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<document id="A31076D0EE82ED6C66828B5E6B965BCE" ID-DOI="10.5281/zenodo.186931" ID-GBIF-Dataset="6c338ef0-b784-415e-9451-4ee5547a85dd" ID-ISSN="1175-5326" ID-Zenodo-Dep="186931" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059447910" checkinUser="plazi" docAuthor="Korsós, Zoltán &amp; Johns, Peter M." docDate="2009" docId="03D8DB743E14FFA8FF614E9FB490FD35" docLanguage="en" docName="zt02065p024.pdf" docOrigin="Zootaxa 2065" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eumastigonus Chamberlin 1920" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FFE1A30C3E17FFAFFFF64807B04EFFB1" masterDocTitle="Introduction to the taxonomy of Iulomorphidae of New Zealand, with descriptions of two new species of Eumastigonus Chamberlin, 1920 (Diplopoda: Spirostreptida: Epinannolenidea)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="4" updateTime="1698581847651" updateUser="plazi">
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<mods:title id="D121D2CA808843714478E0A6F0BEED21">Introduction to the taxonomy of Iulomorphidae of New Zealand, with descriptions of two new species of Eumastigonus Chamberlin, 1920 (Diplopoda: Spirostreptida: Epinannolenidea)</mods:title>
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<mods:namePart id="8B6892BB3818D7039DA4BCCBB3193034">Korsós, Zoltán</mods:namePart>
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<mods:date id="605AA8EDE5E889D38AAB83DC97DA2818">2009</mods:date>
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<treatment id="03D8DB743E14FFA8FF614E9FB490FD35" ID-DOI="http://doi.org/10.5281/zenodo.6225520" ID-GBIF-Taxon="119613815" ID-Zenodo-Dep="6225520" LSID="urn:lsid:plazi:treatment:03D8DB743E14FFA8FF614E9FB490FD35" httpUri="http://treatment.plazi.org/id/03D8DB743E14FFA8FF614E9FB490FD35" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
<subSubSection id="C36B39E93E14FFACFF614E9FB2CAF903" box="[151,644,1688,1714]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BCE6A623E14FFACFF614E9FB2CAF903" blockId="3.[151,644,1688,1714]" box="[151,644,1688,1714]" pageId="3" pageNumber="4">
<heading id="D086DD0E3E14FFACFF614E9FB2CAF903" bold="true" box="[151,644,1688,1714]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B905B6703E14FFACFF614E9FB2CAF903" bold="true" box="[151,644,1688,1714]" pageId="3" pageNumber="4">
Genus
<taxonomicName id="4C7111E13E14FFACFF184E9FB2CAF903" ID-CoL="7PF6X" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" box="[238,644,1688,1714]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B905B6703E14FFACFF184E9FB1D2F903" bold="true" box="[238,412,1688,1714]" italics="true" pageId="3" pageNumber="4">Eumastigonus</emphasis>
<bibRefCitation id="EFE017933E14FFACFE524E9FB2CAF903" author="Chamberlin" box="[420,644,1688,1714]" pageId="3" pageNumber="23" refString="Chamberlin, R. V. (1920) The Myriapoda of the Australian region. Bulletin of the Museum of Comparitive Zoology, 64 (1), 1 - 269." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="8BCE6A623E14FFACFF614EE5B0B8F8A9" blockId="3.[151,1437,1761,1881]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB614D4C3E14FFACFF614EE5B0B8F8A9" pageId="3" pageNumber="4">
<taxonomicName id="4C7111E13E14FFACFF614EE5B251F949" ID-CoL="7PF6X" authority="Chamberlin, 1920: 62" authorityName="Chamberlin" authorityPageNumber="62" authorityYear="1920" box="[151,543,1761,1785]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B905B6703E14FFACFF614EE5B160F948" box="[151,302,1762,1785]" italics="true" pageId="3" pageNumber="4">Eumastigonus</emphasis>
<treatmentCitation id="0AD04C733E14FFACFEC24EE6B251F949" author="Chamberlin" box="[308,543,1761,1784]" page="62" pageId="3" pageNumber="4" year="1920">
<bibRefCitation id="EFE017933E14FFACFEC24EE6B1B6F949" author="Chamberlin" box="[308,504,1761,1784]" pageId="3" pageNumber="23" refString="Chamberlin, R. V. (1920) The Myriapoda of the Australian region. Bulletin of the Museum of Comparitive Zoology, 64 (1), 1 - 269." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
: 62
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFE017933E14FFACFDDC4EE6B2E2F949" author="Jeekel" box="[554,684,1761,1784]" lastPageId="3" lastPageNumber="24" pageId="3" pageNumber="23" refString="Jeekel, C. A. W. (1970) Nomenclator generum et familiarum Diplopodorum. Monografie No. 5, Nederlandse Entomologische Vereniging, Amsterdam, 142 pp." type="book" year="1970">Jeekel 1970</bibRefCitation>
: 09;
<treatmentCitation id="0AD04C733E14FFACFD2B4EE6B3FFF949" author="Hoffman" box="[733,945,1761,1784]" page="200" pageId="3" pageNumber="4" year="1972">
<bibRefCitation id="EFE017933E14FFACFD2B4EE6B332F949" author="Hoffman" box="[733,892,1761,1784]" pageId="3" pageNumber="23" refString="Hoffman, R. L. (1972) On the identity of three genera of Cambaloid millipeds from the Australian region (Spirostreptida). Psyche, Cambridge, USA 79 (3), 200 - 208." type="journal article" year="1972">Hoffman 1972</bibRefCitation>
: 200
</treatmentCitation>
;
<treatmentCitation id="0AD04C733E14FFACFC4D4EE6B4C9F949" author="Mauries" box="[955,1159,1761,1784]" page="256" pageId="3" pageNumber="4" year="1983">
<bibRefCitation id="EFE017933E14FFACFC4D4EE6B41CF949" author="Mauries" box="[955,1106,1761,1784]" pageId="3" pageNumber="24" refString="Mauries, J. - P. (1983) Cambalides nouveaux et peu connus d'Asie, d'Amerique et d'Oceanie. I. Cambalidae et Cambalopsidae (Myriapoda: Diplopoda). Bulletin du Museum national d'Histoire naturelle, serie 4, 5 A (1), 247 - 276." type="journal article" year="1983">Mauriès 1983</bibRefCitation>
: 256
</treatmentCitation>
; 1987: 198;
<treatmentCitation id="0AD04C733E14FFACFAFF4EE6B0BCF8A9" author="Shelley" page="196" pageId="3" pageNumber="4" year="2003">
<bibRefCitation id="EFE017933E14FFACFAFF4EE6B5D6F949" author="Shelley" box="[1289,1432,1761,1784]" pageId="3" pageNumber="24" refString="Shelley, R. M. (2003) A revised, annotated family-level classification of the Diplopoda. Arthropoda Selecta 11 (3), 187 - 207." type="journal article" year="2003">Shelley 2003</bibRefCitation>
: 196
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCE6A623E14FFACFF614F25B346F889" blockId="3.[151,1437,1761,1881]" box="[151,776,1825,1849]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB614D4C3E14FFACFF614F25B346F889" box="[151,776,1825,1849]" pageId="3" pageNumber="4">
<taxonomicName id="4C7111E13E14FFACFF614F25B1F3F889" ID-CoL="7PB2Y" authority="Attems, 1903" authorityName="Attems" authorityYear="1903" box="[151,445,1825,1849]" class="Diplopoda" family="Cambalidae" genus="Dimerogonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B905B6703E14FFACFF614F25B169F888" box="[151,295,1826,1849]" italics="true" pageId="3" pageNumber="4">Dimerogonus</emphasis>
<bibRefCitation id="EFE017933E14FFACFEDB4F26B1F3F889" author="Attems" box="[301,445,1825,1848]" pageId="3" pageNumber="23" refString="Attems, C. (1903) Beitrage zur Myriopodenkunde. II. Reisebeute der Herren Dr. Michaelsen, Dr. Plate, Dr. Burger und Dr. Schauinsland. Zoologischer Jahrbucher, Abteilung fur Systematik 18 (1), 79 - 92." type="journal article" year="1903">Attems, 1903</bibRefCitation>
</taxonomicName>
partim.
<treatmentCitation id="0AD04C733E14FFACFDE14F26B295F889" author="Verhoeff" box="[535,731,1825,1848]" page="74" pageId="3" pageNumber="4" year="1924">
<bibRefCitation id="EFE017933E14FFACFDE14F26B2FDF889" author="Verhoeff" box="[535,691,1825,1848]" pageId="3" pageNumber="24" refString="Verhoeff, K. W. (1924) Results of Dr E. Mjoberg's Swedish Scientific Expeditions to Australia, 1910 - 1913. 34. Myriapoda: Diplopoda. Arkiv for Zoologi 16 (5), 1 - 142." type="journal article" year="1924">Verhoeff 1924</bibRefCitation>
: 74
</treatmentCitation>
, 82.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCE6A623E14FFACFF614F45B2A5F8E9" blockId="3.[151,1437,1761,1881]" box="[151,747,1857,1881]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB614D4C3E14FFACFF614F45B2A5F8E9" box="[151,747,1857,1881]" pageId="3" pageNumber="4">
<taxonomicName id="4C7111E13E14FFACFF614F45B248F8E9" ID-CoL="0a94a338-6cdf-4aa6-b2f2-ae2b7a10d9c2" authority="Verhoeff, 1944: 31" authorityName="Verhoeff" authorityPageNumber="31" authorityYear="1944" box="[151,518,1857,1881]" class="Diplopoda" family="Cambalidae" genus="Insulocambala" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B905B6703E14FFACFF614F45B178F8E8" box="[151,310,1858,1881]" italics="true" pageId="3" pageNumber="4">Insulocambala</emphasis>
<treatmentCitation id="0AD04C733E14FFACFECA4F46B248F8E9" author="Verhoeff" box="[316,518,1857,1880]" page="31" pageId="3" pageNumber="4" year="1944">
<bibRefCitation id="EFE017933E14FFACFECA4F46B191F8E9" author="Verhoeff" box="[316,479,1857,1880]" pageId="3" pageNumber="24" refString="Verhoeff, K. W. (1944) Zur Kenntnis der Cambaliden und uber einige neue australische Formen derselben. Zoologischer Anzeiger 145, 27 - 45." type="journal article" year="1944">Verhoeff, 1944</bibRefCitation>
: 31
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AD04C733E14FFACFDE44F46B2A9F8E9" author="Hoffman" box="[530,743,1857,1880]" page="200" pageId="3" pageNumber="4" year="1972">
<bibRefCitation id="EFE017933E14FFACFDE44F46B2FFF8E9" author="Hoffman" box="[530,689,1857,1880]" pageId="3" pageNumber="23" refString="Hoffman, R. L. (1972) On the identity of three genera of Cambaloid millipeds from the Australian region (Spirostreptida). Psyche, Cambridge, USA 79 (3), 200 - 208." type="journal article" year="1972">Hoffman 1972</bibRefCitation>
: 200
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36B39E93E14FFACFF614F8FB3E9F85E" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BCE6A623E14FFACFF614F8FB3E9F85E" blockId="3.[151,1436,1928,2031]" pageId="3" pageNumber="4">
<emphasis id="B905B6703E14FFACFF614F8FB154F813" bold="true" box="[151,282,1928,1954]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
An epinannolenidean genus with modified first male legpair, and with well-developed flagellum in anterior male gonopods. Anterior gonopods with strong, hairy or pectinate coxal and telopodal processes which form sheath around straight and slender posterior gonopods.
</paragraph>
</subSubSection>
<caption id="DF0E3AEA3E13FFABFF614FDCB4D1F843" box="[151,1183,2011,2035]" httpUri="https://zenodo.org/record/186932/files/figure.png" pageId="4" pageNumber="5" targetBox="[177,1415,218,1972]" targetPageId="4">
<paragraph id="8BCE6A623E13FFABFF614FDCB4D1F843" blockId="4.[151,1183,2011,2035]" box="[151,1183,2011,2035]" pageId="4" pageNumber="5">
<emphasis id="B905B6703E13FFABFF614FDCB157F842" bold="true" box="[151,281,2011,2035]" pageId="4" pageNumber="5">FIGURE 1.</emphasis>
Position of
<taxonomicName id="4C7111E13E13FFABFE6D4FDCB276F843" box="[411,568,2011,2034]" class="Diplopoda" family="Iulomorphidae" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Iulomorphidae</taxonomicName>
in the order
<taxonomicName id="4C7111E13E13FFABFD364FDCB318F843" box="[704,854,2011,2034]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
according to different authors.
</paragraph>
</caption>
<caption id="DF0E3AEA3E12FFAAFF614F03B0BEF88F" pageId="5" pageNumber="6">
<paragraph id="8BCE6A623E12FFAAFF614F03B0BEF88F" blockId="5.[151,1436,1796,2027]" pageId="5" pageNumber="6">
<emphasis id="B905B6703E12FFAAFF614F03B10BF8AD" bold="true" box="[151,325,1796,1820]" pageId="5" pageNumber="6">FIGURES 23.</emphasis>
<taxonomicName id="4C7111E13E12FFAAFEBA4F03B203F8AA" box="[332,589,1796,1819]" class="Diplopoda" family="Cambalidae" genus="Dimerogonus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="orophilus">
<emphasis id="B905B6703E12FFAAFEBA4F03B203F8AA" box="[332,589,1796,1819]" italics="true" pageId="5" pageNumber="6">Dimerogonus orophilus</emphasis>
</taxonomicName>
: male first legpair. 2: Anterior view (hf= hairy field), 3: posterior view. Scale 0.5 mm.
</paragraph>
</caption>
<caption id="DF0E3AEA3E12FFAAFF614F4EB086F85A" httpUri="https://zenodo.org/record/186933/files/figure.png" pageId="5" pageNumber="6" targetBox="[311,1272,202,1748]" targetPageId="5">
<paragraph id="8BCE6A623E12FFAAFF614F4EB086F85A" blockId="5.[151,1436,1796,2027]" pageId="5" pageNumber="6">
<emphasis id="B905B6703E12FFAAFF614F4EB10AF8D0" bold="true" box="[151,324,1865,1889]" pageId="5" pageNumber="6">FIGURES 49.</emphasis>
<taxonomicName id="4C7111E13E12FFAAFEBA4F4DB204F8D0" box="[332,586,1866,1889]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kaorinus">
<emphasis id="B905B6703E12FFAAFEBA4F4DB204F8D0" box="[332,586,1866,1889]" italics="true" pageId="5" pageNumber="6">Eumastigonus kaorinus</emphasis>
</taxonomicName>
. 4: Gnathochilarium of paratype female (MCZ 4868), ventral view, 5: head of holotype male (MCZ 4867), left lateral view, 6: first male legpair, posterior view, 7: anterior gonopods, anterior view (
<emphasis id="B905B6703E12FFAAFA774F6BB5C3F832" box="[1409,1421,1900,1923]" italics="true" pageId="5" pageNumber="6">c</emphasis>
= coxa,
<emphasis id="B905B6703E12FFAAFF204F88B0A7F817" box="[214,233,1935,1958]" italics="true" pageId="5" pageNumber="6">m</emphasis>
= median lobe), 8: right gonopods, right lateral view, posterior gonopod removed from embrace of anterior ones, flagellum omitted (
<emphasis id="B905B6703E12FFAAFE934FB5B134F878" box="[357,378,1970,1993]" italics="true" pageId="5" pageNumber="6">tp</emphasis>
= telopodite process), 9: posterior gonopods, posterior view (
<emphasis id="B905B6703E12FFAAFBFC4FB5B459F878" box="[1034,1047,1970,1993]" italics="true" pageId="5" pageNumber="6">a</emphasis>
= apex). Scales 45: 1 mm, 69: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C36B39E93E11FFA9FF304890B4BAFF4E" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BCE6A623E11FFA9FF304890B430FF03" blockId="6.[151,1438,151,2010]" box="[198,1150,151,178]" pageId="6" pageNumber="7">
<emphasis id="B905B6703E11FFA9FF304890B125FF00" bold="true" box="[198,363,151,177]" pageId="6" pageNumber="7">
<typeStatus id="54CAD4C03E11FFA9FF304890B14DFF00" box="[198,259,151,177]" pageId="6" pageNumber="7">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7111E13E11FFA9FE84489FB32FFF03" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" box="[370,865,152,178]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kaorinus">
<emphasis id="B905B6703E11FFA9FE84489FB2C8FF00" box="[370,646,152,177]" italics="true" pageId="6" pageNumber="7">Eumastigonus kaorinus</emphasis>
<bibRefCitation id="EFE017933E11FFA9FD7B489FB32FFF03" author="Chamberlin" box="[653,865,152,178]" pageId="6" pageNumber="23" refString="Chamberlin, R. V. (1920) The Myriapoda of the Australian region. Bulletin of the Museum of Comparitive Zoology, 64 (1), 1 - 269." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
<paragraph id="8BCE6A623E11FFA9FF3048B9B4BAFF4E" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">
Other included species:
<taxonomicName id="4C7111E13E11FFA9FE1A48B8B30AFF69" authority="Attems, 1903" authorityName="Attems" authorityYear="1903" box="[492,836,190,216]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B905B6703E11FFA9FE1A48B8B2CEFF69" box="[492,640,191,216]" italics="true" pageId="6" pageNumber="7">E. insulanus</emphasis>
(
<bibRefCitation id="EFE017933E11FFA9FD6048B9B375FF69" author="Attems" box="[662,827,190,216]" pageId="6" pageNumber="23" refString="Attems, C. (1903) Beitrage zur Myriopodenkunde. II. Reisebeute der Herren Dr. Michaelsen, Dr. Plate, Dr. Burger und Dr. Schauinsland. Zoologischer Jahrbucher, Abteilung fur Systematik 18 (1), 79 - 92." type="journal article" year="1903">Attems, 1903</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C7111E13E11FFA9FCA048B8B3E6FF69" box="[854,936,191,216]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B905B6703E11FFA9FCA048B8B3E6FF69" box="[854,936,191,216]" italics="true" pageId="6" pageNumber="7">E. ater</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7111E13E11FFA9FC4B48B8B419FF69" box="[957,1111,191,216]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="distinctior">
<emphasis id="B905B6703E11FFA9FC4B48B8B419FF69" box="[957,1111,191,216]" italics="true" pageId="6" pageNumber="7">E. distinctior</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7111E13E11FFA9FB9A48B8B497FF69" box="[1132,1241,191,216]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maior">
<emphasis id="B905B6703E11FFA9FB9A48B8B497FF69" box="[1132,1241,191,216]" italics="true" pageId="6" pageNumber="7">E. maior</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7111E13E11FFA9FB1948B8B528FF69" box="[1263,1382,191,216]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B905B6703E11FFA9FB1948B8B528FF69" box="[1263,1382,191,216]" italics="true" pageId="6" pageNumber="7">E. parvus</emphasis>
</taxonomicName>
(all
<bibRefCitation id="EFE017933E11FFA9FF6148E2B121FF4E" author="Chamberlin" box="[151,367,229,255]" pageId="6" pageNumber="23" refString="Chamberlin, R. V. (1920) The Myriapoda of the Australian region. Bulletin of the Museum of Comparitive Zoology, 64 (1), 1 - 269." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
),
<taxonomicName id="4C7111E13E11FFA9FE7548E2B2BEFF4E" authority="Mauries, 1983" authorityName="Mauries" authorityYear="1983" box="[387,752,229,255]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hemmingseni">
<emphasis id="B905B6703E11FFA9FE7548E2B270FF4F" box="[387,574,229,254]" italics="true" pageId="6" pageNumber="7">E. hemmingseni</emphasis>
<bibRefCitation id="EFE017933E11FFA9FDB348E2B2BEFF4E" author="Mauries" box="[581,752,229,255]" pageId="6" pageNumber="24" refString="Mauries, J. - P. (1983) Cambalides nouveaux et peu connus d'Asie, d'Amerique et d'Oceanie. I. Cambalidae et Cambalopsidae (Myriapoda: Diplopoda). Bulletin du Museum national d'Histoire naturelle, serie 4, 5 A (1), 247 - 276." type="journal article" year="1983">Mauriès, 1983</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C7111E13E11FFA9FCD948E2B388FF4F" box="[815,966,229,254]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hallelujah" status="sp. nov.">
<emphasis id="B905B6703E11FFA9FCD948E2B388FF4F" box="[815,966,229,254]" italics="true" pageId="6" pageNumber="7">E. hallelujah</emphasis>
</taxonomicName>
<emphasis id="B905B6703E11FFA9FC3B48E2B45EFF4E" bold="true" box="[973,1040,229,255]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A2360B0B3E11FFA9FC3B48E2B45EFF4E" box="[973,1040,229,255]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C7111E13E11FFA9FBE848E2B4E5FF4F" box="[1054,1195,229,254]" class="Diplopoda" family="Cambalidae" genus="Eumastigonus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="waitahae" status="sp. nov.">
<emphasis id="B905B6703E11FFA9FBE848E2B4E5FF4F" box="[1054,1195,229,254]" italics="true" pageId="6" pageNumber="7">E. waitahae</emphasis>
</taxonomicName>
<emphasis id="B905B6703E11FFA9FB4748E2B4BAFF4E" bold="true" box="[1201,1268,229,255]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A2360B0B3E11FFA9FB4748E2B4BAFF4E" box="[1201,1268,229,255]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36B39E93E11FFA8FF30490CB3F1FEC7" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="8BCE6A623E11FFA9FF30490CB33AFEFD" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">
<emphasis id="B905B6703E11FFA9FF30490CB12FFE94" bold="true" box="[198,353,267,293]" pageId="6" pageNumber="7">Description:</emphasis>
Size: Length from 15 up to
<quantity id="4C89C7873E11FFA9FD3E490BB36FFE97" box="[712,801,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="mm" value="80.0">80 mm</quantity>
, maximum midbody diameter between 1.44.5 mm, number of podous rings: 3059, with 04 apodous, and telson.
</paragraph>
<paragraph id="8BCE6A623E11FFA9FF33495EB41EFD19" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">
Head as in
<figureCitation id="134A76E73E11FFA9FEB8495EB1D7FEC2" box="[334,409,345,371]" captionStart="FIGURES 4 9" captionStartId="5.[151,270,1865,1889]" captionTargetBox="[311,1272,202,1748]" captionTargetId="figure@5.[251,1322,194,1761]" captionTargetPageId="5" captionText="FIGURES 4 9. Eumastigonus kaorinus. 4: Gnathochilarium of paratype female (MCZ 4868), ventral view, 5: head of holotype male (MCZ 4867), left lateral view, 6: first male legpair, posterior view, 7: anterior gonopods, anterior view (c = coxa, m = median lobe), 8: right gonopods, right lateral view, posterior gonopod removed from embrace of anterior ones, flagellum omitted (tp = telopodite process), 9: posterior gonopods, posterior view (a = apex). Scales 4 5: 1 mm, 6 9: 0.5 mm." httpUri="https://zenodo.org/record/186933/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
; gnathochilarium generalized epinannolenidean
<typeStatus id="54CAD4C03E11FFA9FC13495EB457FEC2" box="[997,1049,345,371]" pageId="6" pageNumber="7">type</typeStatus>
(cf. with
<figureCitation id="134A76E73E11FFA9FB67495EB492FEC2" box="[1169,1244,345,371]" captionStart="FIGURES 4 9" captionStartId="5.[151,270,1865,1889]" captionTargetBox="[311,1272,202,1748]" captionTargetId="figure@5.[251,1322,194,1761]" captionTargetPageId="5" captionText="FIGURES 4 9. Eumastigonus kaorinus. 4: Gnathochilarium of paratype female (MCZ 4868), ventral view, 5: head of holotype male (MCZ 4867), left lateral view, 6: first male legpair, posterior view, 7: anterior gonopods, anterior view (c = coxa, m = median lobe), 8: right gonopods, right lateral view, posterior gonopod removed from embrace of anterior ones, flagellum omitted (tp = telopodite process), 9: posterior gonopods, posterior view (a = apex). Scales 4 5: 1 mm, 6 9: 0.5 mm." httpUri="https://zenodo.org/record/186933/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
, see also
<bibRefCitation id="EFE017933E11FFA9FAA7495EB09AFE2B" author="Jeekel" pageId="6" pageNumber="24" refString="Jeekel, C. A. W. (1985) The distribution of the Diplocheta and the &quot; lost &quot; continent Pacifica. - Bijdragen tot de Dierkunde 55 (1), 100 - 112." type="journal article" year="1985">Jeekel 1985</bibRefCitation>
) where mentum and promentum separate; mentum large, trapezoid, widely separating gnathochilarial stipites from each other; promentum undivided, large, triangular, separating laminae linguales which contact each other at their tips, with two setae each, close to the mouth opening. Gnathochilarial stipites widening and slightly bulging frontally, with two setae along their anterior margins and more on their basal parts; in males with pair of gnathochilarial pits (
<emphasis id="B905B6703E11FFA9FDD64A1DB273FD82" box="[544,573,538,563]" italics="true" pageId="6" pageNumber="7">gp</emphasis>
) each with one short seta, whose function is probably sensory. Posterior surface of gnathochilarial stipites and mentum rugose, but with a smooth half-circle centrally. Gular (hypostomal) plate in males with two lateral depressions (probably for the anterior projections of the first legpair to fit in). Mandibular stipites in males slightly widening. No frontal setae.
</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334AB2B4BAFCAE" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Ocelli (10-45) usually in 2-5 rows or irregularly arranged in a rectangular, oval, triangular or almondshaped ocellarium, posterior row parallel with anterior margin of collum. Antenna generally short, antennomeres 26th subequal in length, 1st and 7th shorter, small, with four apical sensory cones.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334B2BB205FC22" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Collum smooth, or with three or four weak marginal furrows on its lateral anterior corner; prozonae with irregular lines, whereas metazonae with longitudinal sutures, but only in the ventral third of the segmental part below ozopores. No metazonal setae.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334BA4B361FC55" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Ozopores only from 6th ring caudad, behind vertical segmental suture, usually small, inconspicuous, and in some cases hardly detectable on preserved specimens.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334BF7B3D9FB80" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Telson invariably without projection, paraprocts rounded, smooth, hairless, or 2+2 setae along the margin of anal opening, or rarely (in one species) entirely densely setose.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334C3AB4B0FB43" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Colouration: Typically uniformly dark or lighter brown, but sometimes with characteristic colour pattern: interocular region, collum anteriorly and laterally, and telson with dark brown markings, prozonae dark, metazonae, paraprocts and legs lighter. Terga sometimes with dark middorsal stripe or series of dark middorsal spots, accompanied by two longitudinal light stripes, or one wide middorsal light stripe with two dark parallel longitudinal stripes, or dorsum generally dark brown, sides below ozopores lighter.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334CF8B26CF9E0" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">
Male sexual characters: Mandibular stipites strongly widening, trapezoidal, their ventral sides not sclerotized, slightly swollen. Gnathochilarial stipites anteriorly with one sensory pit each in a seemingly soft, slightly swollen field. 1st legpair modified, short, stout, generally hidden between head and 2nd legpair, coxa fused with sternum (coxosternum), anteriorly provided with setae at the margin, prefemur short and wide, twice as wide as long, with two anterior blunt processes whose function is unknown (but
<bibRefCitation id="EFE017933E11FFA9FB1F4D9BB5D3FA07" author="Krabbe" box="[1257,1437,1436,1462]" pageId="6" pageNumber="24" refString="Krabbe, E. (1979) The first legpair of legs in male Spirostreptidae: Their function and taxonomic importance. In: Camatini, M. (Ed.), Myriapod biology. Academic Press, London, pp. 59 - 72." type="book chapter" year="1979">Krabbe (1979)</bibRefCitation>
suggested a clasping role during mating for similar processes in
<taxonomicName id="4C7111E13E11FFA9FC554DC4B410FA6C" box="[931,1118,1475,1501]" class="Diplopoda" family="Spirostreptidae" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
), femur almost normal, its length twice its width, mesal side straight, lateral side strongly concave, three apical podomeres (postfemur, tibia, and tarsus) subsimilar to each other, small, subequal in lengths and widths, tarsus without (or occasionally with) claw (Fig. 13).
</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334E66B2BDF910" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Penis closely behind 2nd legpair, deeply bifurcated, with two long apical branches, reaching almost to the end of prefemur, tips tube-like with hairy openings.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334EB6B408F943" blockId="6.[151,1438,151,2010]" pageId="6" pageNumber="7">Protrusion on 7th ring variable, sometimes glabrous and simple, in other cases rugose and significantly protruding, hence protecting the gonopods which emerge slightly between them.</paragraph>
<paragraph id="8BCE6A623E11FFA9FF334EF9B4C7F8A9" blockId="6.[151,1438,151,2010]" box="[197,1161,1790,1816]" pageId="6" pageNumber="7">Legpairs before and behind gonopodal ring with or without tarsal or femoral pads.</paragraph>
<paragraph id="8BCE6A623E11FFA8FF334F22B2B6FF69" blockId="6.[151,1438,151,2010]" lastBlockId="7.[151,1437,152,644]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Gonopods: Anterior gonopods: sternum forming a strong, bilobed plate in anterior view; coxal processes long, promerite-like, each with one inner process and one or two marginally hairy or pectinate, median and/or lateral lobes or processes. Telopodite caudo-basally with a field of spine-like setae. All these lobes or processes form an envelope into which the posterior gonopod is accommodated; flagellum long, welldeveloped, curving into a groove on posterior gonopods but then, together with them, comes back in between the processes of the anterior gonopods; posterior gonopods slender, long, more simple, usually with hairy coxal knob and thin, bent lamella (telopodite) at tip.</paragraph>
<paragraph id="8BCE6A623E10FFA8FF3048EFB3F1FEC7" blockId="7.[151,1437,152,644]" pageId="7" pageNumber="8">Female sexual characters: Vulvae embedded behind 2nd pair of legs in 3rd ring, relatively large, strongly flattened, disk-shaped, valves of bursa closely pressed against each other. Operculum small, narrow, situated anterio-laterally, tightly attached to bursa, thus whole vulva is very compact. Bursal valves with or without marginal hairs. Apodematic tube and ampulla could not be observed.</paragraph>
</subSubSection>
<subSubSection id="C36B39E93E10FFA8FF304985B361FE72" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BCE6A623E10FFA8FF304985B361FE72" blockId="7.[151,1437,152,644]" pageId="7" pageNumber="8">
<emphasis id="B905B6703E10FFA8FF304985B127FE2D" bold="true" box="[198,361,386,412]" pageId="7" pageNumber="8">Distribution:</emphasis>
<collectingCountry id="F3662AF23E10FFA8FE8C4985B26AFE2D" box="[378,548,386,412]" name="New Zealand" pageId="7" pageNumber="8">New Zealand</collectingCountry>
. Estimated to include about 30 species altogether. Some species are widespread, whereas others have restricted distributions.
</paragraph>
</subSubSection>
<subSubSection id="C36B39E93E10FFA8FF3049D7B490FD35" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BCE6A623E10FFA8FF3049D7B490FD35" blockId="7.[151,1437,152,644]" pageId="7" pageNumber="8">
<emphasis id="B905B6703E10FFA8FF3049D7B10CFE5B" bold="true" box="[198,322,464,490]" pageId="7" pageNumber="8">Remarks:</emphasis>
The group of spines on the posterior surface of the telopodite of the anterior gonopods can be considered as the remnant of the second telopodomere.
<bibRefCitation id="EFE017933E10FFA8FCC449F0B3ADFDA0" author="Enghoff" box="[818,995,503,529]" pageId="7" pageNumber="23" refString="Enghoff, H. (1985) The millipede family Nemasomatidae. With the description of a new genus, and a revision of Orinisobates (Diplopoda: Julida). Entomologica scandinavica 16, 27 - 67." type="journal article" year="1985">Enghoff (1985</bibRefCitation>
,
<bibRefCitation id="EFE017933E10FFA8FC0649F0B463FDA0" author="Enghoff" box="[1008,1069,503,529]" pageId="7" pageNumber="23" refString="Enghoff, H. (1991) A revised cladistic analysis and classification of the millipede order Julida. With establishment of four new families and description of a new nemasomatoid genus from Japan. Zeitschrift fur zoologische Systematik und Evolutionsforschung 29, 241 - 263." type="journal article" year="1991">1991</bibRefCitation>
) noticed a similar structure on the posterior gonopods of certain Nemasomatoidea (cf.
<figureCitation id="134A76E73E10FFA8FCB74A1AB3D9FD86" box="[833,919,541,567]" captionStart="FIGURES 40 43. E" captionStartId="18.[151,270,1905,1929]" captionTargetBox="[189,1390,195,1877]" captionTargetId="figure@18.[151,1436,148,1887]" captionTargetPageId="18" captionText="FIGURES 40 43. E. waitahae male from Okuku Pass, Canterbury Region. 40: Anterior gonopods, anterior view (tp = telopodite process, i = inner lobe, n = needle-like tip, st = sternal plate), 41: anterior gonopods, posterior view (tp = telopodite process, n = needle-like tip of inner lobe), 42: posterior gonopods, anterior view (sg = seminal groove, tp = telopodite, c = coxa, st = sternal plate, a = apex), 43: posterior gonopods, posterior view. Scales 0.5 mm." httpUri="https://zenodo.org/record/186940/files/figure.png" pageId="7" pageNumber="8">Fig. 42</figureCitation>
<emphasis id="B905B6703E10FFA8FC614A19B3E1FD86" box="[919,943,542,567]" italics="true" pageId="7" pageNumber="8">ss</emphasis>
in
<bibRefCitation id="EFE017933E10FFA8FC2E4A1AB4CCFD86" author="Enghoff" box="[984,1154,541,567]" pageId="7" pageNumber="23" refString="Enghoff, H. (1985) The millipede family Nemasomatidae. With the description of a new genus, and a revision of Orinisobates (Diplopoda: Julida). Entomologica scandinavica 16, 27 - 67." type="journal article" year="1985">Enghoff 1985</bibRefCitation>
, or
<figureCitation id="134A76E73E10FFA8FB474A1AB546FD86" box="[1201,1288,541,567]" captionStart="FIGURES 22 24" captionStartId="12.[151,268,1595,1619]" captionTargetBox="[180,1398,1265,1571]" captionTargetId="figure@12.[151,1436,1259,1572]" captionTargetPageId="12" captionText="FIGURES 22 24. Female vulvae (op = operculum). 22: E. kaorinus from Whangaruru headland, Northland Region, right vulva, anterior view, 23: E. distinctior from Porirua, Wellington Region, Galathea Expedition, 1951 (ZMUC), left vulva, posterior view, 24: E. hemmingseni from Akatarawa, Wellington Region, right vulva, posterior view. Scales 0.5 mm." httpUri="https://zenodo.org/record/186936/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
<emphasis id="B905B6703E10FFA8FAFE4A19B55EFD86" box="[1288,1296,542,567]" italics="true" pageId="7" pageNumber="8">f</emphasis>
in
<bibRefCitation id="EFE017933E10FFA8FACC4A1AB096FDEF" author="Enghoff" pageId="7" pageNumber="23" refString="Enghoff, H. (1991) A revised cladistic analysis and classification of the millipede order Julida. With establishment of four new families and description of a new nemasomatoid genus from Japan. Zeitschrift fur zoologische Systematik und Evolutionsforschung 29, 241 - 263." type="journal article" year="1991">Enghoff 1991</bibRefCitation>
). Nemastomatoids are considered to be close to the groundplan of
<taxonomicName id="4C7111E13E10FFA8FBFB4A43B417FDEF" box="[1037,1113,580,606]" class="Diplopoda" kingdom="Animalia" order="Julida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="order">Julida</taxonomicName>
, and
<taxonomicName id="4C7111E13E10FFA8FB6D4A43B500FDEF" box="[1179,1358,580,606]" class="Diplopoda" family="Iulomorphidae" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Iulomorphidae</taxonomicName>
to the groundplan of Epinannolenidea, so this character can be seen as a possible symplesiomorphy.
</paragraph>
</subSubSection>
</treatment>
</document>