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<document ID-DOI="http://doi.org/10.5281/zenodo.4758482" ID-GBIF-Dataset="f0f8f265-10f1-448f-8b15-6c56fc140166" ID-GBIF-Taxon="182067784" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758482" ID-ZooBank="8BFBE3F7-E0EF-4EDD-BA82-9EA087F74688" LSID="urn:lsid:zoobank.org:act:73C51A6E-A31D-43D2-8C5A-89BA5148AAA0" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620834297829" checkinUser="felipe" docAuthor="Grubbs, Scott A." docDate="2015" docId="03DD8784FF9DA71FFE8058346513FCD7" docLanguage="en" docName="Illiesia.11.12.147-166.pdf" docOrigin="Illiesia 11 (12)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Leuctra schusteri Grubbs 2015, sp. n." docType="treatment" docUuid="73C51A6E-A31D-43D2-8C5A-89BA5148AAA0" docUuidSource="ZooBank" docVersion="7" lastPageId="7" lastPageNumber="154" masterDocId="FFE4FFFCFF98A718FFF65E23604AFFEB" masterDocTitle="Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa" masterLastPageNumber="166" masterPageNumber="147" pageId="5" pageNumber="152" updateTime="1632012907386" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Grubbs, Scott A.</mods:namePart>
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<mods:title>Illiesia</mods:title>
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<mods:date>2015</mods:date>
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<mods:number>11</mods:number>
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<mods:detail type="issue">
<mods:number>12</mods:number>
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<mods:start>147</mods:start>
<mods:end>166</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4752768</mods:identifier>
<mods:identifier type="GBIF-Dataset">f0f8f265-10f1-448f-8b15-6c56fc140166</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4758482" ID-GBIF-Taxon="182067784" ID-Zenodo-Dep="4758482" LSID="urn:lsid:zoobank.org:act:73C51A6E-A31D-43D2-8C5A-89BA5148AAA0" httpUri="http://treatment.plazi.org/id/03DD8784FF9DA71FFE8058346513FCD7" lastPageId="7" lastPageNumber="154" pageId="5" pageNumber="152">
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<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[374,578,1559,1583]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis bold="true" box="[374,578,1559,1583]" italics="true" pageId="5" pageNumber="152">Leuctra schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[585,650,1559,1583]" pageId="5" pageNumber="152" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph blockId="5.[189,790,1595,1691]" pageId="5" pageNumber="152">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName box="[579,710,1595,1619]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.specie sfile.org:TaxonName:472643
</paragraph>
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<subSubSection pageId="5" pageNumber="152" type="description">
<paragraph blockId="5.[189,790,1595,1691]" box="[420,567,1667,1691]" pageId="5" pageNumber="152">
(
<figureCitation box="[428,559,1667,1691]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[189,241,1588,1612]" captionStartId-1="6.[189,241,1153,1177]" captionTargetBox-0="[249,1383,254,1542]" captionTargetBox-1="[249,1383,254,1107]" captionTargetId-0="figure-193@4.[249,1383,254,1542]" captionTargetId-1="figure-422@6.[249,1383,254,1107]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-1="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." figureDoi-0="http://doi.org/10.5281/zenodo.4752776" figureDoi-1="http://doi.org/10.5281/zenodo.4752778" httpUri-0="https://zenodo.org/record/4752776/files/figure.png" httpUri-1="https://zenodo.org/record/4752778/files/figure.png" pageId="5" pageNumber="152">Figs. 1018</figureCitation>
)
</paragraph>
<paragraph blockId="5.[189,797,1739,1908]" lastBlockId="5.[835,1444,228,1907]" pageId="5" pageNumber="152">
<emphasis bold="true" box="[189,258,1739,1763]" pageId="5" pageNumber="152">Male.</emphasis>
Forewing length
<quantity box="[487,630,1739,1763]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="5" pageNumber="152" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
, body length 4.56.0 mm (n=15). General body color and wings brown. Cerci tubular and unmodified. Abdominal terga 16 unmodified. Dorsal process on abdominal tergum 7 emanating from anterior margin of segment, tapering markedly to the posterior tip that is slightly bulbous and expanded medially, overall shape of process triangular (
<figureCitation captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="5" pageNumber="152">Figs. 1011</figureCitation>
). Abdominal tergum 8 unmodified except for a quadrate-shaped lightly-pigmented medial sclerite. Epiproct sclerite unmodified and narrow throughout length, tapering slightly posteriorly. Specilla broadest basally, tapering little throughout length (
<figureCitation box="[936,1077,516,540]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="5" pageNumber="152">Figs. 1415</figureCitation>
), generally lacking tubercles along distal outer margins (
<figureCitation box="[1166,1298,552,576]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="5" pageNumber="152">Figs. 1215</figureCitation>
), tip bears a conspicuous, laterally-directed spur (
<figureCitation box="[1292,1427,588,612]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="5" pageNumber="152">Figs. 1213</figureCitation>
). Subanal lobes flat, slightly recurved anteriorly, and subequal in length to specilla (
<figureCitation box="[1205,1335,659,684]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="5" pageNumber="152">Figs. 1415</figureCitation>
). Vesicle small and subtriangular, length ca. equal to width at base (
<figureCitation box="[931,1009,732,756]" captionStart="Figs" captionStartId="6.[189,241,1153,1177]" captionTargetBox="[249,1383,254,1107]" captionTargetId="figure-422@6.[249,1383,254,1107]" captionTargetPageId="6" captionText="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." figureDoi="http://doi.org/10.5281/zenodo.4752778" httpUri="https://zenodo.org/record/4752778/files/figure.png" pageId="5" pageNumber="152">Fig. 18</figureCitation>
).
</paragraph>
<paragraph blockId="5.[835,1444,228,1907]" pageId="5" pageNumber="152">
<emphasis bold="true" box="[835,929,768,792]" pageId="5" pageNumber="152">Female.</emphasis>
Forewing length 5.57.0 mm, body length
<quantity box="[835,974,804,828]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="5" pageNumber="152" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.57.5 mm</quantity>
(n=21). General body color similar to male. Abdominal sternum 7 with a pentagonalshaped medial sclerite. Abdominal sternum 8 sclerotized medially in anterior ½; subgenital plate lobes rounded posterolaterally and angular along the mediolateral corners; lobes separated by a membranous diamond-shaped medial notch, extending maximally over the anterior ¼ of abdominal sternum 9 (
<figureCitation box="[1103,1232,1092,1116]" captionStart="Figs" captionStartId="6.[189,241,1153,1177]" captionTargetBox="[249,1383,254,1107]" captionTargetId="figure-422@6.[249,1383,254,1107]" captionTargetPageId="6" captionText="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." figureDoi="http://doi.org/10.5281/zenodo.4752778" httpUri="https://zenodo.org/record/4752778/files/figure.png" pageId="5" pageNumber="152">Figs. 1617</figureCitation>
).
</paragraph>
<paragraph blockId="5.[835,1444,228,1907]" box="[835,1074,1127,1151]" pageId="5" pageNumber="152">
<emphasis bold="true" box="[835,910,1127,1151]" pageId="5" pageNumber="152">Larva.</emphasis>
Undescribed.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="153" pageId="5" pageNumber="152" type="materials_examined">
<paragraph blockId="5.[835,1444,228,1907]" lastBlockId="6.[835,1444,1297,1932]" lastPageId="6" lastPageNumber="153" pageId="5" pageNumber="152">
<emphasis bold="true" box="[835,1074,1163,1187]" pageId="5" pageNumber="152">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111206385" collectingDate="2015-09-15" collectionCode="USNM" collectorName="S. A. Grubbs" country="United States of America" county="Hart Co." latitude="37.1974" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.0476" municipality="Cooper Spring" pageId="5" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="holotype">
<typeStatus box="[1085,1196,1163,1187]" pageId="5" pageNumber="152">Holotype</typeStatus>
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,
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<collectingCountry box="[1243,1301,1163,1187]" name="United States of America" pageId="5" pageNumber="152">USA</collectingCountry>
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,
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<collectingRegion box="[1318,1436,1163,1187]" country="United States of America" name="Kentucky" pageId="5" pageNumber="152">Kentucky</collectingRegion>
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,
<collectingCounty box="[835,960,1200,1224]" pageId="5" pageNumber="152">Hart Co.</collectingCounty>
,
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,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFCB55AF064F8FB00" box="[835,1202,1235,1260]" country="United States of America" county="Hart Co." latitude="37.1974" longLatPrecision="6" longitude="-86.0476" municipality="Cooper Spring" name="Mammoth Cave National Park" pageId="5" pageNumber="152" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[1215,1438,1235,1259]" pageId="5" pageNumber="152" value="2015-09-15">
<collectingDate box="[1215,1438,1235,1259]" pageId="5" pageNumber="152" value="2015-09-15">15 September 2015</collectingDate>
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,
<collectorName box="[835,981,1271,1295]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
(
<collectionCode box="[999,1084,1271,1295]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="152" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206374" collectingDate="2013-10-08" collectionCode="INHS, USNM, WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Allen Co." latitude="36.6777" location="Smyrna Creek" longLatPrecision="6" longitude="-86.3913" pageId="5" pageNumber="152" specimenCount="60" specimenCount-female="43" specimenCount-male="17" stateProvince="Kentucky" typeStatus="paratype">
<typeStatus box="[1109,1229,1272,1296]" pageId="5" pageNumber="152">Paratypes</typeStatus>
:
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<collectingRegion box="[1243,1361,1271,1295]" country="United States of America" name="Kentucky" pageId="5" pageNumber="152">Kentucky</collectingRegion>
</emphasis>
,
<collectingCounty pageId="5" pageNumber="152">Allen Co.</collectingCounty>
, spring into
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFBE35B3F648DFAD8" box="[1045,1223,1307,1332]" country="United States of America" county="Allen Co." latitude="36.6777" longLatPrecision="6" longitude="-86.3913" name="Smyrna Creek" pageId="5" pageNumber="152" stateProvince="Kentucky">Smyrna Creek</location>
,
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SE Gold City
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,
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,
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</date>
,
<collectorName pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[941,988,1379,1404]" pageId="5" pageNumber="152" type="male">17♂</specimenCount>
,
<specimenCount box="[1007,1054,1379,1404]" pageId="5" pageNumber="152" type="female">43♀</specimenCount>
(
<collectionCode box="[1074,1144,1380,1404]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="5" pageNumber="152" type="Other type of research institution/biorepository">INHS</collectionCode>
,
<collectionCode box="[1160,1245,1380,1404]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="152" type="Museum">USNM</collectionCode>
,
<collectionCode box="[1262,1352,1380,1404]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206334" collectingDate="2013-10-03" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Adair Co." latitude="37.202" location="Spout Spring Branch" longLatPrecision="6" longitude="-85.148" pageId="5" pageNumber="152" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingCounty pageId="5" pageNumber="152">Adair Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFC6E5BA46487FA74" box="[920,1229,1415,1439]" country="United States of America" county="Adair Co." latitude="37.202" longLatPrecision="6" longitude="-85.148" name="Spout Spring Branch" pageId="5" pageNumber="152" stateProvince="Kentucky">Spout Spring Branch</location>
,
<locationDeviation location="Campbellsville" pageId="5" pageNumber="152">
<quantity box="[1271,1374,1415,1439]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="152" unit="km" value="24.0">24 km</quantity>
SE Campbellsville
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,
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,
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<date box="[1251,1439,1451,1476]" pageId="5" pageNumber="152" value="2013-10-03">
<collectingDate box="[1251,1439,1451,1476]" pageId="5" pageNumber="152" value="2013-10-03">3 October 2013</collectingDate>
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<collectorName box="[835,981,1487,1511]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[994,1027,1487,1511]" pageId="5" pageNumber="152" type="male">1♂</specimenCount>
,
<specimenCount box="[1042,1076,1487,1511]" pageId="5" pageNumber="152" type="female">3♀</specimenCount>
(
<collectionCode box="[1091,1182,1487,1511]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206311" collectingDate="2013-10-03" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.1346" location="Sulphur Creek" longLatPrecision="6" longitude="-85.1946" pageId="5" pageNumber="152" specimenCount="14" specimenCount-female="13" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
tributary to
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFAB55BEC63C1F9E0" country="United States of America" latitude="37.1346" longLatPrecision="6" longitude="-85.1946" name="Sulphur Creek" pageId="5" pageNumber="152" stateProvince="Kentucky">Sulphur Creek</location>
,
<locationDeviation box="[925,1189,1523,1548]" location="Columbia" pageId="5" pageNumber="152">
<quantity box="[925,1004,1523,1548]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="152" unit="km" value="10.0">10 km</quantity>
NE Columbia
</locationDeviation>
,
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,
<geoCoordinate box="[1314,1412,1523,1547]" degrees="85.1946" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-85.1946">-85.1946</geoCoordinate>
,
<date pageId="5" pageNumber="152" value="2013-10-03">
<collectingDate pageId="5" pageNumber="152" value="2013-10-03">3 October 2013</collectingDate>
</date>
,
<collectorName box="[1027,1184,1559,1583]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[1207,1241,1559,1583]" pageId="5" pageNumber="152" type="male">1♂</specimenCount>
,
<specimenCount box="[1267,1314,1559,1583]" pageId="5" pageNumber="152" type="female">13♀</specimenCount>
(
<collectionCode box="[1340,1427,1559,1583]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206360" collectingDate="2015-09-15" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Edmonson Co." latitude="37.1983" location="Sal Hollow Trail" longLatPrecision="6" longitude="-86.1443" municipality="Hollow" pageId="5" pageNumber="152" specimenCount="10" specimenCount-female="9" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingCounty box="[835,1017,1595,1620]" pageId="5" pageNumber="152">Edmonson Co.</collectingCounty>
,
<collectingMethod box="[1034,1334,1595,1619]" pageId="5" pageNumber="152">springhead in Fish Trap</collectingMethod>
<collectingMunicipality box="[1347,1438,1595,1619]" pageId="5" pageNumber="152">Hollow</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFCB5587C644DF99C" box="[835,1031,1631,1655]" country="United States of America" county="Edmonson Co." latitude="37.1983" longLatPrecision="6" longitude="-86.1443" municipality="Hollow" name="Sal Hollow Trail" pageId="5" pageNumber="152" stateProvince="Kentucky">Sal Hollow Trail</location>
,
<geoCoordinate box="[1042,1131,1632,1656]" degrees="37.1983" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="37.1983">37.1983</geoCoordinate>
,
<geoCoordinate box="[1143,1239,1631,1655]" degrees="86.1443" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-86.1443">-86.1443</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFB12587C63A0F970" country="United States of America" county="Edmonson Co." latitude="37.1983" longLatPrecision="6" longitude="-86.1443" municipality="Hollow" name="Mammoth Cave National Park" pageId="5" pageNumber="152" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[1013,1236,1667,1691]" pageId="5" pageNumber="152" value="2015-09-15">
<collectingDate box="[1013,1236,1667,1691]" pageId="5" pageNumber="152" value="2015-09-15">15 September 2015</collectingDate>
</date>
,
<collectorName box="[1246,1391,1667,1691]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[1403,1437,1667,1691]" pageId="5" pageNumber="152" type="male">1♂</specimenCount>
,
<specimenCount box="[835,869,1703,1728]" pageId="5" pageNumber="152" type="female">9♀</specimenCount>
(
<collectionCode box="[884,971,1704,1728]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206316" collectingDate="2015-09-15" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.2393" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1476" municipality="Ugly Creek" pageId="5" pageNumber="152" specimenCount="15" specimenCount-female="10" specimenCount-male="5" stateProvince="Kentucky" typeStatus="paratype">
spring run into
<collectingMunicipality box="[1183,1321,1703,1727]" pageId="5" pageNumber="152">Ugly Creek</collectingMunicipality>
,
<geoCoordinate box="[1333,1422,1704,1728]" degrees="37.2393" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="37.2393">37.2393</geoCoordinate>
,
<geoCoordinate degrees="86.1476" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-86.1476">- 86.1476</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFC4A58E86522F908" box="[956,1384,1739,1763]" country="United States of America" latitude="37.2393" longLatPrecision="6" longitude="-86.1476" municipality="Ugly Creek" name="Mammoth Cave National Park" pageId="5" pageNumber="152" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date pageId="5" pageNumber="152" value="2015-09-15">
<collectingDate pageId="5" pageNumber="152" value="2015-09-15">15 September 2015</collectingDate>
</date>
,
<collectorName box="[1049,1200,1775,1800]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[1219,1252,1775,1799]" pageId="5" pageNumber="152" type="male">5♂</specimenCount>
,
<specimenCount box="[1272,1319,1775,1800]" pageId="5" pageNumber="152" type="female">10♀</specimenCount>
(
<collectionCode box="[1342,1427,1776,1800]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206384" collectingDate="2015-09-15" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.2374" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1092" municipality="Ugly Creek" pageId="5" pageNumber="152" specimenCount="17" specimenCount-female="9" specimenCount-male="8" stateProvince="Kentucky" typeStatus="paratype">
spring run into
<collectingMunicipality box="[1056,1205,1811,1835]" pageId="5" pageNumber="152">Ugly Creek</collectingMunicipality>
,
<geoCoordinate box="[1228,1318,1811,1835]" degrees="37.2374" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="37.2374">37.2374</geoCoordinate>
,
<geoCoordinate box="[1341,1439,1811,1835]" degrees="86.1092" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-86.1092">-86.1092</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9DA71DFCB5591464F8F8A4" box="[835,1202,1847,1871]" country="United States of America" latitude="37.2374" longLatPrecision="6" longitude="-86.1092" municipality="Ugly Creek" name="Mammoth Cave National Park" pageId="5" pageNumber="152" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[1215,1438,1847,1871]" pageId="5" pageNumber="152" value="2015-09-15">
<collectingDate box="[1215,1438,1847,1871]" pageId="5" pageNumber="152" value="2015-09-15">15 September 2015</collectingDate>
</date>
,
<collectorName box="[835,994,1883,1907]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[1020,1053,1883,1907]" pageId="5" pageNumber="152" type="male">8♂</specimenCount>
,
<specimenCount box="[1081,1115,1883,1907]" pageId="5" pageNumber="152" type="female">9♀</specimenCount>
(
<collectionCode box="[1144,1231,1883,1907]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206369" collectingDate="2015-09-17" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" lastPageId="6" lastPageNumber="153" latitude="37.2096" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1476" municipality="Good Church Spring" pageId="5" pageNumber="152" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Kentucky" typeStatus="paratype">
<collectingMunicipality lastPageId="6" lastPageNumber="153" pageId="5" pageNumber="152">Good Church Spring</collectingMunicipality>
,
<geoCoordinate box="[282,368,1297,1321]" degrees="37.2096" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.2096">37.2096</geoCoordinate>
,
<geoCoordinate box="[384,482,1297,1321]" degrees="86.1476" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.1476">-86.1476</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFE185B3260BCFAA6" country="United States of America" latitude="37.2096" longLatPrecision="6" longitude="-86.1476" municipality="Good Church Spring" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[267,509,1333,1357]" pageId="6" pageNumber="153" value="2015-09-17">
<collectingDate box="[267,509,1333,1357]" pageId="6" pageNumber="153" value="2015-09-17">17 September 2015</collectingDate>
</date>
,
<collectorName box="[531,685,1333,1357]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[706,740,1333,1357]" pageId="6" pageNumber="153" type="male">2♂</specimenCount>
,
<specimenCount box="[764,797,1333,1357]" pageId="6" pageNumber="153" type="female">5♀</specimenCount>
(
<collectionCode box="[197,284,1369,1393]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206340" collectingDate="2015-09-24" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.1716" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1836" municipality="Silent Grove Springhouse Cave" pageId="6" pageNumber="153" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
spring out of
<collectingMunicipality pageId="6" pageNumber="153">Silent Grove Springhouse Cave</collectingMunicipality>
,
<geoCoordinate box="[267,357,1405,1429]" degrees="37.1716" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.1716">37.1716</geoCoordinate>
,
<geoCoordinate box="[373,470,1405,1429]" degrees="86.1836" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.1836">-86.1836</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFE105B5E60BCFA52" country="United States of America" latitude="37.1716" longLatPrecision="6" longitude="-86.1836" municipality="Silent Grove Springhouse Cave" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[267,509,1441,1465]" pageId="6" pageNumber="153" value="2015-09-24">
<collectingDate box="[267,509,1441,1465]" pageId="6" pageNumber="153" value="2015-09-24">24 September 2015</collectingDate>
</date>
,
<collectorName box="[531,685,1441,1465]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[707,740,1441,1465]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
,
<specimenCount box="[764,797,1441,1465]" pageId="6" pageNumber="153" type="female">2♀</specimenCount>
(
<collectionCode box="[197,284,1477,1501]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206335" collectingDate="2015-10-05" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.2424" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1797" municipality="Blue Spring" pageId="6" pageNumber="153" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Kentucky" typeStatus="paratype">
<collectingMunicipality box="[314,462,1477,1501]" pageId="6" pageNumber="153">Blue Spring</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFE175BE660BCF9EA" country="United States of America" latitude="37.2424" longLatPrecision="6" longitude="-86.1797" municipality="Blue Spring" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<geoCoordinate box="[273,360,1513,1537]" degrees="37.2424" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.2424">37.2424</geoCoordinate>
,
<geoCoordinate box="[390,485,1513,1537]" degrees="86.1797" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.1797">-86.1797</geoCoordinate>
,
<date box="[514,723,1513,1537]" pageId="6" pageNumber="153" value="2015-10-05">
<collectingDate box="[514,723,1513,1537]" pageId="6" pageNumber="153" value="2015-10-05">5 October 2015</collectingDate>
</date>
,
<collectorName pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[293,327,1549,1573]" pageId="6" pageNumber="153" type="male">3♂</specimenCount>
,
<specimenCount box="[344,377,1549,1573]" pageId="6" pageNumber="153" type="female">3♀</specimenCount>
(
<collectionCode box="[395,486,1549,1573]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206387" collectingDate="2015-10-11" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.234" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1547" municipality="Sand Spring" pageId="6" pageNumber="153" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingMunicipality box="[508,658,1549,1573]" pageId="6" pageNumber="153">Sand Spring</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFD56582E61FAF9A2" country="United States of America" latitude="37.234" longLatPrecision="6" longitude="-86.1547" municipality="Sand Spring" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<geoCoordinate box="[447,534,1585,1609]" degrees="37.2340" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.234">37.2340</geoCoordinate>
,
<geoCoordinate box="[552,648,1585,1609]" degrees="86.1547" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.1547">-86.1547</geoCoordinate>
,
<date pageId="6" pageNumber="153" value="2015-10-11">
<collectingDate pageId="6" pageNumber="153" value="2015-10-11">11 October 2015</collectingDate>
</date>
,
<collectorName box="[261,411,1621,1645]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[428,462,1621,1645]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
(
<collectionCode box="[481,572,1621,1645]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206304" collectingDate="2015-10-11" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.2404" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.1738" municipality="Wet Prong Buffalo Creek" pageId="6" pageNumber="153" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
tributary to
<collectingMunicipality pageId="6" pageNumber="153">Wet Prong Buffalo Creek</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFE2D585A60BCF95E" country="United States of America" latitude="37.2404" longLatPrecision="6" longitude="-86.1738" municipality="Wet Prong Buffalo Creek" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<geoCoordinate box="[271,358,1693,1717]" degrees="37.2404" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.2404">37.2404</geoCoordinate>
,
<geoCoordinate box="[385,481,1693,1717]" degrees="86.1738" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.1738">-86.1738</geoCoordinate>
,
<date box="[507,725,1693,1717]" pageId="6" pageNumber="153" value="2015-10-11">
<collectingDate box="[507,725,1693,1717]" pageId="6" pageNumber="153" value="2015-10-11">11 October 2015</collectingDate>
</date>
,
<collectorName pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[296,329,1729,1753]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
,
<specimenCount box="[349,382,1729,1753]" pageId="6" pageNumber="153" type="female">1♀</specimenCount>
(
<collectionCode box="[403,494,1729,1753]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206375" collectingDate="2013-10-01" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Green Co." latitude="37.3441" location="Rte." longLatPrecision="6" longitude="-85.6236" municipality="Big Brush Creek" pageId="6" pageNumber="153" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kentucky" typeStatus="paratype">
<collectingCounty box="[518,645,1729,1753]" pageId="6" pageNumber="153">Green Co.</collectingCounty>
, spring into
<collectingMunicipality box="[189,412,1765,1789]" pageId="6" pageNumber="153">Big Brush Creek</collectingMunicipality>
, nr.
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFE1B58C66251F917" box="[493,539,1765,1788]" country="United States of America" county="Green Co." latitude="37.3441" longLatPrecision="6" longitude="-85.6236" municipality="Big Brush Creek" name="Rte." pageId="6" pageNumber="153" stateProvince="Kentucky">Rte.</location>
569, 9 km WNW
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFF4B592A6128F8CA" box="[189,354,1801,1825]" country="United States of America" county="Green Co." latitude="37.3441" longLatPrecision="6" longitude="-85.6236" municipality="Big Brush Creek" name="Summersville" pageId="6" pageNumber="153" stateProvince="Kentucky">Summersville</location>
,
<geoCoordinate box="[374,463,1801,1825]" degrees="37.3441" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.3441">37.3441</geoCoordinate>
,
<geoCoordinate box="[483,580,1801,1825]" degrees="85.6236" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-85.6236">-85.6236</geoCoordinate>
,
<date box="[600,793,1801,1825]" pageId="6" pageNumber="153" value="2013-10-01">
<collectingDate box="[600,793,1801,1825]" pageId="6" pageNumber="153" value="2013-10-01">1 October 2013</collectingDate>
</date>
,
<collectorName box="[189,334,1837,1861]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[346,379,1837,1861]" pageId="6" pageNumber="153" type="male">2♂</specimenCount>
,
<specimenCount box="[393,427,1837,1861]" pageId="6" pageNumber="153" type="female">3♀</specimenCount>
(
<collectionCode box="[442,533,1837,1861]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206383" collectingDate="2001-07-17" collectionCode="WKUC" collectorName="S. A. Grubbs &amp; J. M Taylor" country="United States of America" county="Hart Co." latitude="37.4105" location="Roundstone Creek" longLatPrecision="6" longitude="-85.9823" pageId="6" pageNumber="153" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingCounty box="[552,656,1837,1861]" pageId="6" pageNumber="153">Hart Co.</collectingCounty>
, spring into
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFF4B597261E7F882" box="[189,429,1873,1897]" country="United States of America" county="Hart Co." latitude="37.4105" longLatPrecision="6" longitude="-85.9823" name="Roundstone Creek" pageId="6" pageNumber="153" stateProvince="Kentucky">Roundstone Creek</location>
,
<locationDeviation box="[459,792,1873,1897]" location="Bonnieville" pageId="6" pageNumber="153">
<quantity box="[459,556,1873,1897]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="153" unit="km" value="7.5">7.5 km</quantity>
NW Bonnieville
</locationDeviation>
,
<geoCoordinate box="[189,279,1909,1933]" degrees="37.4105" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.4105">37.4105</geoCoordinate>
,
<geoCoordinate box="[290,388,1909,1933]" degrees="85.9823" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-85.9823">-85.9823</geoCoordinate>
,
<date box="[399,543,1909,1933]" pageId="6" pageNumber="153" value="2001-07-17">
<collectingDate box="[399,543,1909,1933]" pageId="6" pageNumber="153" value="2001-07-17">17 July 2001</collectingDate>
</date>
,
<collectorName box="[554,696,1909,1933]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
and
<collectorName box="[755,915,1297,1932]" pageId="6" pageNumber="153">J.M Taylor</collectorName>
,
<specimenCount box="[935,968,1297,1321]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
, 3 nymphs (
<collectionCode box="[1139,1230,1297,1321]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206344" collectingDate="2002-09-07" collectingDateMax="2002-09-28" collectingDateMin="2002-09-07" collectionCode="WKUC" collectorName="Grubbs" country="United States of America" pageId="6" pageNumber="153" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Kentucky" typeStatus="paratype">
same but
<date pageId="6" pageNumber="153" value="2002-09-07" valueMax="2002-09-28" valueMin="2002-09-07">
<collectingDate pageId="6" pageNumber="153" value="2002-09-07" valueMax="2002-09-28" valueMin="2002-09-07">728 September 2002</collectingDate>
</date>
, S.A
<collectorName box="[1114,1204,1333,1357]" pageId="6" pageNumber="153">Grubbs</collectorName>
,
<specimenCount box="[1225,1259,1333,1357]" pageId="6" pageNumber="153" type="male">5♂</specimenCount>
,
<specimenCount box="[1283,1316,1333,1357]" pageId="6" pageNumber="153" type="female">6♀</specimenCount>
(
<collectionCode box="[1341,1431,1333,1357]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206349" collectingDate="2003-09-26" collectionCode="WKUC" collectorName="Grubbs" country="United States of America" pageId="6" pageNumber="153" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Kentucky" typeStatus="paratype">
same but
<date box="[957,1187,1369,1393]" pageId="6" pageNumber="153" value="2003-09-26">
<collectingDate box="[957,1187,1369,1393]" pageId="6" pageNumber="153" value="2003-09-26">26 September 2003</collectingDate>
</date>
, S.A
<collectorName box="[1254,1343,1369,1393]" pageId="6" pageNumber="153">Grubbs</collectorName>
,
<specimenCount box="[1359,1392,1369,1393]" pageId="6" pageNumber="153" type="male">7♂</specimenCount>
,
<specimenCount box="[1410,1443,1369,1393]" pageId="6" pageNumber="153" type="female">6♀</specimenCount>
(
<collectionCode box="[843,933,1405,1429]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206305" collectingDate="2013-10-01" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Lynn Camp Creek" latitude="37.3545" location="Summersville" longLatPrecision="6" longitude="-85.7128" municipality="Rte." pageId="6" pageNumber="153" specimenCount="14" specimenCount-female="13" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
spring into
<collectingCounty box="[1105,1335,1405,1430]" pageId="6" pageNumber="153">Lynn Camp Creek</collectingCounty>
, nr.
<collectingMunicipality box="[1397,1443,1406,1429]" pageId="6" pageNumber="153">Rte.</collectingMunicipality>
569, 14 km NW
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFBFD5B8264FAFA52" box="[1035,1200,1441,1465]" country="United States of America" county="Lynn Camp Creek" latitude="37.3545" longLatPrecision="6" longitude="-85.7128" municipality="Rte." name="Summersville" pageId="6" pageNumber="153" stateProvince="Kentucky">Summersville</location>
,
<geoCoordinate box="[1214,1303,1441,1465]" degrees="37.3545" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.3545">37.3545</geoCoordinate>
,
<geoCoordinate box="[1318,1416,1441,1465]" degrees="85.7128" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-85.7128">-85.7128</geoCoordinate>
,
<date pageId="6" pageNumber="153" value="2013-10-01">
<collectingDate pageId="6" pageNumber="153" value="2013-10-01">1 October 2013</collectingDate>
</date>
,
<collectorName box="[1004,1149,1477,1501]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[1162,1196,1477,1501]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
,
<specimenCount box="[1210,1257,1477,1501]" pageId="6" pageNumber="153" type="female">13♀</specimenCount>
(
<collectionCode box="[1272,1362,1477,1501]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206337" collectionCode="WKUC" country="United States of America" pageId="6" pageNumber="153" specimenCount="14" specimenCount-female="10" specimenCount-male="4" stateProvince="Kentucky" typeStatus="paratype">
same as
<typeStatus box="[877,984,1513,1537]" pageId="6" pageNumber="153">holotype</typeStatus>
,
<specimenCount box="[1007,1040,1513,1537]" pageId="6" pageNumber="153" type="male">4♂</specimenCount>
,
<specimenCount box="[1064,1111,1513,1537]" pageId="6" pageNumber="153" type="female">10♀</specimenCount>
(
<collectionCode box="[1136,1226,1513,1537]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206341" collectingDate="2015-09-15" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="37.2061" location="Mammoth Cave National Park" longLatPrecision="6" longitude="-86.0489" municipality="Collins Spring" pageId="6" pageNumber="153" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingMunicipality box="[1256,1439,1513,1537]" pageId="6" pageNumber="153">Collins Spring</collectingMunicipality>
,
<geoCoordinate box="[835,924,1549,1573]" degrees="37.2061" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.2061">37.2061</geoCoordinate>
,
<geoCoordinate box="[941,1039,1549,1573]" degrees="86.0489" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.0489">-86.0489</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFBD6582E65D4F9CE" box="[1056,1438,1549,1573]" country="United States of America" latitude="37.2061" longLatPrecision="6" longitude="-86.0489" municipality="Collins Spring" name="Mammoth Cave National Park" pageId="6" pageNumber="153" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<date box="[835,1063,1585,1609]" pageId="6" pageNumber="153" value="2015-09-15">
<collectingDate box="[835,1063,1585,1609]" pageId="6" pageNumber="153" value="2015-09-15">15 September 2015</collectingDate>
</date>
,
<collectorName box="[1077,1222,1585,1609]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[1240,1273,1585,1609]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
,
<specimenCount box="[1290,1323,1585,1609]" pageId="6" pageNumber="153" type="female">4♀</specimenCount>
(
<collectionCode box="[1340,1427,1585,1609]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206310" collectingDate="2013-10-04" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Warren Co." latitude="37.139" location="Shanty Hollow Lake" longLatPrecision="6" longitude="-86.3849" pageId="6" pageNumber="153" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<collectingCounty box="[835,977,1621,1645]" pageId="6" pageNumber="153">Warren Co.</collectingCounty>
, tributary to
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9DA71FFE8058346513FCD7:8EAB6049FF9EA71EFB8058766538F986" box="[1142,1394,1621,1645]" country="United States of America" county="Warren Co." latitude="37.139" longLatPrecision="6" longitude="-86.3849" name="Shanty Hollow Lake" pageId="6" pageNumber="153" stateProvince="Kentucky">Shanty Hollow Lake</location>
,
<locationDeviation location="Richardsville" pageId="6" pageNumber="153">
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="153" unit="km" value="8.5">8.5 km</quantity>
NE Richardsville
</locationDeviation>
,
<geoCoordinate box="[1107,1196,1657,1681]" degrees="37.1390" direction="north" orientation="latitude" pageId="6" pageNumber="153" precision="5" value="37.139">37.1390</geoCoordinate>
,
<geoCoordinate box="[1212,1310,1657,1681]" degrees="86.3849" direction="west" orientation="longitude" pageId="6" pageNumber="153" precision="5" value="-86.3849">-86.3849</geoCoordinate>
,
<date pageId="6" pageNumber="153" value="2013-10-04">
<collectingDate pageId="6" pageNumber="153" value="2013-10-04">4 October 2013</collectingDate>
</date>
,
<collectorName box="[902,1046,1693,1717]" pageId="6" pageNumber="153">S.A. Grubbs</collectorName>
,
<specimenCount box="[1057,1091,1693,1717]" pageId="6" pageNumber="153" type="male">1♂</specimenCount>
,
<specimenCount box="[1104,1138,1693,1717]" pageId="6" pageNumber="153" type="female">4♀</specimenCount>
(
<collectionCode box="[1152,1243,1693,1717]" pageId="6" pageNumber="153">WKUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4752778" ID-Zenodo-Dep="4752778" httpUri="https://zenodo.org/record/4752778/files/figure.png" pageId="6" pageNumber="153" startId="6.[189,241,1153,1177]" targetBox="[249,1383,254,1107]" targetPageId="6">
<paragraph blockId="6.[189,1443,1153,1213]" pageId="6" pageNumber="153">
Figs. 1618.
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[336,524,1153,1177]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[336,524,1153,1177]" italics="true" pageId="6" pageNumber="153">Leuctra schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[533,599,1154,1177]" pageId="6" pageNumber="153" rank="species">sp. n.</taxonomicNameLabel>
, Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X.
</paragraph>
</caption>
<subSubSection pageId="6" pageNumber="153" type="etymology">
<paragraph blockId="6.[835,1444,1297,1932]" pageId="6" pageNumber="153">
<emphasis bold="true" box="[835,973,1729,1753]" pageId="6" pageNumber="153">Etymology.</emphasis>
This species is named in honor of Dr. Guenter Schuster, Foundation Professor of Biological Sciences, Eastern
<collectingRegion box="[1183,1297,1801,1825]" country="United States of America" name="Kentucky" pageId="6" pageNumber="153">Kentucky</collectingRegion>
University. Dr. Schuster and his students have spent many years studying the aquatic fauna of
<collectingRegion box="[1256,1373,1873,1897]" country="United States of America" name="Kentucky" pageId="6" pageNumber="153">Kentucky</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="154" pageId="6" pageNumber="153" type="diagnosis">
<paragraph blockId="6.[835,1444,1297,1932]" lastBlockId="7.[189,798,228,1908]" lastPageId="7" lastPageNumber="154" pageId="6" pageNumber="153">
<emphasis bold="true" box="[835,964,1908,1932]" pageId="6" pageNumber="153">Diagnosis.</emphasis>
The combination of the well-defined dorsal process on the 7
<superScript attach="left" box="[471,486,228,242]" fontSize="6" pageId="7" pageNumber="154">th</superScript>
abdominal tergum and a small triangular vesicle places
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[562,689,264,288]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[562,689,264,288]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[699,764,265,288]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
in the
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[233,328,301,324]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[233,328,301,324]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
species group (
<bibRefCitation author="Harper, P. P. &amp; F. Harper" box="[514,785,300,324]" pageId="7" pageNumber="154" pagination="467 - 472" refId="ref9363" refString="Harper, P. P. &amp; F. Harper. 1997. The genus Leuctra Stephens in North America: a preliminary report. Pages 467 - 472. In Landolt, P &amp; M. Sartori, editors. Ephemeroptera &amp; Plecoptera: Biology-Ecology-Systematics. Fribourg, MTL." type="book chapter" year="1997">Harper &amp; Harper 1997</bibRefCitation>
). The posteriorly tapering process on the 7
<superScript attach="right" box="[783,798,337,351]" fontSize="6" pageId="7" pageNumber="154">th</superScript>
abdominal tergum of
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[447,573,372,396]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[447,573,372,396]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[580,643,372,395]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[659,788,372,396]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="7" pageNumber="154">Figs. 1011</figureCitation>
) is similar only to
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[410,508,409,432]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[410,508,409,432]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
(
<figureCitation box="[529,661,408,432]" captionStart-0="Figs" captionStart-1="Figs 49" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionStartId-2="4.[189,241,1588,1612]" captionStartId-3="6.[189,241,1153,1177]" captionStartId-4="8.[189,241,1152,1176]" captionStartId-5="10.[189,241,1584,1608]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetBox-2="[249,1383,254,1542]" captionTargetBox-3="[249,1383,254,1107]" captionTargetBox-4="[249,1383,254,1105]" captionTargetBox-5="[249,1383,254,1538]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetId-2="figure-193@4.[249,1383,254,1542]" captionTargetId-3="figure-422@6.[249,1383,254,1107]" captionTargetId-4="figure-432@8.[249,1383,254,1105]" captionTargetId-5="figure-197@10.[249,1383,254,1538]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-2="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-3="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." captionText-4="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." captionText-5="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" figureDoi-2="http://doi.org/10.5281/zenodo.4752776" figureDoi-3="http://doi.org/10.5281/zenodo.4752778" figureDoi-4="http://doi.org/10.5281/zenodo.4752780" figureDoi-5="http://doi.org/10.5281/zenodo.4752784" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" httpUri-2="https://zenodo.org/record/4752776/files/figure.png" httpUri-3="https://zenodo.org/record/4752778/files/figure.png" httpUri-4="https://zenodo.org/record/4752780/files/figure.png" httpUri-5="https://zenodo.org/record/4752784/files/figure.png" pageId="7" pageNumber="154">Figs. 2325</figureCitation>
), and very distinct from
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[364,527,444,468]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[364,527,444,468]" italics="true" pageId="7" pageNumber="154">L. carolinensis</emphasis>
</taxonomicName>
(
<figureCitation box="[551,660,444,468]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="7" pageNumber="154">Figs. 13</figureCitation>
),
<taxonomicName box="[691,797,444,468]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[691,797,444,468]" italics="true" pageId="7" pageNumber="154">L. tenella</emphasis>
</taxonomicName>
(
<figureCitation box="[198,276,480,504]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="7" pageNumber="154">Fig. 19</figureCitation>
),
<taxonomicName authorityName=": Needham and Claassen" authorityYear="1925" box="[298,396,480,504]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="triloba">
<emphasis box="[298,396,480,504]" italics="true" pageId="7" pageNumber="154">L. triloba</emphasis>
</taxonomicName>
(
<figureCitation box="[412,541,480,504]" captionStart-0="Figs" captionStart-1="Figs 49" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStart-7="Figs" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionStartId-2="4.[189,241,1588,1612]" captionStartId-3="6.[189,241,1153,1177]" captionStartId-4="8.[189,241,1152,1176]" captionStartId-5="10.[189,241,1584,1608]" captionStartId-6="11.[189,241,1592,1616]" captionStartId-7="13.[189,241,1706,1730]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetBox-2="[249,1383,254,1542]" captionTargetBox-3="[249,1383,254,1107]" captionTargetBox-4="[249,1383,254,1105]" captionTargetBox-5="[249,1383,254,1538]" captionTargetBox-6="[249,1383,254,1546]" captionTargetBox-7="[249,1383,362,1658]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetId-2="figure-193@4.[249,1383,254,1542]" captionTargetId-3="figure-422@6.[249,1383,254,1107]" captionTargetId-4="figure-432@8.[249,1383,254,1105]" captionTargetId-5="figure-197@10.[249,1383,254,1538]" captionTargetId-6="figure-151@11.[249,1383,254,1546]" captionTargetId-7="figure-160@13.[249,1383,362,1659]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="13" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-2="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-3="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." captionText-4="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." captionText-5="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." captionText-6="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." captionText-7="Figs. 3438. Leuctra triloba. 34, 7th abdominal process, dorsal view, 350X, Alabama, tributary to West Fork Hatchett Creek; 35, 7th abdominal process, dorsal view, 350X, North Carolina, Wine Spring Creek; 36, L. triloba, 7th abdominal process, dorsal view, 500X, Tennessee, tributary to South Suck Creek; 37, paraprocts, caudal view, 750X, North Carolina, Wine Spring Creek; 38, L. triloba, paraprocts, caudal view, 1500X, North Carolina, Wine Spring Creek. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" figureDoi-2="http://doi.org/10.5281/zenodo.4752776" figureDoi-3="http://doi.org/10.5281/zenodo.4752778" figureDoi-4="http://doi.org/10.5281/zenodo.4752780" figureDoi-5="http://doi.org/10.5281/zenodo.4752784" figureDoi-6="http://doi.org/10.5281/zenodo.4752786" figureDoi-7="http://doi.org/10.5281/zenodo.4752788" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" httpUri-2="https://zenodo.org/record/4752776/files/figure.png" httpUri-3="https://zenodo.org/record/4752778/files/figure.png" httpUri-4="https://zenodo.org/record/4752780/files/figure.png" httpUri-5="https://zenodo.org/record/4752784/files/figure.png" httpUri-6="https://zenodo.org/record/4752786/files/figure.png" httpUri-7="https://zenodo.org/record/4752788/files/figure.png" pageId="7" pageNumber="154">Figs. 3436</figureCitation>
),
<taxonomicName authorityName="Grubbs" authorityYear="2010" box="[563,637,480,504]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="usdi">
<emphasis box="[563,637,480,504]" italics="true" pageId="7" pageNumber="154">L. usdi</emphasis>
</taxonomicName>
(
<figureCitation box="[653,729,480,504]" captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="7" pageNumber="154">Fig. 43</figureCitation>
), and
<taxonomicName authorityName="Hanson" authorityYear="1941" box="[189,317,516,540]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis box="[189,317,516,540]" italics="true" pageId="7" pageNumber="154">L. variabilis</emphasis>
</taxonomicName>
(
<figureCitation box="[333,410,516,540]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="7" pageNumber="154">Fig. 48</figureCitation>
).
</paragraph>
<paragraph blockId="7.[189,798,228,1908]" pageId="7" pageNumber="154">
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[227,413,552,576]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[227,413,552,576]" italics="true" pageId="7" pageNumber="154">Leuctra schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[419,482,552,575]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[189,287,589,612]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[189,287,589,612]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
by paraproct characteristics. The subanal lobes of
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[300,430,624,648]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[300,430,624,648]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[443,512,624,647]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
are approximately the same length as the specilla (
<figureCitation box="[544,678,659,684]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="7" pageNumber="154">Figs. 1415</figureCitation>
) whereas with
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[255,354,697,720]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[255,354,697,720]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
the subanal lobes extend ca. ¾ the length of the specilla (
<figureCitation box="[458,631,732,756]" captionStart="Figs" captionStartId="11.[189,241,1592,1616]" captionTargetBox="[249,1383,254,1546]" captionTargetId="figure-151@11.[249,1383,254,1546]" captionTargetPageId="11" captionText="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752786" httpUri="https://zenodo.org/record/4752786/files/figure.png" pageId="7" pageNumber="154">Figs. 2830, 32</figureCitation>
). The specilla of
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[222,348,768,792]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[222,348,768,792]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[357,422,768,791]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
possess a prominent, laterallydirected spur (
<figureCitation box="[366,496,804,828]" captionStart-0="Figs" captionStart-1="Figs 49" captionStart-2="Figs" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionStartId-2="4.[189,241,1588,1612]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetBox-2="[249,1383,254,1542]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetId-2="figure-193@4.[249,1383,254,1542]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-2="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" figureDoi-2="http://doi.org/10.5281/zenodo.4752776" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" httpUri-2="https://zenodo.org/record/4752776/files/figure.png" pageId="7" pageNumber="154">Figs. 1212</figureCitation>
). In contrast, the specilla of
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[227,328,841,864]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[227,328,841,864]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
is rounded distally and lack a spur (
<figureCitation box="[198,295,876,900]" captionStart="Figs" captionStartId="10.[189,241,1584,1608]" captionTargetBox="[249,1383,254,1538]" captionTargetId="figure-197@10.[249,1383,254,1538]" captionTargetPageId="10" captionText="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752784" httpUri="https://zenodo.org/record/4752784/files/figure.png" pageId="7" pageNumber="154">Figs. 27</figureCitation>
,
<figureCitation box="[317,391,876,900]" captionStart="Figs" captionStartId="11.[189,241,1592,1616]" captionTargetBox="[249,1383,254,1546]" captionTargetId="figure-151@11.[249,1383,254,1546]" captionTargetPageId="11" captionText="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752786" httpUri="https://zenodo.org/record/4752786/files/figure.png" pageId="7" pageNumber="154">31, 33</figureCitation>
). The dorsal process on the 7
<superScript attach="right" box="[783,798,876,890]" fontSize="6" pageId="7" pageNumber="154">th</superScript>
abdominal tergum of
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[447,573,912,936]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[447,573,912,936]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[580,643,912,935]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[659,788,912,936]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="7" pageNumber="154">Figs. 1011</figureCitation>
) and
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[242,339,948,971]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[242,339,948,971]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
(
<figureCitation box="[356,486,948,972]" captionStart-0="Figs" captionStart-1="Figs 49" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionStartId-2="4.[189,241,1588,1612]" captionStartId-3="6.[189,241,1153,1177]" captionStartId-4="8.[189,241,1152,1176]" captionStartId-5="10.[189,241,1584,1608]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetBox-2="[249,1383,254,1542]" captionTargetBox-3="[249,1383,254,1107]" captionTargetBox-4="[249,1383,254,1105]" captionTargetBox-5="[249,1383,254,1538]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetId-2="figure-193@4.[249,1383,254,1542]" captionTargetId-3="figure-422@6.[249,1383,254,1107]" captionTargetId-4="figure-432@8.[249,1383,254,1105]" captionTargetId-5="figure-197@10.[249,1383,254,1538]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-2="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-3="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." captionText-4="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." captionText-5="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" figureDoi-2="http://doi.org/10.5281/zenodo.4752776" figureDoi-3="http://doi.org/10.5281/zenodo.4752778" figureDoi-4="http://doi.org/10.5281/zenodo.4752780" figureDoi-5="http://doi.org/10.5281/zenodo.4752784" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" httpUri-2="https://zenodo.org/record/4752776/files/figure.png" httpUri-3="https://zenodo.org/record/4752778/files/figure.png" httpUri-4="https://zenodo.org/record/4752780/files/figure.png" httpUri-5="https://zenodo.org/record/4752784/files/figure.png" pageId="7" pageNumber="154">Figs. 2325</figureCitation>
) are too similar in overall appearance, namely shape and size, to provide reliable diagnostic value.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="154" type="discussion">
<paragraph blockId="7.[189,798,228,1908]" pageId="7" pageNumber="154">
As is the case for most Nearctic
<taxonomicName box="[708,790,1056,1079]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis box="[708,790,1056,1079]" italics="true" pageId="7" pageNumber="154">Leuctra</emphasis>
</taxonomicName>
, possessing females definitively associated with males is the easiest means of differentiating between species. Females of
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[539,666,1163,1187]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[539,666,1163,1187]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[676,742,1164,1187]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[189,307,1200,1223]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[189,307,1200,1223]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
are very similar morphologically, particularly in the shape of the subgenital plate lobes and pigmentation patterns in darker individuals. Both species have lateral indentations that essentially demarcate the anterior plane of the lobes.
</paragraph>
<paragraph blockId="7.[189,798,228,1908]" pageId="7" pageNumber="154">
<emphasis bold="true" box="[189,304,1415,1439]" pageId="7" pageNumber="154">Remarks.</emphasis>
The type locality is a small spring run that emanates from a limestone bluff in southcentral
<collectingRegion box="[282,396,1487,1511]" country="United States of America" name="Kentucky" pageId="7" pageNumber="154">Kentucky</collectingRegion>
(
<figureCitation box="[417,552,1487,1511]" captionStart="Figs" captionStartId="17.[189,241,1793,1817]" captionTargetBox="[324,1307,255,1746]" captionTargetId="figure-111@17.[324,1307,254,1747]" captionTargetPageId="17" captionText="Figs. 53-54. 53, Location of the Leuctra schusteri sp. n. collection sites. IL = Illinois, IN = Indiana, KY = Kentucky, MO = Missouri, OH = Ohio, TN = Tennessee, VA = Virginia, WV = West Virginia. TL = type locality; 54, Leuctra schusteri sp. n. type locality (Cooper Spring, Mammoth Cave National Park, Kentucky, USA) facing downstream." figureDoi="http://doi.org/10.5281/zenodo.4752796" httpUri="https://zenodo.org/record/4752796/files/figure.png" pageId="7" pageNumber="154">Figs. 5354</figureCitation>
). Most adults were collected proximal to the spring source. All
<typeStatus box="[189,294,1560,1584]" pageId="7" pageNumber="154">paratype</typeStatus>
localities are also springheads or small spring runs. No other species of stoneflies were collected in the adult stage.
</paragraph>
<paragraph blockId="7.[189,798,228,1908]" lastBlockId="7.[835,1444,228,828]" pageId="7" pageNumber="154">
The type and all
<typeStatus box="[430,535,1668,1692]" pageId="7" pageNumber="154">paratype</typeStatus>
localities (
<figureCitation box="[665,744,1667,1691]" captionStart="Figs" captionStartId="17.[189,241,1793,1817]" captionTargetBox="[324,1307,255,1746]" captionTargetId="figure-111@17.[324,1307,254,1747]" captionTargetPageId="17" captionText="Figs. 53-54. 53, Location of the Leuctra schusteri sp. n. collection sites. IL = Illinois, IN = Indiana, KY = Kentucky, MO = Missouri, OH = Ohio, TN = Tennessee, VA = Virginia, WV = West Virginia. TL = type locality; 54, Leuctra schusteri sp. n. type locality (Cooper Spring, Mammoth Cave National Park, Kentucky, USA) facing downstream." figureDoi="http://doi.org/10.5281/zenodo.4752796" httpUri="https://zenodo.org/record/4752796/files/figure.png" pageId="7" pageNumber="154">Fig. 53</figureCitation>
) are located within the Interior Plateau Level III Ecoregion, a region characterized by Mississippianage limestone, extensive karst development, and emergent spring systems (
<bibRefCitation author="Woods, A. J. &amp; J. M. Omernik &amp; W. H. Martin &amp; G. J. Pond &amp; W. M. Andrews &amp; S. M. Call &amp; J. A. Comstock &amp; D. D. Taylor" box="[510,727,1811,1835]" pageId="7" pageNumber="154" refId="ref9893" refString="Woods, A. J., J. M. Omernik, W. H. Martin, G. J. Pond, W. M. Andrews, S. M. Call, J. A. Comstock &amp; D. D. Taylor. 2002. Ecoregions of Kentucky (color poster with map, descriptive text, summary tables, and photographs): Reston, VA, U. S Geographical Survey (map scale 1: 1,000,000)." type="book" year="2002">Woods et al. 2002</bibRefCitation>
). The range of this species likely extends southward into
<collectingRegion box="[189,310,1884,1907]" country="United States of America" name="Tennessee" pageId="7" pageNumber="154">Tennessee</collectingRegion>
since
<specimenCount box="[394,554,1884,1908]" pageId="7" pageNumber="154" type="generic" typeStatus="paratype">one paratype</specimenCount>
locality (Allen Co., spring into Smyrna Creek) is only ca.
<quantity box="[1282,1340,228,252]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="154" unit="km" value="3.0">3 km</quantity>
north of the state border. Northward into unglaciated southern
<collectingRegion box="[950,1041,300,324]" country="United States of America" name="Indiana" pageId="7" pageNumber="154">Indiana</collectingRegion>
and southern
<collectingRegion box="[1220,1279,300,324]" country="United States of America" name="Ohio" pageId="7" pageNumber="154">Ohio</collectingRegion>
,
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[1295,1393,301,324]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[1295,1393,301,324]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
has been the only autumn-emergent
<taxonomicName box="[1239,1321,337,360]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis box="[1239,1321,337,360]" italics="true" pageId="7" pageNumber="154">Leuctra</emphasis>
</taxonomicName>
collected. Similarly, only
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[1053,1166,373,396]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[1053,1166,373,396]" italics="true" pageId="7" pageNumber="154">L. tenuis</emphasis>
</taxonomicName>
has been collected eastward into the Appalachian Plateau region of eastern
<collectingRegion box="[927,1041,444,468]" country="United States of America" name="Kentucky" pageId="7" pageNumber="154">Kentucky</collectingRegion>
and
<collectingRegion box="[1099,1265,444,468]" country="United States of America" name="West Virginia" pageId="7" pageNumber="154">West Virginia</collectingRegion>
.
</paragraph>
<paragraph blockId="7.[835,1444,228,828]" pageId="7" pageNumber="154">
<bibRefCitation author="Grubbs, S. A. &amp; C. M. Thomas &amp; B. T. Hutchins &amp; J. M. Taylor" box="[873,1127,480,504]" pageId="7" pageNumber="154" pagination="321 - 332" refId="ref9225" refString="Grubbs, S. A., C. M. Thomas, B. T. Hutchins, &amp; J. M. Taylor. 2006. Life cycles of Leuctra spp. and Allocapnia recta (Plecoptera: Leuctridae and Capniidae) across a flow gradient in a headwater karst valley. Southeastern Naturalist, 5: 321 - 332. Available online: http: // www. bioone. org / doi / full / 10.1656 / 1528 - 7092 % 282006 % 295 % 5 B 321 % 3 ALCOARA % 5 D 2." type="journal article" year="2006">Grubbs et al. (2006)</bibRefCitation>
showed that
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[1307,1443,480,504]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[1307,1443,480,504]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[835,903,516,539]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
(as
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[957,1094,516,540]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis box="[957,972,517,540]" italics="true" pageId="7" pageNumber="154">L</emphasis>
. cf.
<emphasis box="[1027,1094,517,540]" italics="true" pageId="7" pageNumber="154">tenuis</emphasis>
</taxonomicName>
) exhibited a univoltine-slow life cycle. Nymphs were present by early January and appear to require continuous flow in the spring run during the dry summer months.
<taxonomicName box="[1361,1443,625,648]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis box="[1361,1443,625,648]" italics="true" pageId="7" pageNumber="154">Leuctra</emphasis>
</taxonomicName>
nymphs present through summer were presumably only
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[1051,1181,696,720]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="154" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[1051,1181,696,720]" italics="true" pageId="7" pageNumber="154">L. schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1192,1260,696,719]" pageId="7" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
since this was the only adult species collected from July through early October. The common name, Karst Needlefly, is proposed for this species (
<bibRefCitation author="Stark, B. P. &amp; K. W. Stewart &amp; S. W. Szczytko &amp; R. W. Baumann &amp; B. C. Kondratieff" box="[1168,1358,804,828]" pageId="7" pageNumber="154" pagination="1 - 20" refId="ref9825" refString="Stark, B. P., K. W. Stewart, S. W. Szczytko, R. W. Baumann, &amp; B. C. Kondratieff. 2012. Scientific and common names of stoneflies of Nearctic stoneflies (Plecoptera), with corrections and additions to the list. The Caddis Press, Miscellaneous Contributions, 1: 1 - 20." type="journal article" year="2012">Stark et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>