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<document id="313E63C34B8D5A381301213935E98F4A" ID-DOI="10.11646/zootaxa.3846.4.2" ID-GBIF-Dataset="ea136b1c-dc96-4c41-b078-01ca553453a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="252519" ID-ZooBank="71A80CC3-CDC1-4291-863B-FA7501D60AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467899762" checkinUser="plazi" docAuthor="Câmara, J. T. &amp; Rafael, J. A." docDate="2014" docId="03F6943FE079FF91C7C7FE36AD4B1604" docLanguage="en" docName="zt03846p546.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opeatocerata chaetohypandrialis Câmara &amp; Rafael, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="514" masterDocId="FFCFEC47E075FF9DC750FFEAA8211077" masterDocTitle="Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)" masterLastPageNumber="546" masterPageNumber="502" pageNumber="514" updateTime="1698305230595" updateUser="plazi">
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<mods:title id="205F30109AC7A68BF236BAC4298AEF36">Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)</mods:title>
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<mods:namePart id="625969E46FA1F61D86B9CE3941B50147">Câmara, J. T.</mods:namePart>
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<mods:namePart id="E738D8787566F8273E4FC73251FF41B7">Rafael, J. A.</mods:namePart>
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<treatment id="03F6943FE079FF91C7C7FE36AD4B1604" ID-DOI="http://doi.org/10.5281/zenodo.5285211" ID-GBIF-Taxon="119514690" ID-Zenodo-Dep="5285211" LSID="urn:lsid:plazi:treatment:03F6943FE079FF91C7C7FE36AD4B1604" httpUri="http://treatment.plazi.org/id/03F6943FE079FF91C7C7FE36AD4B1604" lastPageNumber="514" pageId="12" pageNumber="514">
<subSubSection id="C34576A2E079FF91C7C7FE36A9061260" pageId="12" pageNumber="514" type="nomenclature">
<paragraph id="8BE02529E079FF91C7C7FE36AAAB1181" blockId="12.[151,650,476,535]" box="[151,650,476,502]" pageId="12" pageNumber="514">
<heading id="D0A89245E079FF91C7C7FE36AAAB1181" bold="true" box="[151,650,476,502]" fontSize="11" level="1" pageId="12" pageNumber="514" reason="1">
<emphasis id="B92BF93BE079FF91C7C7FE36AAAB1181" bold="true" box="[151,650,476,502]" pageId="12" pageNumber="514">
<taxonomicName id="4C5F5EAAE079FF91C7C7FE36AA071181" ID-CoL="8D3LG" box="[151,550,476,502]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="514" phylum="Arthropoda" rank="species" species="chaetohypandrialis" status="sp. nov.">
<emphasis id="B92BF93BE079FF91C7C7FE36AA071181" bold="true" box="[151,550,476,502]" italics="true" pageId="12" pageNumber="514">Opeatocerata chaetohypandrialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E079FF91C57DFE36AAAB1181" box="[557,650,476,502]" pageId="12" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE02529E079FF91C7C7FE14A9061260" blockId="12.[151,650,476,535]" box="[151,295,510,535]" pageId="12" pageNumber="514">
(
<figureCitation id="136439ACE079FF91C7CFFE14A93E1260" box="[159,287,510,535]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Figs 4757</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E079FF91C7C7FDACAB6612D0" pageId="12" pageNumber="514" type="diagnosis">
<paragraph id="8BE02529E079FF91C7C7FDACAB6612D0" blockId="12.[151,1437,582,1651]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C7C7FDACA92B1228" bold="true" box="[151,266,582,607]" pageId="12" pageNumber="514">Diagnosis</emphasis>
. Tergites 36 with basal subrectangular and apical transversal brown spots (
<figureCitation id="136439ACE079FF91C327FDACACEC1228" box="[1143,1229,582,607]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 50</figureCitation>
); hypandium with long bristles at apex (
<figureCitation id="136439ACE079FF91C6D8FD80A9FC12F4" box="[392,477,618,643]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 57</figureCitation>
); phallus with large dorsal subapical appendix in lateral view (
<figureCitation id="136439ACE079FF91C3CEFD80ACD112F4" box="[1182,1264,618,643]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 55</figureCitation>
) and spiniform projections laterally, best visualized in ventral view (
<figureCitation id="136439ACE079FF91C5B6FD64AB1B12D0" box="[742,826,654,679]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 56</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E079FF91C797FD58AB391524" pageId="12" pageNumber="514" type="description">
<paragraph id="8BE02529E079FF91C797FD58ABE71558" blockId="12.[151,1437,582,1651]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FD58A97012BC" bold="true" box="[199,337,690,715]" pageId="12" pageNumber="514">Description</emphasis>
.
<emphasis id="B92BF93BE079FF91C630FD58A9BC12BC" bold="true" box="[352,413,690,715]" pageId="12" pageNumber="514">Male</emphasis>
(
<figureCitation id="136439ACE079FF91C6FDFD58AA2312BC" box="[429,514,690,715]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 47</figureCitation>
).
<emphasis id="B92BF93BE079FF91C546FD58AA7712BC" bold="true" box="[534,598,690,715]" pageId="12" pageNumber="514">Head</emphasis>
: Face parallel-sided, dark brown with gray pruinescence visible in ventral view, about
<date id="FFE103E9E079FF91C673FD3DA9671298" box="[291,326,727,751]" pageId="12" pageNumber="514">4X</date>
longer than lower width. Ocellar tubercle protuberant, dark brown with gray puinescence; ocelli yellow. Scape and pedicel yellow with yellow bristles; postpedicel brown about
<date id="FFE103E9E079FF91C36FFD11AC401364" box="[1087,1121,763,787]" pageId="12" pageNumber="514">2X</date>
longer than pedicel; stylus about
<date id="FFE103E9E079FF91C78CFCF5A8DF1340" box="[220,254,799,823]" pageId="12" pageNumber="514">2X</date>
longer than postpedicel. Proboscis yellow shorter than head height.
<emphasis id="B92BF93BE079FF91C4AAFCF4AC721340" bold="true" box="[1018,1107,798,823]" pageId="12" pageNumber="514">Thorax</emphasis>
yellow, shiny (
<figureCitation id="136439ACE079FF91C3AEFCF4AD731340" box="[1278,1362,798,823]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 47</figureCitation>
). Legs yellow, except for hind femur and tibia with black apical ring and all tarsomeres 45 black; hind tibia and tarsus with anterodorsal and posterodorsal rows of longer bristles.
<emphasis id="B92BF93BE079FF91C4D2FC8CABE21308" bold="true" box="[898,963,870,895]" pageId="12" pageNumber="514">Wing</emphasis>
(
<figureCitation id="136439ACE079FF91C48CFC8CAC171308" box="[988,1078,870,895]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 48</figureCitation>
) hyaline, pterostigma brown, conspicuous, about 2.5X longer than wide. Halter yellow.
<emphasis id="B92BF93BE079FF91C408FC60ABED13D4" bold="true" box="[856,972,906,931]" pageId="12" pageNumber="514">Abdomen</emphasis>
yellow, shiny; tergites 36 with basal subrectangular and apical transversal brown spots (
<figureCitation id="136439ACE079FF91C5B7FC44AB5113B0" box="[743,880,942,967]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Figs 49, 50</figureCitation>
); tergite 8 divided in two subrectangular plates; sternite 8 divided in two trapezoidal plates.
<emphasis id="B92BF93BE079FF91C52DFC38AB2B139C" bold="true" box="[637,778,978,1003]" pageId="12" pageNumber="514">Terminalia:</emphasis>
Anterior cercus with a reclinate dorsal projection in lateral view (
<figureCitation id="136439ACE079FF91C78CFC1CA97F1478" box="[220,350,1014,1039]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Figs 49, 51</figureCitation>
), subrectangular in dorsal view (
<figureCitation id="136439ACE079FF91C59DFC1CAB001478" box="[717,801,1014,1039]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 52</figureCitation>
); descendant plate subtrapezoidal in posterior view (
<figureCitation id="136439ACE079FF91C23FFC1CA8961444" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 53</figureCitation>
). Posterior cercus with rounded apex in lateral view (
<figureCitation id="136439ACE079FF91C414FBF0ABF21444" box="[836,979,1050,1075]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Figs 49, 51</figureCitation>
); without dorsal projections (
<figureCitation id="136439ACE079FF91C265FBF0ADB11444" box="[1333,1424,1050,1075]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 52</figureCitation>
). Hypoproct suboval in lateral view (
<figureCitation id="136439ACE079FF91C572FBD4AA521420" box="[546,627,1086,1111]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 51</figureCitation>
) and comma-shaped in posterior view (
<figureCitation id="136439ACE079FF91C37EFBD4ACA31420" box="[1070,1154,1086,1111]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 53</figureCitation>
). Epandrial with bilobate ventral lobe (
<figureCitation id="136439ACE079FF91C67FFB88A994140C" box="[303,437,1122,1147]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Figs 49, 51</figureCitation>
); spiniform bristles mesially on dorsal lobe and longer bristles apically on ventral lobe (
<figureCitation id="136439ACE079FF91C7CFFB6CA8D214E8" box="[159,243,1158,1183]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 51</figureCitation>
). Hypandrium longer than wide, with truncate apex and median sinus, with long bristles at apex (
<figureCitation id="136439ACE079FF91C26BFB6CADB114E8" box="[1339,1424,1158,1183]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 57</figureCitation>
). Phallus with narrower base, extending towards the apex, longer than hypandrium, with a larger dorsal subapical appendix in lateral view (
<figureCitation id="136439ACE079FF91C698FB24AA3F1490" box="[456,542,1230,1255]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 55</figureCitation>
) and spiniform projections laterally in ventral view (
<figureCitation id="136439ACE079FF91C3C7FB24ACD11490" box="[1175,1264,1230,1255]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 56</figureCitation>
). Subepandrial sclerite wider than long, V-shaped (
<figureCitation id="136439ACE079FF91C572FB18AA57157C" box="[546,630,1266,1291]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 54</figureCitation>
). Bacilliform sclerite without folds and longer than subepandrial sclerite (
<figureCitation id="136439ACE079FF91C7CFFAFCA8D51558" box="[159,244,1302,1327]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 54</figureCitation>
).
<emphasis id="B92BF93BE079FF91C657FAFCA9EA1558" bold="true" box="[263,459,1302,1327]" pageId="12" pageNumber="514">Specimen length</emphasis>
:
<quantity id="4CA788CCE079FF91C68AFAFCAA121559" box="[474,563,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="514" unit="mm" value="3.5">3.5 mm</quantity>
;
<emphasis id="B92BF93BE079FF91C56FFAFCAA591558" bold="true" box="[575,632,1302,1327]" pageId="12" pageNumber="514">wing</emphasis>
:
<quantity id="4CA788CCE079FF91C5D7FAFDAAC11559" box="[647,736,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="514" unit="mm" value="4.2">4.2 mm</quantity>
.
<emphasis id="B92BF93BE079FF91C5BBFAFCAB631558" bold="true" box="[747,834,1302,1327]" pageId="12" pageNumber="514">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BE02529E079FF91C797FAD0AB391524" blockId="12.[151,1437,582,1651]" box="[199,792,1338,1363]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FAD0A9DC1524" bold="true" box="[199,509,1338,1363]" pageId="12" pageNumber="514">Geographical distribution</emphasis>
.
<collectingCountry id="F34865B9E079FF91C55BFAD0AA6E1524" box="[523,591,1338,1363]" name="Brazil" pageId="12" pageNumber="514">Brazil</collectingCountry>
(Pará) (
<figureCitation id="136439ACE079FF91C5FAFAD0AB2D1524" box="[682,780,1338,1363]" captionStart="FIGURE 211" captionStartId="42.[151,250,1955,1977]" captionTargetBox="[235,1368,193,1934]" captionTargetId="figure@42.[216,1371,193,1934]" captionTargetPageId="42" captionText="FIGURE 211. Geographic distribution of species of Opeatocerata." httpUri="https://zenodo.org/record/252546/files/figure.png" pageId="12" pageNumber="514">Fig. 211</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E079FF91C797FAB4ACCE15C8" pageId="12" pageNumber="514" type="materials_examined">
<paragraph id="8BE02529E079FF91C797FAB4AB8E15EC" blockId="12.[151,1437,582,1651]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FAB4A9501500" bold="true" box="[199,369,1374,1399]" pageId="12" pageNumber="514">
<typeStatus id="54E49B8BE079FF91C797FAB4A9231500" box="[199,258,1374,1399]" pageId="12" pageNumber="514">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54E49B8BE079FF91C6D2FAB4AA011500" box="[386,544,1374,1399]" pageId="12" pageNumber="514" type="holotype">
<emphasis id="B92BF93BE079FF91C6D2FAB4AA011500" bold="true" box="[386,544,1374,1399]" pageId="12" pageNumber="514">HOLOTYPE</emphasis>
</typeStatus>
♂, labelled:
<collectingCountry id="F34865B9E079FF91C5EDFAB4AB221500" box="[701,771,1374,1399]" name="Brazil" pageId="12" pageNumber="514">Brasil</collectingCountry>
, Pará, Santo A[ntonio] de Tauá.
<date id="FFE103E9E079FF91C3D3FAB4ACDC1500" box="[1155,1277,1374,1399]" pageId="12" pageNumber="514" value="1975-04-07">07.iv.1975</date>
. I.S. Gorayeb (
<collectionCode id="ED4EBDECE079FF91C7CFFA69A8D215EC" box="[159,243,1411,1435]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="12" pageNumber="514">MPEG</collectionCode>
).
<typeStatus id="54E49B8BE079FF91C655FA68A95C15EC" box="[261,381,1410,1435]" pageId="12" pageNumber="514" type="paratype">
<emphasis id="B92BF93BE079FF91C655FA68A95C15EC" bold="true" box="[261,381,1410,1435]" pageId="12" pageNumber="514">Paratypes</emphasis>
</typeStatus>
: same data as
<typeStatus id="54E49B8BE079FF91C570FA68AAA315EC" box="[544,642,1410,1435]" pageId="12" pageNumber="514" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D59EEA0E079FF91C5C2FA69AA9F15ED" box="[658,702,1411,1435]" pageId="12" pageNumber="514" type="generic">2 ♂</specimenCount>
,
<collectionCode id="ED4EBDECE079FF91C59BFA69AB2C15EC" box="[715,781,1411,1435]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="12" pageNumber="514">INPA</collectionCode>
;
<specimenCount id="9D59EEA0E079FF91C448FA69AB6515ED" box="[792,836,1411,1435]" pageId="12" pageNumber="514" type="generic">1 ♂</specimenCount>
,
<collectionCode id="ED4EBDECE079FF91C400FA69AB8215EC" box="[848,931,1411,1435]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="12" pageNumber="514">MPEG</collectionCode>
).
</paragraph>
<paragraph id="8BE02529E079FF91C797FA4CACCE15C8" blockId="12.[151,1437,582,1651]" box="[199,1263,1446,1471]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FA4CA98A15C8" bold="true" box="[199,427,1446,1471]" pageId="12" pageNumber="514">
<typeStatus id="54E49B8BE079FF91C797FA4CA91515C8" box="[199,308,1446,1471]" pageId="12" pageNumber="514" type="holotype">Holotype</typeStatus>
condition
</emphasis>
. Good; wing mounted in microslide; abdomen in microtube with glycerin.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E079FF91C797FA20AD281594" box="[199,1289,1482,1507]" pageId="12" pageNumber="514" type="etymology">
<paragraph id="8BE02529E079FF91C797FA20AD281594" blockId="12.[151,1437,582,1651]" box="[199,1289,1482,1507]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FA20A9671594" bold="true" box="[199,326,1482,1507]" pageId="12" pageNumber="514">Etymology</emphasis>
. From the Greek
<emphasis id="B92BF93BE079FF91C55CFA20AA761594" box="[524,599,1482,1507]" italics="true" pageId="12" pageNumber="514">chaeto</emphasis>
(bristles), referring to the bristles present on the hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E079FF91C797FA04AD4B1604" pageId="12" pageNumber="514" type="discussion">
<paragraph id="8BE02529E079FF91C797FA04AD4B1604" blockId="12.[151,1437,582,1651]" pageId="12" pageNumber="514">
<emphasis id="B92BF93BE079FF91C797FA04A91A1670" bold="true" box="[199,315,1518,1543]" pageId="12" pageNumber="514">Remarks.</emphasis>
<taxonomicName id="4C5F5EAAE079FF91C614FA05AA9E1670" box="[324,703,1518,1543]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="514" phylum="Arthropoda" rank="species" species="chaetohypandrialis" status="sp. nov.">
<emphasis id="B92BF93BE079FF91C614FA05AA9E1670" box="[324,703,1518,1543]" italics="true" pageId="12" pageNumber="514">Opeatocerata chaetohypandrialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E079FF91C599FA04AB051670" box="[713,804,1518,1543]" pageId="12" pageNumber="514" rank="species">
<emphasis id="B92BF93BE079FF91C599FA04AB051670" bold="true" box="[713,804,1518,1543]" pageId="12" pageNumber="514">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other species by the following characters: tergites 36 with basal subrectangular and apical transversal brown spots (
<figureCitation id="136439ACE079FF91C48AF9F8AC0D165C" box="[986,1068,1554,1579]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 50</figureCitation>
) (other species have paramedian spots on the tergites); hypandrium with bristles at apex (
<figureCitation id="136439ACE079FF91C45BF9DCAB7D1638" box="[779,860,1590,1615]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 57</figureCitation>
) (other species without bristles on the hypandium); phallus with spiniform projections laterally (
<figureCitation id="136439ACE079FF91C5DAF9B0AAFA1604" box="[650,731,1626,1651]" captionStart="FIGURES 47 57" captionStartId="13.[151,264,1814,1837]" captionTargetBox="[151,1417,194,1792]" captionTargetId="figure@13.[151,1436,193,1794]" captionTargetPageId="13" captionText="FIGURES 47 57. Opeatocerata chaetohypandrialis sp. nov., holotype ♂. 47. habitus, lateral view; 48. wing; 49. abdomen from tergite 4 to terminalia, lateral view; 50. tergites 3 8, dorsal view; 51. terminalia, lateral view; 52. lobes of cerci, dorsal view; 53. descendant plate of anterior cercus and hypropoct, posterior view; 54. subepandrial sclerite and bacilliform sclerite, dorsal view; 55. ejaculatory apodeme, phallus and hypandrium, lateral view; 56. apex of phallus, ventral view; 57. hypandrium, ventral view." httpUri="https://zenodo.org/record/252526/files/figure.png" pageId="12" pageNumber="514">Fig. 56</figureCitation>
) (in other species without spiniform projections laterally).
</paragraph>
</subSubSection>
</treatment>
</document>