272 lines
57 KiB
XML
272 lines
57 KiB
XML
<document ID-DOI="10.35929/RSZ.0083" ID-ISSN="0035-418" ID-Zenodo-Dep="7761487" checkinTime="1679514280647" checkinUser="valdenar" docAuthor="Schwendinger, Peter J., Huber, Siegfried, Lehmann-Graber, Christina, Ono, Hirotsugu, Aung, Mu Mu & Hongpadharakiree, Komsan" docDate="2022" docId="03E18D7172400361B2C7FC505119FEC9" docLanguage="en" docName="RevSuisseZool.129.2.375-424.pdf" docOrigin="Revue suisse de Zoologie 129 (2)" docSource="http://dx.doi.org/10.35929/rsz.0083" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Liphistius lahu Schwendinger 1998" docType="treatment" docVersion="3" lastPageNumber="382" masterDocId="FFD8F50972460369B239FFEC5305FF92" masterDocTitle="A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species" masterLastPageNumber="424" masterPageNumber="375" pageNumber="380" updateTime="1679571001943" updateUser="valdenar">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Liphistius birmanicus-group (Araneae: Liphistiidae) with the description of five new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schwendinger, Peter J.</mods:namePart>
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<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Huber, Siegfried</mods:namePart>
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<mods:affiliation>Ottenbohlstrasse 12, D- 88690 Mühlhofen, Germany</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lehmann-Graber, Christina</mods:namePart>
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<mods:affiliation>Muséum d’histoire naturelle de Genève, C. P. 6404, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ono, Hirotsugu</mods:namePart>
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<mods:affiliation>Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aung, Mu Mu</mods:namePart>
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<mods:affiliation>Forest Research Institute, Forest Department, Ministry of Natural Resources and Environmental Conservation, Yezin, Nay Pyi Taw, Myanmar</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hongpadharakiree, Komsan</mods:namePart>
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<mods:affiliation>Sirinart Rajini Mangrove Ecosystem Learning Center, Pranburi, Prachuab Khiri Khan Province, Thailand</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Revue suisse de Zoologie</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-10-03</mods:number>
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<mods:detail type="volume">
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<mods:number>129</mods:number>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:start>375</mods:start>
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<mods:end>424</mods:end>
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<mods:url>http://dx.doi.org/10.35929/rsz.0083</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.35929/RSZ.0083</mods:identifier>
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<mods:identifier type="ISSN">0035-418</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7761487</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7761499" ID-Zenodo-Dep="7761499" LSID="urn:lsid:plazi:treatment:03E18D7172400361B2C7FC505119FEC9" httpUri="http://treatment.plazi.org/id/03E18D7172400361B2C7FC505119FEC9" lastPageId="8" lastPageNumber="382" pageId="6" pageNumber="380">
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<subSubSection box="[254,653,956,980]" pageId="6" pageNumber="380" type="nomenclature">
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<paragraph blockId="6.[254,653,956,1013]" box="[254,653,956,980]" pageId="6" pageNumber="380">
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<heading bold="true" box="[254,653,956,980]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="380" reason="2">
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<taxonomicName ID-CoL="72J5J" authority="Schwendinger, 1998" authorityName="Schwendinger" authorityYear="1998" box="[254,653,956,980]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="lahu">
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<emphasis bold="true" box="[254,653,956,980]" pageId="6" pageNumber="380">
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<emphasis bold="true" box="[254,420,956,980]" italics="true" pageId="6" pageNumber="380">Liphistius lahu</emphasis>
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<bibRefCitation author="Schwendinger P. J." box="[427,653,957,980]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[396,511,990,1013]" pageId="6" pageNumber="380" type="description">
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||
<paragraph blockId="6.[254,653,956,1013]" box="[396,511,990,1013]" pageId="6" pageNumber="380">
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<figureCitation box="[396,462,990,1013]" captionStart="Fig" captionStartId="7.[151,186,1431,1452]" captionTargetBox="[151,1436,257,1395]" captionTargetId="figure-270@7.[151,1436,257,1395]" captionTargetPageId="7" captionText="Fig. 1. Localities of species in the Liphistius birmanicus-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads." figureDoi="http://doi.org/10.5281/zenodo.7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="6" pageNumber="380">Figs 1</figureCitation>
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,
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<figureCitation box="[475,511,990,1013]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[151,186,1824,1845]" captionStartId-1="11.[150,185,1911,1932]" captionTargetBox-0="[155,1431,649,1777]" captionTargetBox-1="[154,1432,769,1861]" captionTargetId-0="figure-152@10.[151,1436,644,1782]" captionTargetId-1="figure-264@11.[151,1436,765,1865]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." captionText-1="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi-0="http://doi.org/10.5281/zenodo.7761497" figureDoi-1="http://doi.org/10.5281/zenodo.7761501" httpUri-0="https://zenodo.org/record/7761497/files/figure.png" httpUri-1="https://zenodo.org/record/7761501/files/figure.png" pageId="6" pageNumber="380">4-5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="380" type="reference_group">
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<paragraph blockId="6.[151,756,1037,1147]" pageId="6" pageNumber="380">
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<taxonomicName authority="Schwendinger, 1998: 17 - 19" authorityName="Schwendinger" authorityPageNumber="17 - 19" authorityYear="1998" box="[151,625,1037,1058]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="lahu">
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<emphasis box="[151,309,1037,1058]" italics="true" pageId="6" pageNumber="380">Liphistius lahu</emphasis>
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<bibRefCitation author="Schwendinger P. J." box="[329,625,1037,1058]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998: 17-19</bibRefCitation>
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</taxonomicName>
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, fig. 1A-H (description of males and females). –
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<bibRefCitation author="Schwendinger P. J." pageId="6" pageNumber="380" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
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: fig. 1A-H (reprint of illustrations in
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<bibRefCitation author="Schwendinger P. J." pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="380" type="materials_examined">
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<paragraph blockId="6.[151,756,1172,1293]" pageId="6" pageNumber="380">
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<materialsCitation pageId="6" pageNumber="380">
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<emphasis bold="true" box="[151,266,1172,1195]" pageId="6" pageNumber="380">Holotype:</emphasis>
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MHNG-ARTO-0024737; male; Thailand, Chiang Mai Province, Fang District, Doi Angkhang, 1540 m, 19°55’10”N, 99°02’55”E; 27.VIII.1990; leg. P.J. Schwendinger.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="6.[151,756,1318,1440]" pageId="6" pageNumber="380">
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<materialsCitation pageId="6" pageNumber="380">
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<emphasis bold="true" box="[151,275,1318,1341]" pageId="6" pageNumber="380">Paratypes:</emphasis>
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MHNG-ARTO-0024739; 1 male; collected together with the holotype. – MHNG-ARTO-0024740 to 24745; 6 female paratypes; collected at the type locality; 27.VIII.1990 and 25.IX.1986; leg. P.J. Schwendinger
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="6.[151,756,1465,1586]" pageId="6" pageNumber="380">
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<emphasis bold="true" box="[151,337,1465,1488]" pageId="6" pageNumber="380">Other material:</emphasis>
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<materialsCitation pageId="6" pageNumber="380">MHNG; 1 female; Thailand, Chiang Mai Province, Fang District, Doi Pha Luang, 1600 m, 20°02’37”N, 99°06’14”E; 3.XI.1990; leg. P.J. Schwendinger. No new material available.</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="380" type="diagnosis">
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<paragraph blockId="6.[151,756,1611,1996]" pageId="6" pageNumber="380">
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<emphasis bold="true" box="[151,271,1611,1634]" pageId="6" pageNumber="380">Diagnosis:</emphasis>
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Males distinguished by a moderately deep tibial apophysis (depth/length ratio ~ 1.4,
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<figureCitation box="[622,707,1644,1667]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4F</figureCitation>
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), by a widely rounded retrolateral-proximal heel on paracymbium (
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<figureCitation box="[281,358,1710,1733]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
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H-J), by an indistinctly elevated prolateral part of the distal contrategular edge (
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<figureCitation captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4E, G</figureCitation>
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), and by short para-embolic plate not separated by an invagination from retroventral edge of embolus complex (
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<figureCitation box="[261,328,1841,1864]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
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C-E). Females distinguished by ventral side of poreplate without anterolateral processes and by an indistinct step between anterior lobes and lateral poreplate margins; posterior stalk axe-blade-shaped, about half as long as poreplate (
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<figureCitation box="[491,556,1973,1996]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="6" pageNumber="380">Fig. 5</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="7" lastPageNumber="381" pageId="6" pageNumber="380" type="description">
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<paragraph blockId="6.[151,756,2021,2077]" pageId="6" pageNumber="380">
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<emphasis bold="true" box="[151,445,2021,2044]" pageId="6" pageNumber="380">Additions to description:</emphasis>
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Medium-sized spiders with brown colouration in both sexes and annulated legs
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</paragraph>
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<paragraph blockId="6.[831,1437,910,2077]" lastBlockId="7.[151,756,1554,2029]" lastPageId="7" lastPageNumber="381" pageId="6" pageNumber="380">
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and palps in females and juveniles (in large spiders less distinct than in small ones, annulations thus fading as females become older and larger). Males with scopula weak on tarsi I-II, slightly denser on tarsi III-IV, covering distal 4/5 of tarsus I and distal 5/6 of tarsi II-IV. Male palps with moderately deep tibial apophysis (depth/length ratio ~ 1.4), not set back from distal margin of tibia, carrying four long, pointed apical megaspines (
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||
<figureCitation box="[1053,1140,1164,1187]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4F</figureCitation>
|
||
;
|
||
<bibRefCitation author="Schwendinger P. J." box="[1157,1386,1164,1187]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
: fig. 1A-C;
|
||
<bibRefCitation author="Schwendinger P. J." box="[910,1139,1196,1219]" pageId="6" pageNumber="380" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
|
||
: fig. 1A-C); paracymbium short, with an almost flat distal surface and with a widely arched retrolateral-proximal heel (
|
||
<figureCitation box="[1310,1381,1259,1282]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
H-J); cumulus indistinct, carrying a group of 5-6 long, strong bristles (
|
||
<figureCitation box="[1015,1088,1323,1346]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
H-J;
|
||
<bibRefCitation author="Schwendinger P. J." box="[1149,1383,1323,1346]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
: fig. 1B-C;
|
||
<bibRefCitation author="Schwendinger P. J." box="[914,1148,1355,1378]" pageId="6" pageNumber="380" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
|
||
: fig. 1B-C); subtegulum without apophysis (
|
||
<figureCitation box="[1064,1158,1386,1409]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4E</figureCitation>
|
||
); proventral process of contrategulum conical, with narrowly rounded apex in dorsal view (
|
||
<figureCitation box="[978,1046,1450,1473]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
A-C); prolateral part of distal edge of contrategulum developed as a low (not elevated as in
|
||
<taxonomicName box="[862,991,1513,1536]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="metopiae" status="sp. nov.">
|
||
<emphasis box="[862,991,1513,1536]" italics="true" pageId="6" pageNumber="380">L. metopiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1000,1087,1513,1536]" pageId="6" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) keel (
|
||
<figureCitation box="[1165,1285,1513,1536]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4E, G</figureCitation>
|
||
); no wrinkles on dorsal side of contrategulum, a pronounced proximal ledge on its retrodorsal side (
|
||
<figureCitation box="[1171,1241,1577,1600]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
A-C); distal edge of contrategulum very wide, with narrowly rounded dorsal apex (
|
||
<figureCitation box="[994,1070,1641,1664]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
A-C;
|
||
<bibRefCitation author="Schwendinger P. J." box="[1142,1379,1641,1664]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
: fig. 1D-E;
|
||
<bibRefCitation author="Schwendinger P. J." box="[907,1135,1672,1695]" pageId="6" pageNumber="380" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
|
||
: fig. 1D-E); tegulum large, its distal margin not elevated, its proximal edge widely arched, coarsely serrate, bent and distinctly overhanging membranous area of contrategulum below it (
|
||
<figureCitation captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
C-D;
|
||
<bibRefCitation author="Schwendinger P. J." box="[910,1138,1799,1822]" pageId="6" pageNumber="380" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
: fig. 1A-B;
|
||
<bibRefCitation author="Schwendinger P. J." pageId="6" pageNumber="380" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
|
||
: fig. 1A-B); para-embolic plate very short, not separated from retroventral edge of embolus complex by an invagination (
|
||
<figureCitation box="[1075,1149,1895,1918]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="6" pageNumber="380">Fig. 4</figureCitation>
|
||
C-E); sclerotised part of embolus proper strengthened by 3-4 distally dentate longitudinal ribs reaching apex, narrowly divided from distinctly shorter membranous embolus part; at base of membranous embolus part a weakly pigmented area with only 3-4 longitudinal wrinkles and with a wide and asymmetrical distal margin (
|
||
<figureCitation box="[480,551,1554,1577]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="7" pageNumber="381">Fig. 4</figureCitation>
|
||
C-D, G). Females with more or less distinctly annulated legs and palps; uterus externus with a small pair of lateral pockets (as illustrated for
|
||
<taxonomicName box="[306,390,1651,1674]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
|
||
<emphasis box="[306,390,1651,1674]" italics="true" pageId="7" pageNumber="381">L. ferox</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[398,485,1651,1674]" pageId="7" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation box="[497,572,1651,1674]" captionStart="Fig" captionStartId="21.[153,188,1845,1866]" captionTargetId="figure-7@20.[151,1439,257,2021]" captionTargetPageId="20" captionText="Fig. 11. Liphistius ferox, vulvae of seven females; dorsal view (A, C, E, G, I, K, N, P), ventral view (B, D, F, H, J, L-M, O). (A-B) Thandaung Gyi, moulted 21.VII.2015; with uterus externus attached (in B hidden on other side). (C-D) Thandaung Gyi, moulted 8.1.2015. (E-F) Thandaung Gyi, moulted 15.VII.2015; with uterus externus attached (in F only pocket of one side visible). (G-H) Thandaung Gyi, moulted 16.VII.2015; vulval plate. (I-J) Same female; isolated uterus externus. (K-L) Naw Bubaw, moulted 26.II.2015. (M) Allotype; Thandaung Lay, moulted 10.VII.2015, vulval plate. (N) Same female; isolated uterus externus. (O) Thandaung Gyi, moulted 20.IX.2014; vulval plate. (P) Same female; isolated uterus externus. Arrows indicating paired lateral pockets of still attached membranous uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761521" httpUri="https://zenodo.org/record/7761521/files/figure.png" pageId="7" pageNumber="381">Fig. 11</figureCitation>
|
||
I-J, N, P and
|
||
<figureCitation captionStart="Fig" captionStartId="21.[151,186,1651,1672]" captionTargetBox="[156,1436,901,1604]" captionTargetId="figure-411@21.[151,1439,879,1625]" captionTargetPageId="21" captionText="Fig. 12. Liphistius ferox, vulvae of three females; dorsal view (A, D, F), ventral view (B-C, E). (A-B) Thandaung Gyi, moulted 26.X.2014; vulval plate. (C) Thandaung Gyi, moulted 11.1.2015; vulval plate. (D) Same female; isolated uterus externus. (E) Thandaung Gyi, moulted 6.XII.2014; vulval plate. (F) Same female; isolated uterus externus. Scale lines 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7761519" httpUri="https://zenodo.org/record/7761519/files/figure.png" pageId="7" pageNumber="381">Fig 12D, F</figureCitation>
|
||
); vulval plate with several hairs on lateral folds; poreplate wider than long, with a pair of pronounced, rounded lobes on anterior margin, without anterolateral processes; an indistinct step between anterior lobes and lateral poreplate margins (
|
||
<figureCitation box="[450,518,1813,1836]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="7" pageNumber="381">Fig. 5</figureCitation>
|
||
E-F, H); receptacular cluster racemose, quite long, almost reaching or slightly surpassing anterior margin of poreplate (
|
||
<figureCitation box="[683,750,1877,1900]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="7" pageNumber="381">Fig. 5</figureCitation>
|
||
;
|
||
<bibRefCitation author="Schwendinger P. J." box="[151,382,1909,1932]" pageId="7" pageNumber="381" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998</bibRefCitation>
|
||
: fig. 1F-H;
|
||
<bibRefCitation author="Schwendinger P. J." box="[522,753,1909,1932]" pageId="7" pageNumber="381" pagination="371 - 377" refId="ref38160" refString="Schwendinger P. J. 1999. Erratum: Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 28 (3 - 4): 371 - 377." type="journal article" year="1999">Schwendinger, 1999</bibRefCitation>
|
||
: fig. 1F-H); posterior stalk axe-blade-shaped, its anterior portion narrow, its posterior margin wide and arched, narrower than poreplate.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7761489" ID-Zenodo-Dep="7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="7" pageNumber="381" startId="7.[151,186,1431,1452]" targetBox="[151,1436,257,1395]" targetPageId="7">
|
||
<paragraph blockId="7.[151,1436,1431,1481]" pageId="7" pageNumber="381">
|
||
Fig. 1. Localities of species in the
|
||
<taxonomicName authorityName="Thorell" authorityYear="1897" box="[485,694,1431,1452]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis box="[485,694,1431,1452]" italics="true" pageId="7" pageNumber="381">Liphistius birmanicus</emphasis>
|
||
</taxonomicName>
|
||
-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[151,756,2054,2077]" box="[151,756,2054,2077]" pageId="7" pageNumber="381">
|
||
<emphasis bold="true" box="[151,269,2054,2077]" pageId="7" pageNumber="381">Variation:</emphasis>
|
||
For carapace measurements and prefoveal
|
||
</paragraph>
|
||
<paragraph blockId="7.[831,1437,1554,1745]" pageId="7" pageNumber="381">
|
||
setae counts see
|
||
<tableCitation box="[1015,1097,1554,1577]" captionStart="Table 1" captionStartId="46.[151,204,1454,1475]" captionTargetId="graphics-627@46.[151,1436,1545,2074]" captionTargetPageId="46" captionText="Table 1. Carapace measurements (in mm; including data from the literature) and prefoveal setae counts of Liphistius birmanicus group species." httpUri="http://table.plazi.org/id/DF376CEF72680347B2AEFA425282FA71" pageId="7" pageNumber="381" tableUuid="DF376CEF72680347B2AEFA425282FA71">Table 1</tableCitation>
|
||
. All specimens examined have well-developed AME. Variation in the shape of the paracymbium and of the distal edge plus the proventral process of the contrategulum of males is shown in
|
||
<figureCitation captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="7" pageNumber="381">Fig. 4</figureCitation>
|
||
H-J and
|
||
<figureCitation box="[937,1000,1688,1711]" captionStart="Fig" captionStartId="10.[151,186,1824,1845]" captionTargetBox="[155,1431,649,1777]" captionTargetId="figure-152@10.[151,1436,644,1782]" captionTargetPageId="10" captionText="Fig. 4. Liphistius lahu, palp of male holotype (B-I) and of male paratype (A, J). (A-B) Contrategulum of right palp, distal view (dorsal side up). (C) Left palpal organ, distal and slightly retrodorsal view (dorsal side up). (D) Distal part of left palpal organ, retroventral and slightly distal view; bent proximal edge of tegulum indicated by arrow. (E) Distal part of left palp, vental and slightly proximal view. (F) Distal part of left tibia, retrolateral and slightly proximal view. (G) Palpal organ and distal margin of tibia of left palp, prolateral and slightly proximal view. (H) Proximal part of tarsus of right palp, ventral view. (I-J) Same of left palp. Scale lines 1.0 mm (A-G; H-J). Abbreviations: c - cumulus; cp - proventral contrategular process; da - dorsal apex of distal contrategular edge; de - prolateral part of distal edge of contrategulum; me - membranous part of embolus proper; pc - paracymbium; pl - proximal ledge on retrodorsal side of contrategulum; pp - para-embolic plate; pt - palpal tarsus; re - retroventral edge of embolus complex; se - sclerotised part of embolus proper; st - subtegulum; t - tegulum; ta - tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7761497" httpUri="https://zenodo.org/record/7761497/files/figure.png" pageId="7" pageNumber="381">Fig. 4</figureCitation>
|
||
A-C, respectively. Variation in the shape of the vulval plates of five females is shown in
|
||
<figureCitation box="[1335,1400,1722,1745]" captionStart="Fig" captionStartId="11.[150,185,1911,1932]" captionTargetBox="[154,1432,769,1861]" captionTargetId="figure-264@11.[151,1436,765,1865]" captionTargetPageId="11" captionText="Fig. 5. Liphistius lahu, vulval plates of five females; dorsal view (A, D, G), ventral view (B, E-F, H-I) and anteroventral view (C). (A-C) Paratype MHNG-ARTO-0024744 (taken from specimen). (D-E) Paratype MHNG-ARTO-0024741 (from exuvia). (F) Paratype MHNG-ARTO-0024745 (from specimen). (G-H) Paratype MHNG-ARTO-0024742 (from exuvia). (I) Female from Doi Pa Luang (from specimen). Scale lines 1.0 mm.Arrows indicating indistinct step between anterior lobes and lateral margin of poreplate.Abbreviations: al - anterior lobe of poreplate; CDO - central dorsal opening of poreplate; ga - genital atrium; lf - lateral fold of vulval plate; ps - posterior stalk; rc - receptacular cluster." figureDoi="http://doi.org/10.5281/zenodo.7761501" httpUri="https://zenodo.org/record/7761501/files/figure.png" pageId="7" pageNumber="381">Fig. 5</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[831,1436,1770,1894]" pageId="7" pageNumber="381">
|
||
<emphasis bold="true" box="[831,998,1770,1793]" pageId="7" pageNumber="381">Relationships:</emphasis>
|
||
Palp morphology of males is very similar in
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[947,1027,1804,1827]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[947,1027,1804,1827]" italics="true" pageId="7" pageNumber="381">L. lahu</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[1086,1216,1804,1827]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="metopiae" status="sp. nov.">
|
||
<emphasis box="[1086,1216,1804,1827]" italics="true" pageId="7" pageNumber="381">L. metopiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1226,1315,1804,1827]" pageId="7" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, indicating that these two species are not only geographically but also phylogenetically close to each other.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="381" type="distribution">
|
||
<paragraph blockId="7.[831,1436,1920,2077]" pageId="7" pageNumber="381">
|
||
<emphasis bold="true" box="[831,983,1920,1943]" pageId="7" pageNumber="381">Distribution:</emphasis>
|
||
<taxonomicName authorityName="Schwendinger" authorityYear="1998" box="[1001,1176,1920,1943]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="lahu">
|
||
<emphasis box="[1001,1176,1920,1943]" italics="true" pageId="7" pageNumber="381">Liphistius lahu</emphasis>
|
||
</taxonomicName>
|
||
is known from two localities in the mountains of the northern Thai province of Chiang Mai, at and close to the border with the Shan State of Myanmar (
|
||
<figureCitation box="[1040,1104,2020,2043]" captionStart="Fig" captionStartId="7.[151,186,1431,1452]" captionTargetBox="[151,1436,257,1395]" captionTargetId="figure-270@7.[151,1436,257,1395]" captionTargetPageId="7" captionText="Fig. 1. Localities of species in the Liphistius birmanicus-group in northern Thailand, eastern Myanmar and southern China. Blue lines indicate major rivers, black lines national borders, grey lines major roads." figureDoi="http://doi.org/10.5281/zenodo.7761489" httpUri="https://zenodo.org/record/7761489/files/figure.png" pageId="7" pageNumber="381">Fig. 1</figureCitation>
|
||
). This species can probably be found on both sides of that border.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="382" type="biology_ecology">
|
||
<paragraph blockId="8.[151,756,258,347]" pageId="8" pageNumber="382">
|
||
<emphasis bold="true" box="[151,246,258,281]" pageId="8" pageNumber="382">Biology:</emphasis>
|
||
Information is given in the original description (
|
||
<bibRefCitation author="Schwendinger P. J." box="[158,432,291,314]" pageId="8" pageNumber="382" pagination="17 - 30" refId="ref38130" refString="Schwendinger P. J. 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30." type="journal article" year="1998">Schwendinger, 1998: 19</bibRefCitation>
|
||
). No new specimens or new biological information are available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |