" captionStart="FIGURE 9" captionStartId="12.[151,250,1467,1492]" captionTargetBox="[157,1430,190,1438]" captionTargetId="figure@12.[151,1436,181,1442]" captionTargetPageId="12" captionText="FIGURE 9. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3690520" httpUri="https://zenodo.org/record/3690520/files/figure.png" pageId="9" pageNumber="460">Fig. 9</figureCitation>
).
<emphasis id="B900EA8F5878CC6FD26D4DF6FDDCFF6E" box="[376,523,153,179]" italics="true" pageId="9" pageNumber="460">Epimera 13</emphasis>
acutely projected posteroventrally, each with lateral ridge, ventral margins with 3, 2, 5 (1 lost) robust setae respectively.
<emphasis id="B900EA8F5878CC6FD1714DD1FD29FF0A" box="[612,766,189,215]" italics="true" pageId="9" pageNumber="460">Pleopods 13</emphasis>
, pleopod 3 shortest; peduncles with 2, 2, 4 coupling hooks respectively; both rami each with 79 articles.
<emphasis id="B900EA8F5878CC6FD1B04D8DFCC7FF26" box="[677,784,225,251]" italics="true" pageId="9" pageNumber="460">Uropod 1</emphasis>
, peduncle with 15 dorsolateral and 11 dorsomedial robust setae, bearing large distolateral robust seta; outer ramus subequal length to inner ramus, about 0.75 times as long as peduncle, with 4 dorsolateral, 4 dorsomedial and 4 terminal robust setae; inner ramus with 9 dorsomedial and 4 terminal robust setae.
<emphasis id="B900EA8F5878CC6FD2844C21FE2AFEBA" box="[401,509,333,359]" italics="true" pageId="9" pageNumber="460">Uropod 2</emphasis>
about 0.6 times length of uropod 1; peduncle laterodistal corner projected, dorsolateral margin, dorsomedial margin and distomedial corner with 11, 6 and 3 robust setae, respectively; outer ramus about 0.85 times length of inner ramus, about 95% length of peduncle, with 8 dorsomedial and 3? terminal robust setae; inner ramus with 4 lateral, 9 dorsomedial and 5 terminal robust setae.
<emphasis id="B900EA8F5878CC6FD0FB4CD5FB8EFE0E" box="[1006,1113,441,467]" italics="true" pageId="9" pageNumber="460">Uropod 3</emphasis>
long, about 175% as long as uropod 2; peduncle, lateral, medial and ventral margins with 3, 1 and 2 robust setae, respectively, mediodistal corner bearing 4 robust setae; both rami rectilinear, outer ramus about 115% length of inner ramus, about 2.5 times length of peduncle, with vestigial second article; lateral and medial margins of both rami lined with robust setae, distal margins bearing several setae, longest seta about 0.15 times length of outer ramus.
<emphasis id="B900EA8F5878CC6FD7574F25FB5DFDBE" box="[1090,1162,585,611]" italics="true" pageId="9" pageNumber="460">Telson</emphasis>
about 1.25 times longer than wide, each lateral margin with 4 small robust setae, tips acutely projected, dorsal surfaces of each lobe with a robust seta and 3 sensory setae.
</paragraph>
<paragraph id="8BCB369D5878CC6FD3D24FD8FE57FD2E" blockId="9.[151,1437,153,972]" pageId="9" pageNumber="460">
<emphasis id="B900EA8F5878CC6FD3D24FD8FE46FD12" bold="true" box="[199,401,692,719]" pageId="9" pageNumber="460">Coloration in life</emphasis>
(
<figureCitation id="134F2A185878CC6FD2B54FD9FE26FD12" box="[416,497,693,719]" captionStart="FIGURE 10" captionStartId="13.[151,250,1541,1566]" captionTargetBox="[151,1435,417,1515]" captionTargetId="figure@13.[151,1435,417,1515]" captionTargetPageId="13" captionText="FIGURE 10. Maera sagamiensis sp. nov. Holotype, male, 10.1 mm (OMNH-Ar-10625). Live specimen, photograph by H. Kohtsuka." figureDoi="http://doi.org/10.5281/zenodo.3690522" httpUri="https://zenodo.org/record/3690522/files/figure.png" pageId="9" pageNumber="460">Fig. 10</figureCitation>
). Eyes black and white, mouth parts orange, other parts faint orange; yellow internal organs seen through.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165878CC6FD3D24F91FC3FFC75" pageId="9" pageNumber="460" type="discussion">
<paragraph id="8BCB369D5878CC6FD3D24F91FAEBFC5E" blockId="9.[151,1437,153,972]" pageId="9" pageNumber="460">
<emphasis id="B900EA8F5878CC6FD3D24F91FEECFCCA" bold="true" box="[199,315,765,791]" pageId="9" pageNumber="460">Remarks.</emphasis>
The presence of small notches on the coxae 13 is a distinct character in the genus, although Japanese specimens of
<taxonomicName id="4C744D1E5878CC6FD2274E4EFE1EFCE6" baseAuthorityName="Bruzelius" baseAuthorityYear="1859" box="[306,457,801,827]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="loveni">
<emphasis id="B900EA8F5878CC6FD2274E4EFE1EFCE6" box="[306,457,801,827]" italics="true" pageId="9" pageNumber="460">Maera loveni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D1E5878CC6FD1144E4EFD7AFCE6" authorityName="G. Karaman &amp; Ruffo" authorityYear="1971" box="[513,685,801,827]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="pachytelson">
<emphasis id="B900EA8F5878CC6FD1144E4EFD7AFCE6" box="[513,685,801,827]" italics="true" pageId="9" pageNumber="460">M. pachytelson</emphasis>
</taxonomicName>
from
<collectingCountry id="F363760D5878CC6FD1E74E4DFCE9FCE6" box="[754,830,801,827]" name="France" pageId="9" pageNumber="460">France</collectingCountry>
and
<collectingCountry id="F363760D5878CC6FD0634E4DFC63FCE6" box="[886,948,801,827]" name="Israel" pageId="9" pageNumber="460">Israel</collectingCountry>
have minute notches on the coxa 2 and the coxae 12, respectively (
<figureCitation id="134F2A185878CC6FD2A54E29FDC5FC82" box="[432,530,837,863]" captionStart="FIGURE 4" captionStartId="6.[151,250,2004,2029]" captionTargetBox="[181,1408,190,1967]" captionTargetId="figure@6.[167,1420,181,1978]" captionTargetPageId="6" captionText="FIGURE 4. Maera loveni (Bruzelius, 1859). Male, 58.3 mm (OMNH-Ar-10623). Scales: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.3690510" httpUri="https://zenodo.org/record/3690510/files/figure.png" pageId="9" pageNumber="460">Fig. 4-G</figureCitation>
<figureCitation id="134F2A185878CC6FD1074E29FDF6FC82" box="[530,545,837,863]" captionStart="FIGURE 2" captionStartId="3.[151,250,1739,1764]" captionTargetBox="[161,1428,694,1706]" captionTargetId="figure@3.[152,1436,687,1714]" captionTargetPageId="3" captionText="FIGURE 2. Maera loveni (Bruzelius, 1859). Male, 58.3 mm (OMNH-Ar-10623). Habitus." figureDoi="http://doi.org/10.5281/zenodo.3690506" httpUri="https://zenodo.org/record/3690506/files/figure.png" pageId="9" pageNumber="460">2</figureCitation>
, G.
<bibRefCitation id="EFE54B6C5878CC6FD15A4E29FC88FC82" author="Karaman, G. &amp; Ruffo, S." box="[591,863,837,863]" pageId="9" pageNumber="460" pagination="113 - 176" refId="ref11856" refString="Karaman, G. &amp; Ruffo, S. (1971) Contributo alla conoscenza della specie Mediterranee del genere Maera. Memorie del Museo Civico di Storia Naturale, Verona, 19, 113 - 176." type="journal article" year="1971">Karaman &amp; Ruffo 1971</bibRefCitation>
).
<taxonomicName id="4C744D1E5878CC6FD0664E2AFB99FC82" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[883,1102,838,863]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sagamiensis" status="sp. nov.">
<emphasis id="B900EA8F5878CC6FD0664E2AFB99FC82" box="[883,1102,838,863]" italics="true" pageId="9" pageNumber="460">Maera sagamiensis</emphasis>
</taxonomicName>
<emphasis id="B900EA8F5878CC6FD7404E29FB78FC82" bold="true" box="[1109,1199,837,863]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="A23357F45878CC6FD7404E29FB78FC82" box="[1109,1199,837,863]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C744D1E5878CC6FD3C04E06FEE9FC5E" baseAuthorityName="Bruzelius" baseAuthorityYear="1859" box="[213,318,873,899]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="loveni">
<emphasis id="B900EA8F5878CC6FD3C04E06FEE9FC5E" box="[213,318,873,899]" italics="true" pageId="9" pageNumber="460">M. loveni</emphasis>
</taxonomicName>
by the shape of the gnathopod 2 palm, and from
<taxonomicName id="4C744D1E5878CC6FD0704E06FBC7FC5E" authorityName="G. Karaman &amp; Ruffo" authorityYear="1971" box="[869,1040,873,899]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="pachytelson">
<emphasis id="B900EA8F5878CC6FD0704E06FBC7FC5E" box="[869,1040,873,899]" italics="true" pageId="9" pageNumber="460">M. pachytelson</emphasis>
</taxonomicName>
by the shape of the telson.
</paragraph>
<paragraph id="8BCB369D5878CC6FD3D24EE0FC3FFC75" blockId="9.[151,1437,153,972]" box="[199,1000,908,936]" pageId="9" pageNumber="460">
<emphasis id="B900EA8F5878CC6FD3D24EE0FEFDFC7A" bold="true" box="[199,298,908,935]" pageId="9" pageNumber="460">Habitat.</emphasis>
Bottom sediment unknown,
<quantity id="4C8C9B785878CC6FD17B4EE1FD3AFC7A" box="[622,749,909,935]" metricMagnitude="2" metricUnit="m" metricValue="2.035" metricValueMax="2.7399999999999998" metricValueMin="1.33" pageId="9" pageNumber="460" unit="m" value="203.5" valueMax="274.0" valueMin="133.0">133274 m</quantity>
depth (present study).
</paragraph>
</subSubSection>
<subSubSection id="C36E65165878CC6FD3D24EDCFCDEFC11" box="[199,777,944,972]" pageId="9" pageNumber="460" type="distribution">
<paragraph id="8BCB369D5878CC6FD3D24EDCFCDEFC11" blockId="9.[151,1437,153,972]" box="[199,777,944,972]" pageId="9" pageNumber="460">
<emphasis id="B900EA8F5878CC6FD3D24EDCFEB6FC16" bold="true" box="[199,353,944,971]" pageId="9" pageNumber="460">Distribution.</emphasis>
Sagami Bay in
<collectingCountry id="F363760D5878CC6FD1064EDDFD84FC16" box="[531,595,945,971]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>