345 lines
49 KiB
XML
345 lines
49 KiB
XML
<document id="D9CDF9A8A78C4A385A209EBC617433DF" ID-CLB-Dataset="26637" ID-DOI="10.11646/zootaxa.4612.3.2" ID-GBIF-Dataset="3835b1c0-ee9d-4a2a-bbcf-9454d050636f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3235112" ID-ZooBank="BDCF5C67-6320-48C9-A449-C4433869E0B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559194293764" checkinUser="plazi" docAuthor="Joharchi, Omid, Halliday, Bruce, Tolstikov, Andrei V. & Trach, Viacheslav A." docDate="2019" docId="03CF87940024FFA3FF3123BEAA30FE31" docLanguage="en" docName="zootaxa.4612.3.2.pdf" docOrigin="Zootaxa 4612 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantholaelaps Joharchi & Halliday & Tolstikov & Trach 2019, gen. nov." docType="treatment" docVersion="10" lastPageNumber="331" masterDocId="FFF6FFEC0027FFA6FFA62431AB20FFBA" masterDocTitle="New records and new species of mites from Cuba, with description of a new genus of Laelapidae (Acari: Mesostigmata)" masterLastPageNumber="350" masterPageNumber="326" pageNumber="329" updateTime="1698732318921" updateUser="ExternalLinkService">
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<mods:mods id="B4A909419637CBBA14BB78C356D92FE1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B44F4DCFC250F7525FD0F52D8F9DFA55">
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<mods:title id="95E6204061D5595497C16C438DB6BD47">New records and new species of mites from Cuba, with description of a new genus of Laelapidae (Acari: Mesostigmata)</mods:title>
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</mods:titleInfo>
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<mods:name id="98565FE39326A766EE2D823481BDD765" type="personal">
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<mods:namePart id="B0D6E1689192771A6E9E618705C5EC4B">Joharchi, Omid</mods:namePart>
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<mods:namePart id="256FDFCD4B36A0B6759210A6948337BF">Halliday, Bruce</mods:namePart>
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<mods:namePart id="935FE749B312B3D8AFBD3220A5D04574">Tolstikov, Andrei V.</mods:namePart>
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<mods:name id="892CDE6ACC98D52340F8ED0969F7EFB7" type="personal">
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<mods:namePart id="06E17BD396A2BFF1C91F17D323B8BE0D">Trach, Viacheslav A.</mods:namePart>
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<mods:typeOfResource id="784AD9373D962EE1411EE87EDDA8EEB5">text</mods:typeOfResource>
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<mods:title id="A8EC976351C9BBED5F6226FA35526BC7">Zootaxa</mods:title>
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<mods:part id="2FA71A76FAA33E9D1EBE24BB85562194">
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<mods:date id="D352265D020FA90399717AFD0BF4CB61">2019</mods:date>
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<mods:number id="CC5A00D3D90DFCDD77A27B3072942A53">2019-05-30</mods:number>
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<mods:number id="9778FD9E4D0C1B963897999DCDA7660E">4612</mods:number>
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<mods:number id="D815BBC168CEEDB0A13A32740D3327A4">3</mods:number>
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<mods:identifier id="776386856EADE8B98DDF02DCF04825F7" type="DOI">10.11646/zootaxa.4612.3.2</mods:identifier>
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<treatment id="03CF87940024FFA3FF3123BEAA30FE31" ID-DOI="http://doi.org/10.5281/zenodo.5697298" ID-GBIF-Taxon="157322166" ID-Zenodo-Dep="5697298" LSID="urn:lsid:plazi:treatment:03CF87940024FFA3FF3123BEAA30FE31" httpUri="http://treatment.plazi.org/id/03CF87940024FFA3FF3123BEAA30FE31" lastPageId="5" lastPageNumber="331" pageId="3" pageNumber="329">
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<subSubSection id="C37C65090024FFA5FF3123BEA93FF810" box="[151,543,1935,1962]" pageId="3" pageNumber="329" type="nomenclature">
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<paragraph id="8BD936820024FFA5FF3123BEA93FF810" blockId="3.[151,543,1935,1962]" box="[151,543,1935,1962]" pageId="3" pageNumber="329">
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<heading id="D09181EE0024FFA5FF3123BEA93FF810" bold="true" box="[151,543,1935,1962]" fontSize="11" level="1" pageId="3" pageNumber="329" reason="1">
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<emphasis id="B912EA900024FFA5FF3123BEA93FF810" bold="true" box="[151,543,1935,1962]" pageId="3" pageNumber="329">
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Genus
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<taxonomicName id="4C664D010024FFA5FF4823BEAA89F810" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[238,425,1935,1962]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="329" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B912EA900024FFA5FF4823BEAA89F810" bold="true" box="[238,425,1935,1962]" italics="true" pageId="3" pageNumber="329">Acantholaelaps</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157EB0024FFA5FE1723A1A93FF810" box="[433,543,1936,1962]" pageId="3" pageNumber="329" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37C65090024FFA5FF3123E5A95EF856" box="[151,638,2003,2028]" pageId="3" pageNumber="329" type="materials_examined">
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<paragraph id="8BD936820024FFA5FF3123E5A95EF856" blockId="3.[151,638,2003,2028]" box="[151,638,2003,2028]" pageId="3" pageNumber="329">
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<typeStatus id="54DD88200024FFA5FF3123E5ABE9F856" box="[151,201,2004,2028]" pageId="3" pageNumber="329">Type</typeStatus>
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species:
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<taxonomicName id="4C664D010024FFA5FE8023E5A907F851" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[294,551,2004,2028]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="329" phylum="Arthropoda" rank="species" species="strategus" status="sp. nov.">
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<emphasis id="B912EA900024FFA5FE8023E5A907F851" box="[294,551,2004,2028]" italics="true" pageId="3" pageNumber="329">Acantholaelaps strategus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157EB0024FFA5FD8B23E5A95EF856" box="[557,638,2004,2028]" pageId="3" pageNumber="329" rank="species">
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<emphasis id="B912EA900024FFA5FD8B23E5A95EF856" bold="true" box="[557,638,2004,2028]" pageId="3" pageNumber="329">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C65090023FFA2FF3124A9A931FCE5" pageId="4" pageNumber="330" type="description">
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<paragraph id="8BD936820023FFA2FF3124A9A90DFD75" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="330">
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<emphasis id="B912EA900023FFA2FF3124A9AA09FF09" bold="true" box="[151,297,152,179]" pageId="4" pageNumber="330">Description.</emphasis>
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<emphasis id="B912EA900023FFA2FE9424A8AAA6FF09" box="[306,390,153,179]" italics="true" pageId="4" pageNumber="330">Female</emphasis>
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.
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<taxonomicName id="4C664D010023FFA2FE3024A8A931FF09" authorityName="A.Berlese" authorityYear="1892" box="[406,529,153,179]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
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with an oval-shaped dorsal shield without lateral incisions, bearing 28 pairs of setae (28 pairs in
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<specimenCount id="9D60FD0B0023FFA2FEFC248CAAD2FF6D" box="[346,498,189,215]" pageId="4" pageNumber="330" type="generic">14 specimens</specimenCount>
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including
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<typeStatus id="54DD88200023FFA2FDCE248CA9EDFF6D" box="[616,717,189,215]" pageId="4" pageNumber="330" type="holotype">holotype</typeStatus>
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, 29 pairs in three specimens [
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<emphasis id="B912EA900023FFA2FBB0248CAF11FF6D" box="[1046,1073,189,215]" italics="true" pageId="4" pageNumber="330">S3</emphasis>
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on shield] and 30 pairs [
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<emphasis id="B912EA900023FFA2FAE3248CAE7EFF6D" box="[1349,1374,189,215]" italics="true" pageId="4" pageNumber="330">s2</emphasis>
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&
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<emphasis id="B912EA900023FFA2FA26248CAEBCFF6D" box="[1408,1436,189,215]" italics="true" pageId="4" pageNumber="330">S3</emphasis>
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on shield] in three specimens), without
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<emphasis id="B912EA900023FFA2FDF024D3A952FF41" box="[598,626,226,251]" italics="true" pageId="4" pageNumber="330">Zx</emphasis>
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setae between the
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<emphasis id="B912EA900023FFA2FCEF24D3A875FF41" box="[841,853,226,251]" italics="true" pageId="4" pageNumber="330">J</emphasis>
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and
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<emphasis id="B912EA900023FFA2FC2824D3A8BEFF41" box="[910,926,226,251]" italics="true" pageId="4" pageNumber="330">Z</emphasis>
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setae; all dorsal shield setae smooth, pointed and variable in length, marginal setae longest and often appearing wavy in slide-mounted specimens,
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<emphasis id="B912EA900023FFA2FA8F2534AE65FEA5" box="[1321,1349,261,287]" italics="true" pageId="4" pageNumber="330">S5</emphasis>
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greatly elongated and wavy in form, at least two-thirds of the body length (
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<figureCitation id="135D2A070023FFA2FC2B2518AF28FEF9" box="[909,1032,297,323]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,1936,1961]" captionTargetBox="[168,1379,197,1881]" captionTargetId="figure@6.[154,1393,176,1898]" captionTargetPageId="6" captionText="FIGURES 12–16. Acantholaelaps strategus sp. nov., female. 12, dorsal idiosoma; 13, ventral idiosoma; 14, subcapitulum; 15, epistome; 16, chelicera." httpUri="https://zenodo.org/record/3235118/files/figure.png" pageId="4" pageNumber="330">Figures 12</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FBB32518AF7DFEF9" box="[1045,1117,297,323]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1641,1666]" captionTargetBox="[232,1358,196,1589]" captionTargetId="figure@7.[227,1361,186,1595]" captionTargetPageId="7" captionText="FIGURES 17–20. DIC micrographs of Acantholaelaps strategus sp. nov., female: 17, 18, idiosoma in dorsal view, showing variation in the length of setae in central area of the shield, 19, 20, idiosoma in ventral view, showing variation in the size of body and level of sternal shield sclerotisation." httpUri="https://zenodo.org/record/3235120/files/figure.png" pageId="4" pageNumber="330">17, 18</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FBCF2518AE28FEFE" box="[1129,1288,297,324]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1935,1960]" captionTargetBox="[227,1358,614,1895]" captionTargetId="figure@8.[227,1361,604,1896]" captionTargetPageId="8" captionText="FIGURES 21–26. SEM micrographs of Acantholaelaps strategus sp. nov., female. 21, idiosoma in dorsal view; 22, idiosoma in ventral view; 23, podonotal area; 24, opisthonotal area; 25, idiosoma in lateral view; 26, long caudal setae." httpUri="https://zenodo.org/record/3235122/files/figure.png" pageId="4" pageNumber="330">21, 23, 24, 26</figureCitation>
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). Lateral soft skin and opisthogastric area strongly hypertrichous (
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<figureCitation id="135D2A070023FFA2FD7A257CA86CFEDD" box="[732,844,332,359]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1935,1960]" captionTargetBox="[227,1358,614,1895]" captionTargetId="figure@8.[227,1361,604,1896]" captionTargetPageId="8" captionText="FIGURES 21–26. SEM micrographs of Acantholaelaps strategus sp. nov., female. 21, idiosoma in dorsal view; 22, idiosoma in ventral view; 23, podonotal area; 24, opisthonotal area; 25, idiosoma in lateral view; 26, long caudal setae." httpUri="https://zenodo.org/record/3235122/files/figure.png" pageId="4" pageNumber="330">Figure 25</figureCitation>
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); one pair of caudal setae greatly elongated and wavy in form, at least two-thirds of the body length (
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<figureCitation id="135D2A070023FFA2FD042540A83DFE31" box="[674,797,369,395]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,1936,1961]" captionTargetBox="[168,1379,197,1881]" captionTargetId="figure@6.[154,1393,176,1898]" captionTargetPageId="6" captionText="FIGURES 12–16. Acantholaelaps strategus sp. nov., female. 12, dorsal idiosoma; 13, ventral idiosoma; 14, subcapitulum; 15, epistome; 16, chelicera." httpUri="https://zenodo.org/record/3235118/files/figure.png" pageId="4" pageNumber="330">Figures 13</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FC8E2540A868FE31" box="[808,840,369,395]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1935,1960]" captionTargetBox="[227,1358,614,1895]" captionTargetId="figure@8.[227,1361,604,1896]" captionTargetPageId="8" captionText="FIGURES 21–26. SEM micrographs of Acantholaelaps strategus sp. nov., female. 21, idiosoma in dorsal view; 22, idiosoma in ventral view; 23, podonotal area; 24, opisthonotal area; 25, idiosoma in lateral view; 26, long caudal setae." httpUri="https://zenodo.org/record/3235122/files/figure.png" pageId="4" pageNumber="330">22</figureCitation>
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). Sternal shield reduced in size and longer than wide, its anterior margin often poorly defined, bearing two pairs of long setae,
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<emphasis id="B912EA900023FFA2FC6925A4A8D0FE15" box="[975,1008,405,431]" italics="true" pageId="4" pageNumber="330">st1</emphasis>
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off the shield; genital shield with one pair of setae (
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<figureCitation id="135D2A070023FFA2FE942588AA8EFE69" box="[306,430,441,467]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,1936,1961]" captionTargetBox="[168,1379,197,1881]" captionTargetId="figure@6.[154,1393,176,1898]" captionTargetPageId="6" captionText="FIGURES 12–16. Acantholaelaps strategus sp. nov., female. 12, dorsal idiosoma; 13, ventral idiosoma; 14, subcapitulum; 15, epistome; 16, chelicera." httpUri="https://zenodo.org/record/3235118/files/figure.png" pageId="4" pageNumber="330">Figures 13</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FE1D2588A926FE69" box="[443,518,441,467]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1641,1666]" captionTargetBox="[232,1358,196,1589]" captionTargetId="figure@7.[227,1361,186,1595]" captionTargetPageId="7" captionText="FIGURES 17–20. DIC micrographs of Acantholaelaps strategus sp. nov., female: 17, 18, idiosoma in dorsal view, showing variation in the length of setae in central area of the shield, 19, 20, idiosoma in ventral view, showing variation in the size of body and level of sternal shield sclerotisation." httpUri="https://zenodo.org/record/3235120/files/figure.png" pageId="4" pageNumber="330">19, 20</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FDB42588A911FE69" box="[530,561,441,467]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1935,1960]" captionTargetBox="[227,1358,614,1895]" captionTargetId="figure@8.[227,1361,604,1896]" captionTargetPageId="8" captionText="FIGURES 21–26. SEM micrographs of Acantholaelaps strategus sp. nov., female. 21, idiosoma in dorsal view; 22, idiosoma in ventral view; 23, podonotal area; 24, opisthonotal area; 25, idiosoma in lateral view; 26, long caudal setae." httpUri="https://zenodo.org/record/3235122/files/figure.png" pageId="4" pageNumber="330">22</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FD9B2588A9A8FE69" box="[573,648,441,467]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,1092,1117]" captionTargetBox="[230,1359,195,1043]" captionTargetId="figure@10.[227,1361,188,1044]" captionTargetPageId="10" captionText="FIGURES 33–36. SEM micrographs of Acantholaelaps strategus sp. nov., female. 33, sternal shield; 34, genital, anal shield and opisthogastric area; 35, gnathosoma; 36, subcapitulum and tritosternum." httpUri="https://zenodo.org/record/3235126/files/figure.png" pageId="4" pageNumber="330">33, 34</figureCitation>
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). Anal shield with para-anal setae longer than unpaired post-anal seta (
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<figureCitation id="135D2A070023FFA2FF3925ECAA39FE4D" box="[159,281,477,503]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,1936,1961]" captionTargetBox="[168,1379,197,1881]" captionTargetId="figure@6.[154,1393,176,1898]" captionTargetPageId="6" captionText="FIGURES 12–16. Acantholaelaps strategus sp. nov., female. 12, dorsal idiosoma; 13, ventral idiosoma; 14, subcapitulum; 15, epistome; 16, chelicera." httpUri="https://zenodo.org/record/3235118/files/figure.png" pageId="4" pageNumber="330">Figures 13</figureCitation>
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,
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<figureCitation id="135D2A070023FFA2FE8225ECAA64FE4D" box="[292,324,477,503]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,1092,1117]" captionTargetBox="[230,1359,195,1043]" captionTargetId="figure@10.[227,1361,188,1044]" captionTargetPageId="10" captionText="FIGURES 33–36. SEM micrographs of Acantholaelaps strategus sp. nov., female. 33, sternal shield; 34, genital, anal shield and opisthogastric area; 35, gnathosoma; 36, subcapitulum and tritosternum." httpUri="https://zenodo.org/record/3235126/files/figure.png" pageId="4" pageNumber="330">34</figureCitation>
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). Hypostome with six rows of minute teeth; hypostomal seta
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<emphasis id="B912EA900023FFA2FC4B25ECAF29FE4D" box="[1005,1033,477,503]" italics="true" pageId="4" pageNumber="330">h3</emphasis>
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longer than other hypostomal setae. Trochanter of palp bearing two long setae, palp tarsal claw two-tined (
|
||
<figureCitation id="135D2A070023FFA2FC012630AF01FDA1" box="[935,1057,513,539]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,1936,1961]" captionTargetBox="[168,1379,197,1881]" captionTargetId="figure@6.[154,1393,176,1898]" captionTargetPageId="6" captionText="FIGURES 12–16. Acantholaelaps strategus sp. nov., female. 12, dorsal idiosoma; 13, ventral idiosoma; 14, subcapitulum; 15, epistome; 16, chelicera." httpUri="https://zenodo.org/record/3235118/files/figure.png" pageId="4" pageNumber="330">Figures 14</figureCitation>
|
||
,
|
||
<figureCitation id="135D2A070023FFA2FB8B2630AF53FDA1" box="[1069,1139,513,539]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">28, 29</figureCitation>
|
||
&
|
||
<figureCitation id="135D2A070023FFA2FB302630AF96FDA6" box="[1174,1206,513,540]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,1092,1117]" captionTargetBox="[230,1359,195,1043]" captionTargetId="figure@10.[227,1361,188,1044]" captionTargetPageId="10" captionText="FIGURES 33–36. SEM micrographs of Acantholaelaps strategus sp. nov., female. 33, sternal shield; 34, genital, anal shield and opisthogastric area; 35, gnathosoma; 36, subcapitulum and tritosternum." httpUri="https://zenodo.org/record/3235126/files/figure.png" pageId="4" pageNumber="330">36</figureCitation>
|
||
). Peritrematal shield broadly enlarged around stigmatic opening (
|
||
<figureCitation id="135D2A070023FFA2FD252614A9D4FD85" box="[643,756,549,575]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">Figure 27</figureCitation>
|
||
), post-stigmatic section not clearly defined (
|
||
<figureCitation id="135D2A070023FFA2FB472614AE73FD85" box="[1249,1363,549,575]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">Figure 27</figureCitation>
|
||
). Legs I longer than legs II & III, legs IV longest (
|
||
<figureCitation id="135D2A070023FFA2FD262678A9D1FDD9" box="[640,753,585,611]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1935,1960]" captionTargetBox="[227,1358,614,1895]" captionTargetId="figure@8.[227,1361,604,1896]" captionTargetPageId="8" captionText="FIGURES 21–26. SEM micrographs of Acantholaelaps strategus sp. nov., female. 21, idiosoma in dorsal view; 22, idiosoma in ventral view; 23, podonotal area; 24, opisthonotal area; 25, idiosoma in lateral view; 26, long caudal setae." httpUri="https://zenodo.org/record/3235122/files/figure.png" pageId="4" pageNumber="330">Figure 21</figureCitation>
|
||
); greatly elongated macrosetae present on genua and femora I–IV (
|
||
<emphasis id="B912EA900023FFA2FF7A265CAA26FD3D" box="[220,262,621,647]" italics="true" pageId="4" pageNumber="330">pd1</emphasis>
|
||
on legs I & II and
|
||
<emphasis id="B912EA900023FFA2FE7C265CA924FD3D" box="[474,516,621,647]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
on III & IV). Tibia III with eight setae (2 1/1 2/1 1) (
|
||
<figureCitation id="135D2A070023FFA2FBF0265CAFDDFD3D" box="[1110,1277,621,647]" captionStart-0="FIGURES 37–38" captionStart-1="FIGURES 39–40" captionStartId-0="11.[151,264,2018,2043]" captionStartId-1="12.[151,264,1938,1963]" captionTargetBox-0="[177,1404,442,1862]" captionTargetBox-1="[171,1424,227,1850]" captionTargetId-0="figure@11.[153,1438,420,1885]" captionTargetId-1="figure@12.[153,1438,210,1864]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 37–38. Acantholaelaps strategus sp. nov., female. 37, leg I (trochanter-tibia); 38, leg II." captionText-1="FIGURES 39–40. Acantholaelaps strategus sp. nov., female. 39, leg III; 40, leg IV." httpUri-0="https://zenodo.org/record/3235128/files/figure.png" httpUri-1="https://zenodo.org/record/3235130/files/figure.png" pageId="4" pageNumber="330">Figures 37–40</figureCitation>
|
||
). Tarsus II–IV with subterminal blunt spines (
|
||
<emphasis id="B912EA900023FFA2FE5326A0A939FD11" box="[501,537,657,683]" italics="true" pageId="4" pageNumber="330">al1</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FDF726A0A955FD11" box="[593,629,657,683]" italics="true" pageId="4" pageNumber="330">pl1</emphasis>
|
||
); pre-tarsi II–IV with a pair of stout claws, each claw with a basal spur on its inner margin (
|
||
<figureCitation id="135D2A070023FFA2FEDC2684A900FD75" box="[378,544,693,719]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">Figures 31–32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936820023FFA2FF6126E8A931FCE5" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="330">
|
||
<emphasis id="B912EA900023FFA2FF6126E8AA21FD49" box="[199,257,729,755]" italics="true" pageId="4" pageNumber="330">Male</emphasis>
|
||
. As for female except ventral idiosoma with separate sternogenital and anal shields, and an extra irregularly-shaped area of sclerotisation between them (
|
||
<figureCitation id="135D2A070023FFA2FD6C26CCA853FCAD" box="[714,883,765,791]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1689,1714]" captionTargetBox="[242,1348,201,1628]" captionTargetId="figure@15.[227,1361,190,1640]" captionTargetPageId="15" captionText="FIGURES 47–52. DIC micrographs of Acantholaelaps strategus sp. nov., male. 47, idiosoma in ventral view; 48, composite shield; ventral irregular shield & anal shield; 49, ventral irregular shield; 50, chelicera; 51, tibia & tarsus II; 52, tibia & tarsus III." httpUri="https://zenodo.org/record/3235136/files/figure.png" pageId="4" pageNumber="330">Figures 47–49</figureCitation>
|
||
). Tarsi II and III with strong spine-like setae
|
||
<emphasis id="B912EA900023FFA2FAD226CCAEBCFCAD" box="[1396,1436,765,791]" italics="true" pageId="4" pageNumber="330">av1</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FF6F2710ABD2FC81" box="[201,242,801,827]" italics="true" pageId="4" pageNumber="330">av2</emphasis>
|
||
(
|
||
<figureCitation id="135D2A070023FFA2FEA42710AA8CFC81" box="[258,428,800,827]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1689,1714]" captionTargetBox="[242,1348,201,1628]" captionTargetId="figure@15.[227,1361,190,1640]" captionTargetPageId="15" captionText="FIGURES 47–52. DIC micrographs of Acantholaelaps strategus sp. nov., male. 47, idiosoma in ventral view; 48, composite shield; ventral irregular shield & anal shield; 49, ventral irregular shield; 50, chelicera; 51, tibia & tarsus II; 52, tibia & tarsus III." httpUri="https://zenodo.org/record/3235136/files/figure.png" pageId="4" pageNumber="330">Figures 51–52</figureCitation>
|
||
). Spermatodactyl longer than movable digit, sharply bent upward apically into S-shape, with truncate tip (
|
||
<figureCitation id="135D2A070023FFA2FEFB2774AAF9FCE5" box="[349,473,837,863]" captionStart="FIGURES 45–46" captionStartId="14.[151,264,2004,2029]" captionTargetBox="[210,1377,786,1914]" captionTargetId="figure@14.[185,1406,777,1938]" captionTargetPageId="14" captionText="FIGURES 45–46. Acantholaelaps strategus sp. nov., male. 45, ventral idiosoma; 46, chelicera." httpUri="https://zenodo.org/record/3235134/files/figure.png" pageId="4" pageNumber="330">Figures 46</figureCitation>
|
||
,
|
||
<figureCitation id="135D2A070023FFA2FE422775A924FCE5" box="[484,516,836,863]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1689,1714]" captionTargetBox="[242,1348,201,1628]" captionTargetId="figure@15.[227,1361,190,1640]" captionTargetPageId="15" captionText="FIGURES 47–52. DIC micrographs of Acantholaelaps strategus sp. nov., male. 47, idiosoma in ventral view; 48, composite shield; ventral irregular shield & anal shield; 49, ventral irregular shield; 50, chelicera; 51, tibia & tarsus II; 52, tibia & tarsus III." httpUri="https://zenodo.org/record/3235136/files/figure.png" pageId="4" pageNumber="330">50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C65090023FFA3FF612759A8A7FEF9" lastPageId="5" lastPageNumber="331" pageId="4" pageNumber="330" type="diagnosis">
|
||
<paragraph id="8BD936820023FFA2FF612759A8CAFBE1" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="330">
|
||
<emphasis id="B912EA900023FFA2FF612759AAEDFC39" bold="true" box="[199,461,872,899]" pageId="4" pageNumber="330">Differential diagnosis.</emphasis>
|
||
The new genus can easily be distinguished from other related members of the family by a combination of morphological attributes, some of which are unique or rarely observed in laelapid mites, such as (1) dorsal seta
|
||
<emphasis id="B912EA900023FFA2FE982780AA7AFC71" box="[318,346,945,971]" italics="true" pageId="4" pageNumber="330">S5</emphasis>
|
||
markedly elongated (2) very long dorsal setae (macrosetae) on femur and genu of legs I-IV (
|
||
<emphasis id="B912EA900023FFA2FAD42780AEBDFC71" box="[1394,1437,945,971]" italics="true" pageId="4" pageNumber="330">pd1</emphasis>
|
||
on legs I & II and
|
||
<emphasis id="B912EA900023FFA2FEC627E4AAAAFC55" box="[352,394,981,1007]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
on III & IV) (3) tibia III bears eight setae (2 1/1 2/1 1) (4) tarsus II–IV with subterminal blunt spines (
|
||
<emphasis id="B912EA900023FFA2FF4927C8AA33FBA9" box="[239,275,1017,1043]" italics="true" pageId="4" pageNumber="330">al1</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FEED27C8AA4FFBA9" box="[331,367,1017,1043]" italics="true" pageId="4" pageNumber="330">pl1</emphasis>
|
||
) (5) claws on pre-tarsi II–IV with a basal spur on their convex surface; (6) peritrematal shield broadly enlarged around stigmatic opening; (7) sternal shield reduced in size and longer than wide and bearing two pairs setae,
|
||
<emphasis id="B912EA900023FFA2FEBD2070AA1BFBE1" box="[283,315,1089,1115]" italics="true" pageId="4" pageNumber="330">st1</emphasis>
|
||
off the shield; (8) opisthogastric area strongly hypertrichous.
|
||
</paragraph>
|
||
<paragraph id="8BD936820023FFA2FF612054AE08F921" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="330">
|
||
Females of the new genus are similar to
|
||
<taxonomicName id="4C664D010023FFA2FD3C2054A859FBC5" authority="Clark" authorityName="Clark" authorityYear="2012" box="[666,889,1125,1151]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FD3C2054A812FBC5" box="[666,818,1125,1151]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
Clark
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="EFF74B730023FFA2FC0C2054AF89FBC5" author="Clark, J. M. & Hawke, D. J." box="[938,1193,1125,1151]" pageId="4" pageNumber="330" pagination="187 - 199" refId="ref9857" refString="Clark, J. M. & Hawke, D. J. (2012) A new epizoic laelapid mite from the New Zealand sand scarab Pericoptus truncatus larvae and its isotopic ecology. New Zealand Journal of Zoology, 39, 187 - 199. https: // doi. org / 10.1080 / 03014223.2011.628997" type="journal article" year="2012">Clark & Hawke, 2012</bibRefCitation>
|
||
) in having the dorsal shield without lateral incisions, lateral soft skin strongly hypertrichous, sternal shield reduced in size and longer than wide and bearing two pairs setae,
|
||
<emphasis id="B912EA900023FFA2FDEB209CA94EFB7D" box="[589,622,1197,1223]" italics="true" pageId="4" pageNumber="330">st1</emphasis>
|
||
off the shield, legs I longer than II and III, elongated setae (macrosetae) on genua and femora I-IV (
|
||
<emphasis id="B912EA900023FFA2FE6820E0AAD8FB51" box="[462,504,1233,1259]" italics="true" pageId="4" pageNumber="330">pd1</emphasis>
|
||
on legs I & II and
|
||
<emphasis id="B912EA900023FFA2FD7420E0A9DCFB51" box="[722,764,1233,1259]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
on III & IV), tibia III bearing eight setae (2 1/1 2/1 1), and tarsus II-IV with subterminal blunt spines (
|
||
<emphasis id="B912EA900023FFA2FDD820C4A982FAB5" box="[638,674,1269,1295]" italics="true" pageId="4" pageNumber="330">al1</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FD7F20C4A9DDFAB5" box="[729,765,1269,1295]" italics="true" pageId="4" pageNumber="330">pl1</emphasis>
|
||
). The new genus differs from
|
||
<taxonomicName id="4C664D010023FFA2FBF320C4AFCDFAB5" authorityName="Clark" authorityYear="2012" box="[1109,1261,1269,1295]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FBF320C4AFCDFAB5" box="[1109,1261,1269,1295]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
by the 28 pairs of dorsal shield setae (15 podonotal, 13 opisthonotal), with
|
||
<emphasis id="B912EA900023FFA2FC9A212BA87DFA89" box="[828,861,1305,1331]" italics="true" pageId="4" pageNumber="330">j–J</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FC30212BA89FFA89" box="[918,959,1305,1331]" italics="true" pageId="4" pageNumber="330">z–Z</emphasis>
|
||
series complete (vs. 22 pairs (12 podonotal, 10 opisthonotal) and missing many setae from
|
||
<emphasis id="B912EA900023FFA2FD73210FA9D7FAED" box="[725,759,1341,1367]" italics="true" pageId="4" pageNumber="330">j–J</emphasis>
|
||
and
|
||
<emphasis id="B912EA900023FFA2FC96210FA879FAED" box="[816,857,1341,1367]" italics="true" pageId="4" pageNumber="330">z–Z</emphasis>
|
||
series in
|
||
<taxonomicName id="4C664D010023FFA2FC63210CAF7DFAED" authorityName="Clark" authorityYear="2012" box="[965,1117,1341,1367]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FC63210CAF7DFAED" box="[965,1117,1341,1367]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
), setae
|
||
<emphasis id="B912EA900023FFA2FB10210CAFF2FAED" box="[1206,1234,1341,1367]" italics="true" pageId="4" pageNumber="330">S5</emphasis>
|
||
greatly elongated and on dorsal shield (vs not markedly elongated and off the dorsal shield), setae
|
||
<emphasis id="B912EA900023FFA2FB8F2150AF67FAC1" box="[1065,1095,1377,1403]" italics="true" pageId="4" pageNumber="330">Z4</emphasis>
|
||
&
|
||
<emphasis id="B912EA900023FFA2FBCA2150AFAAFAC1" box="[1132,1162,1377,1403]" italics="true" pageId="4" pageNumber="330">Z5</emphasis>
|
||
not markedly elongated (vs markedly elongated), opisthogastric area strongly hypertrichous (one pair of caudal setae greatly elongated, at least two-thirds of the body length) (vs. opisthogastric area hypotrichous in
|
||
<taxonomicName id="4C664D010023FFA2FC422198AF5CFA79" authorityName="Clark" authorityYear="2012" box="[996,1148,1449,1475]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FC422198AF5CFA79" box="[996,1148,1449,1475]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
, lacking very long caudal setae), peritrematal shield broadly enlarged around stigmatic opening (
|
||
<figureCitation id="135D2A070023FFA2FC6421FCAF13FA5D" box="[962,1075,1485,1511]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">Figure 27</figureCitation>
|
||
) and post-stigmatic section not conspicuous (vs. peritrematal shield normal and post-stigmatic section conspicuous in
|
||
<taxonomicName id="4C664D010023FFA2FBF521C0AFCBF9B1" authorityName="Clark" authorityYear="2012" box="[1107,1259,1521,1547]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FBF521C0AFCBF9B1" box="[1107,1259,1521,1547]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
), one elongated seta (
|
||
<emphasis id="B912EA900023FFA2FF732224ABDFF995" box="[213,255,1557,1583]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
) (macroseta) on each genua III–IV, and tibia IV without elongated macrosetae (vs. two elongated setae (
|
||
<emphasis id="B912EA900023FFA2FF072208ABEAF9E9" box="[161,202,1593,1619]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
,
|
||
<emphasis id="B912EA900023FFA2FF7F2208AA24F9E9" box="[217,260,1593,1619]" italics="true" pageId="4" pageNumber="330">pd1</emphasis>
|
||
) (macrosetae) on each genua III–IV and
|
||
<emphasis id="B912EA900023FFA2FD732208A9DFF9E9" box="[725,767,1593,1619]" italics="true" pageId="4" pageNumber="330">ad1</emphasis>
|
||
on tibia IV elongated setae (macrosetae) in
|
||
<taxonomicName id="4C664D010023FFA2FB552208AEABF9E9" authorityName="Clark" authorityYear="2012" box="[1267,1419,1593,1619]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FB552208AEABF9E9" box="[1267,1419,1593,1619]" italics="true" pageId="4" pageNumber="330">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
), and pre-tarsi II–IV with a basal spur on the inner surface of each tarsal claw (
|
||
<figureCitation id="135D2A070023FFA2FC41226CAFADF9CD" box="[999,1165,1629,1655]" captionStart="FIGURES 27–32" captionStartId="9.[151,264,1632,1657]" captionTargetBox="[247,1343,186,1584]" captionTargetId="figure@9.[227,1361,181,1592]" captionTargetPageId="9" captionText="FIGURES 27–32. DIC micrographs of Acantholaelaps strategus sp. nov., female. 27, stigma, post-stigmatic section & peritreme; 28, subcapitulum; 29, palp and tarsal claw; 30, fixed digit of chelicera, dorsal view; 31, pre-tarsus III (arrow shows claw with a basal spur on its inner margin); 32, pre-tarsus IV (arrow shows claw with a basal spur on its inner margin)." httpUri="https://zenodo.org/record/3235124/files/figure.png" pageId="4" pageNumber="330">Figures 31–32</figureCitation>
|
||
). The last character state is otherwise unknown in the free-living
|
||
<taxonomicName id="4C664D010023FFA2FDF122B0A9F5F921" authorityName="A.Berlese" authorityYear="1892" box="[599,725,1665,1691]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
|
||
, and appears to be autapomorphic for the new genus.
|
||
</paragraph>
|
||
<paragraph id="8BD936820023FFA3FF612294A8A7FEF9" blockId="4.[151,1437,152,2015]" lastBlockId="5.[151,1437,153,395]" lastPageId="5" lastPageNumber="331" pageId="4" pageNumber="330">
|
||
The new genus is also very similar to
|
||
<taxonomicName id="4C664D010023FFA2FDD02294A82AF905" box="[630,778,1701,1727]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FDD02294A82AF905" box="[630,778,1701,1727]" italics="true" pageId="4" pageNumber="330">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
. These two genera both have lightly sclerotised idiosomal shields, sternal shield reduced to leave small triangular endopodal fragments, sternal setae
|
||
<emphasis id="B912EA900023FFA2FB3022F8AF97F959" box="[1174,1207,1737,1763]" italics="true" pageId="4" pageNumber="330">st1</emphasis>
|
||
inserted in soft skin anterior to the sternal shield, and greatly elongated setae on the dorsal shield and some leg segments. These appear to be apomorphic character states in both genera. Most species of
|
||
<taxonomicName id="4C664D010023FFA2FC2E2320AF3CF891" box="[904,1052,1809,1835]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FC2E2320AF3CF891" box="[904,1052,1809,1835]" italics="true" pageId="4" pageNumber="330">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
have a posteriorly reduced dorsal shield, which also appears to be apomorphic, but
|
||
<taxonomicName id="4C664D010023FFA2FD6B2304AF52F8F5" authority="Costa & Hunter, 1971" authorityName="Costa & Hunter" authorityYear="1971" box="[717,1138,1845,1871]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="species" species="abnormalis">
|
||
<emphasis id="B912EA900023FFA2FD6B2304A9C0F8F5" box="[717,736,1845,1871]" italics="true" pageId="4" pageNumber="330">C</emphasis>
|
||
.
|
||
<emphasis id="B912EA900023FFA2FD492304A84FF8F5" box="[751,879,1845,1871]" italics="true" pageId="4" pageNumber="330">abnormalis</emphasis>
|
||
<bibRefCitation id="EFF74B730023FFA2FCD12304AF52F8F5" author="Costa, M. & Hunter, P. E." box="[887,1138,1845,1871]" pageId="4" pageNumber="330" pagination="323 - 360" refId="ref9995" refString="Costa, M. & Hunter, P. E. (1971) The genus Coleolaelaps Berlese, 1914 (Acarina: Mesostigmata). Redia, 52, 323 - 360." type="journal article" year="1971">Costa & Hunter, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
has a normal oval-shaped dorsal shield.
|
||
<taxonomicName id="4C664D010023FFA2FE972368A868F8C9" authority="Costa & Hunter, 1971" authorityName="Costa & Hunter" authorityYear="1971" box="[305,840,1881,1907]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="species" species="brachysetus">
|
||
<emphasis id="B912EA900023FFA2FE972368A970F8C9" box="[305,592,1881,1907]" italics="true" pageId="4" pageNumber="330">Coleolaelaps brachysetus</emphasis>
|
||
<bibRefCitation id="EFF74B730023FFA2FDF02368A868F8C9" author="Costa, M. & Hunter, P. E." box="[598,840,1881,1907]" pageId="4" pageNumber="330" pagination="323 - 360" refId="ref9995" refString="Costa, M. & Hunter, P. E. (1971) The genus Coleolaelaps Berlese, 1914 (Acarina: Mesostigmata). Redia, 52, 323 - 360." type="journal article" year="1971">Costa & Hunter, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
has a posteriorly reduced dorsal shield, but is atypical in lacking greatly elongate setae on the dorsal shield and legs. These may be plesiomorphic character states in two early-derivative members of the genus.
|
||
<taxonomicName id="4C664D010023FFA2FDF72390A9DFF801" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[593,767,1953,1979]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="329" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B912EA900023FFA2FDF72390A9DFF801" box="[593,767,1953,1979]" italics="true" pageId="4" pageNumber="330">Acantholaelaps</emphasis>
|
||
</taxonomicName>
|
||
shares some derived character states with
|
||
<taxonomicName id="4C664D010023FFA2FB712390AE4BF801" box="[1239,1387,1953,1979]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900023FFA2FB712390AE4BF801" box="[1239,1387,1953,1979]" italics="true" pageId="4" pageNumber="330">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
, but has a normal oval-shaped dorsal shield similar to that of
|
||
<taxonomicName id="4C664D010023FFA2FCBE23F4A89AF865" authorityName="Costa & Hunter" authorityYear="1971" box="[792,954,1989,2015]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="330" phylum="Arthropoda" rank="species" species="abnormalis">
|
||
<emphasis id="B912EA900023FFA2FCBE23F4A80BF865" box="[792,811,1989,2015]" italics="true" pageId="4" pageNumber="330">C</emphasis>
|
||
.
|
||
<emphasis id="B912EA900023FFA2FC9C23F4A89AF865" box="[826,954,1989,2015]" italics="true" pageId="4" pageNumber="330">abnormalis</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C664D010023FFA2FB9A23F4AFCAF865" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[1084,1258,1989,2015]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="329" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B912EA900023FFA2FB9A23F4AFCAF865" box="[1084,1258,1989,2015]" italics="true" pageId="4" pageNumber="330">Acantholaelaps</emphasis>
|
||
</taxonomicName>
|
||
differs from all species of
|
||
<taxonomicName id="4C664D010022FFA3FEB624A8AA84FF09" box="[272,420,153,179]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900022FFA3FEB624A8AA84FF09" box="[272,420,153,179]" italics="true" pageId="5" pageNumber="331">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
in the presence of thick blunt setae on tarsus II, modified tarsal claws, the relative length of the legs (length of leg I longer than leg III vs. shorter than leg III in
|
||
<taxonomicName id="4C664D010022FFA3FC6E248CAF7CFF6D" box="[968,1116,189,215]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900022FFA3FC6E248CAF7CFF6D" box="[968,1116,189,215]" italics="true" pageId="5" pageNumber="331">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
), and a short post-anal seta. The new genus is compared with other morphologically similar hypoaspidine genera associated with scarab beetles in
|
||
<tableCitation id="C6E403390022FFA3FF122534AA08FEA5" box="[180,296,261,287]" captionStart="TABLE 1" captionStartId="20.[151,239,501,527]" captionTargetPageId="20" captionText="TABLE 1. Diagnostic characters for eight genera of Laelapidae associated with scarab beetles." httpUri="http://table.plazi.org/id/DF19660A0033FFB2FF3125C4AFACFDB5" pageId="5" pageNumber="331" tableUuid="DF19660A0033FFB2FF3125C4AFACFDB5">Table 1. A</tableCitation>
|
||
full analysis of these character states must await the description of more beetle-associated species in genera similar to
|
||
<taxonomicName id="4C664D010022FFA3FEFC2518AACEFEF9" box="[346,494,297,323]" class="Arachnida" family="Laelapidae" genus="Coleolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900022FFA3FEFC2518AACEFEF9" box="[346,494,297,323]" italics="true" pageId="5" pageNumber="331">Coleolaelaps</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C664D010022FFA3FE5A2518A9B4FEF9" authorityName="Clark" authorityYear="2012" box="[508,660,297,323]" class="Arachnida" family="Laelapidae" genus="Mumulaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912EA900022FFA3FE5A2518A9B4FEF9" box="[508,660,297,323]" italics="true" pageId="5" pageNumber="331">Mumulaelaps</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C664D010022FFA3FD742518A8A0FEF9" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[722,896,297,323]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="329" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B912EA900022FFA3FD742518A8A0FEF9" box="[722,896,297,323]" italics="true" pageId="5" pageNumber="331">Acantholaelaps</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C65090022FFA3FF61257CAA30FE31" pageId="5" pageNumber="331" type="etymology">
|
||
<paragraph id="8BD936820022FFA3FF61257CAA30FE31" blockId="5.[151,1437,153,395]" pageId="5" pageNumber="331">
|
||
<emphasis id="B912EA900022FFA3FF61257CAA68FEDD" bold="true" box="[199,328,333,359]" pageId="5" pageNumber="331">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName id="4C664D010022FFA3FE6F257CA957FEDD" authority="Joharchi & Halliday & Tolstikov & Trach, 2019" authorityName="Joharchi & Halliday & Tolstikov & Trach" authorityYear="2019" box="[457,631,333,359]" class="Arachnida" family="Laelapidae" genus="Acantholaelaps" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="329" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B912EA900022FFA3FE6F257CA957FEDD" box="[457,631,333,359]" italics="true" pageId="5" pageNumber="331">Acantholaelaps</emphasis>
|
||
</taxonomicName>
|
||
(Latin
|
||
<emphasis id="B912EA900022FFA3FD61257CA804FEDD" box="[711,804,333,359]" italics="true" pageId="5" pageNumber="331">Acantho</emphasis>
|
||
, spur) refers to the basal spur on the tarsal claws. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |