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<document id="9BC40F0BAC5474EAD64095BBB17718CB" ID-CLB-Dataset="22248" ID-DOI="10.5852/ejt.2013.51" ID-GBIF-Dataset="a5c53b3d-2f01-4805-8cb0-2c788edc9737" ID-ISSN="2118-9773" ID-Zenodo-Dep="3820535" ID-ZooBank="D20CE35E-08CF-41EE-9700-ABDEA6F4413E" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589220100437" checkinUser="carolina" docAuthor="Blaimer, Bonnie B. &amp; Fisher, Brian L." docDate="2013" docId="03BEDE7EFFE0FF955B3A0501FB22FC92" docLanguage="en" docName="ejt-2013-51.pdf.imf" docOrigin="European Journal of Taxonomy 51" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Crematogaster alafara Blaimer &amp; Fisher 2013, sp. nov." docType="treatment" docUuid="957DBC9B-0B2D-4ABF-B179-E18F67603927" docUuidSource="ZooBank" docVersion="12" lastPageNumber="20" masterDocId="FF87A606FFF0FF8659140771FFA1FFC2" masterDocTitle="Taxonomy of the Crematogaster degeeri-species-assemblage in the Malagasy region (Hymenoptera: Formicidae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="17" updateTime="1698839182804" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0FDF6201F082E5BFDB0E4600C39F97AF">Taxonomy of the Crematogaster degeeri-species-assemblage in the Malagasy region (Hymenoptera: Formicidae)</mods:title>
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<mods:name id="536ADB84F64307FC5CB5F6A7FA836C2B" type="personal">
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<mods:namePart id="2029A3F947DA1EEC251A8320F5159773">Blaimer, Bonnie B.</mods:namePart>
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<mods:affiliation id="654D5C78B857207FD05F54B2C144797E">Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA E-mail: bonnieblaimer @ gmail. com (corresponding author) &amp; Department of Entomology, California Academy of Sciences, San Francisco, California, USA &amp; urn: lsid: zoobank. org: author: 2 ADA 98 B 1 - 6 FEC- 446 E-B 056 - 9244 F 417 F 0 BC</mods:affiliation>
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<mods:namePart id="7F8651E37E635C2BE98846F4F38CDDAB">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="BFD0D4AC20FF73E1D67562F63BAD3E91">Department of Entomology, California Academy of Sciences, San Francisco, California, USA &amp; urn: lsid: zoobank. org: author: 7 CFCBBE 5 - 5443 - 4 F 94 - B 2 AC- 04 BFD 4 D 520 F 0</mods:affiliation>
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<mods:title id="DF2CEC682ACC46AA18CCE594CA68F337">European Journal of Taxonomy</mods:title>
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<mods:date id="5772BAB87B1F5DE121AB7E3973834D2E">2013</mods:date>
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<mods:number id="6569CB64590876EAB68B5A4A42C6638C">2013-08-02</mods:number>
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<treatment id="03BEDE7EFFE0FF955B3A0501FB22FC92" ID-DOI="http://doi.org/10.5281/zenodo.3843891" ID-GBIF-Taxon="164044899" ID-Zenodo-Dep="3843891" LSID="urn:lsid:zoobank.org:act:957DBC9B-0B2D-4ABF-B179-E18F67603927" httpUri="http://treatment.plazi.org/id/03BEDE7EFFE0FF955B3A0501FB22FC92" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="C30D3CE3FFE0FF965B3A0501FBA7FD48" box="[558,1030,623,650]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BA86F68FFE0FF965B3A0501FBA7FD48" blockId="16.[365,1223,623,727]" box="[558,1030,623,650]" pageId="16" pageNumber="17">
<heading id="D0E0D804FFE0FF965B3A0501FBA7FD48" box="[558,1030,623,650]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="4C1714EBFFE0FF965B3A0501FC04FD48" authority="Blaimer" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[558,933,623,650]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE0FF965B3A0501FC9EFD48" bold="true" box="[558,831,623,650]" italics="true" pageId="16" pageNumber="17">Crematogaster alafara</emphasis>
Blaimer
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF965AB80501FBA7FD48" box="[940,1030,624,650]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FF86636AFFE0FF96587905E7FB66FD73" box="[365,1223,662,689]" pageId="16" pageNumber="17">
urn:lsid:zoobank.org:act:
<uuid id="FFB155BDFFE0FF965B9F05E7FB66FD73" box="[651,1223,662,689]" pageId="16" pageNumber="17">957DBC9B-0B2D-4ABF-B179-E18F67603927</uuid>
</uri>
</paragraph>
<paragraph id="8BA86F68FFE0FF965BE305CCFC9DFD15" blockId="16.[365,1223,623,727]" box="[759,828,701,727]" pageId="16" pageNumber="17">
<figureCitation id="132C73EDFFE0FF965BE305CCFC9DFD15" box="[759,828,701,727]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[213,1378,276,1881]" captionTargetId="figure@18.[206,1382,265,1890]" captionTargetPageId="18" captionText="Fig. 6. Crematogaster alafara sp. nov., worker, queen and distribution. A-C. Worker (CASENT0492837). A. Full-face. B. Profile. C. Dorsal. D. Species distribution. E-G. Queen (CASENT0050441). E. Profile. F. Full-face. G. Dorsal." figureDoi="http://doi.org/10.5281/zenodo.3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE0FF9659A90473FCD0FB9E" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BA86F68FFE0FF9659A90473FE97FCDE" blockId="16.[189,310,770,796]" box="[189,310,770,796]" pageId="16" pageNumber="17">
<heading id="D0E0D804FFE0FF9659A90473FE97FCDE" bold="true" box="[189,310,770,796]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B963B37AFFE0FF9659A90473FE97FCDE" bold="true" box="[189,310,770,796]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A90445FCD0FB9E" blockId="16.[189,1399,820,1116]" pageId="16" pageNumber="17">
Workers of
<taxonomicName id="4C1714EBFFE0FF9658580445FDFBFC8C" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[332,602,820,846]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE0FF9658580445FDFBFC8C" box="[332,602,820,846]" italics="true" pageId="16" pageNumber="17">Crematogaster alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF965B710445FD63FC8C" box="[613,706,820,846]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
are fairly small (HW 0.72
<emphasis id="B963B37AFFE0FF965D170445FBACFC8F" box="[1027,1037,820,845]" italics="true" pageId="16" pageNumber="17">-</emphasis>
0.88, WL 0.79
<emphasis id="B963B37AFFE0FF965DAE0445FB65FC8F" box="[1210,1220,820,845]" italics="true" pageId="16" pageNumber="17">-</emphasis>
0.91) and have short to medium-sized propodeal spines (SPI 0.06
<emphasis id="B963B37AFFE0FF965A0C042AFC83FCB6" box="[792,802,859,884]" italics="true" pageId="16" pageNumber="17">-</emphasis>
0.10). The characteristic promesonotal characters, with the propodeum usually distinctly set off from the promesonotum in lateral view (as
<figureCitation id="132C73EDFFE0FF965DA904F0FAB6FC5E" box="[1213,1303,897,924]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[213,1378,276,1881]" captionTargetId="figure@18.[206,1382,265,1890]" captionTargetPageId="18" captionText="Fig. 6. Crematogaster alafara sp. nov., worker, queen and distribution. A-C. Worker (CASENT0492837). A. Full-face. B. Profile. C. Dorsal. D. Species distribution. E-G. Queen (CASENT0050441). E. Profile. F. Full-face. G. Dorsal." figureDoi="http://doi.org/10.5281/zenodo.3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="16" pageNumber="17">Fig. 6B</figureCitation>
), render this species moderately easy to recognize from other species in the
<taxonomicName id="4C1714EBFFE0FF965ADE04D9FBE2FC03" authorityName="Forel" authorityYear="1886" box="[970,1091,935,962]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="degeeri">
<emphasis id="B963B37AFFE0FF965ADE04D9FBE2FC03" box="[970,1091,935,962]" italics="true" pageId="16" pageNumber="17">C. degeeri</emphasis>
</taxonomicName>
-assemblage. Possibly this species could be confused with
<taxonomicName id="4C1714EBFFE0FF965B2504BFFD61FC2A" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[561,704,974,1000]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="malahelo" status="sp. nov.">
<emphasis id="B963B37AFFE0FF965B2504BFFD61FC2A" box="[561,704,974,1000]" italics="true" pageId="16" pageNumber="17">C. malahelo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF965BD304BFFC84FC2A" box="[711,805,974,1000]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
; the latter, however, bears sharp lateral margins on the pronotum (absent in
<taxonomicName id="4C1714EBFFE0FF9658CD0484FDF1FBCC" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[473,592,1012,1039]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE0FF9658CD0484FDF1FBCC" box="[473,592,1012,1039]" italics="true" pageId="16" pageNumber="17">C. alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF965B430484FD13FBCD" box="[599,690,1013,1039]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
) and has shorter, more triangular spines (SPI 0.03
<emphasis id="B963B37AFFE0FF965DE30484FAA0FBCC" box="[1271,1281,1013,1038]" italics="true" pageId="16" pageNumber="17">-</emphasis>
0.07) than
<taxonomicName id="4C1714EBFFE0FF9659A9036AFE97FBF7" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[189,310,1051,1077]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE0FF9659A9036AFE97FBF7" box="[189,310,1051,1077]" italics="true" pageId="16" pageNumber="17">C. alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF96582A036AFE34FBF7" box="[318,405,1051,1077]" pageId="16" pageNumber="17" rank="species">sp. nov</taxonomicNameLabel>
.. Mean genetic distance in the barcode region of
<taxonomicName id="4C1714EBFFE0FF965ACF036AFBF5FBF7" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[987,1108,1051,1077]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE0FF965ACF036AFBF5FBF7" box="[987,1108,1051,1077]" italics="true" pageId="16" pageNumber="17">C. alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE0FF965D48036AFB16FBF7" box="[1116,1207,1051,1077]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
to other species within the
<taxonomicName id="4C1714EBFFE0FF96582E0333FE14FB99" authorityName="Forel" authorityYear="1886" box="[314,437,1089,1116]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="degeeri">
<emphasis id="B963B37AFFE0FF96582E0333FE14FB99" box="[314,437,1089,1116]" italics="true" pageId="16" pageNumber="17">C. degeeri</emphasis>
</taxonomicName>
-assemblage is 15.7
<emphasis id="B963B37AFFE0FF965B8F0333FD04FB99" box="[667,677,1090,1115]" italics="true" pageId="16" pageNumber="17">-</emphasis>
22.2 % (
<tableCitation id="C6955AD3FFE0FF965A130330FCC0FB9E" box="[775,865,1089,1116]" captionStart="Table 3" captionStartId="8.[189,258,268,294]" captionTargetId="graphics@8.[189,1403,394,1151]" captionTargetPageId="8" captionText="Table 3. Summary of Tamura-Nei distance matrix. Divergences of cytochrome oxidase I sequences, as estimated under the Tamura-Nei Model (Tamura &amp; Nei 1993) within the software MEGA5 (Tamura et al. 2011). Bold font emphasizes highest levels of divergence within and between species." httpUri="http://table.plazi.org/id/DF683FE0FFF8FF8E59A9067DFB51FEAE" pageId="16" pageNumber="17" tableUuid="DF683FE0FFF8FF8E59A9067DFB51FEAE">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE0FF9659A903F6FAD6FB3E" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BA86F68FFE0FF9659A903F6FEE5FB63" blockId="16.[189,324,1159,1185]" box="[189,324,1159,1185]" pageId="16" pageNumber="17">
<heading id="D0E0D804FFE0FF9659A903F6FEE5FB63" bold="true" box="[189,324,1159,1185]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B963B37AFFE0FF9659A903F6FEE5FB63" bold="true" box="[189,324,1159,1185]" pageId="16" pageNumber="17">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A903C9FAD6FB3E" blockId="16.[189,1399,1208,1276]" pageId="16" pageNumber="17">
The Malagasy word
<taxonomicName id="4C1714EBFFE0FF9658BB03C9FDA4FB10" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[431,517,1208,1234]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alafara">
<emphasis id="B963B37AFFE0FF9658BB03C9FDA4FB10" box="[431,517,1208,1234]" italics="true" pageId="16" pageNumber="17">alafara</emphasis>
</taxonomicName>
means “last forest”, alluding to the restricted distribution of this species as a reminder of forest conservation concerns in
<collectingCountry id="F3002FF8FFE0FF965BC10393FCC4FB3E" box="[725,869,1250,1276]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
. This name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE0FF9659A90233FD33F915" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BA86F68FFE0FF9659A90233FE32FA9D" blockId="16.[189,1399,1346,1529]" box="[189,403,1346,1375]" pageId="16" pageNumber="17">
<emphasis id="B963B37AFFE0FF9659A90233FE32FA9D" bold="true" box="[189,403,1346,1375]" pageId="16" pageNumber="17">Type specimens</emphasis>
</paragraph>
<materialsCitation id="3B7F6535FFE0FF9659FE021BFB6FFA3A" ID-GBIF-Occurrence="2612042306" collectingDate="2003-03-31" collectionCode="CASC" collectorName="Fisher" country="Madagascar" county="Sahamaloatra" elevation="1150" latitude="-21.236666" location="P. N. Ranomafana" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" pageId="16" pageNumber="17" specimenCode="CASENT0492837" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<paragraph id="8BA86F68FFE0FF9659FE021BFEFDFA46" blockId="16.[189,1399,1346,1529]" box="[234,348,1386,1412]" pageId="16" pageNumber="17">
<emphasis id="B963B37AFFE0FF9659FE021BFEFDFA46" bold="true" box="[234,348,1386,1412]" pageId="16" pageNumber="17">
<typeStatus id="54ACD1CAFFE0FF9659FE021BFEFDFA46" box="[234,348,1386,1412]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A902E0FB6FFA3A" blockId="16.[189,1399,1346,1529]" pageId="16" pageNumber="17">
<specimenCount id="9D11A4E1FFE0FF9659A902E0FEB8FA69" box="[189,281,1425,1451]" pageId="16" pageNumber="17" type="worker">Worker</specimenCount>
: pinned,
<specimenCode id="DBB1C713FFE0FF96589D02E0FDC4FA69" box="[393,613,1425,1451]" pageId="16" pageNumber="17">CASENT0492837</specimenCode>
, BLF08619, ex rotten log [imaged on AntWeb]; original locality label: MADGR:
<location id="8EC839B3FFE0FF96588C02C9FDCFFA13" LSID="urn:lsid:plazi:treatment:03BEDE7EFFE0FF955B3A0501FB22FC92:8EC839B3FFE0FF96588C02C9FDCFFA13" box="[408,622,1463,1490]" county="Sahamaloatra" latitude="-21.236666" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" name="P. N. Ranomafana" pageId="16" pageNumber="17">P.N. Ranomafana</location>
,
<collectingCounty id="62C917E4FFE0FF965B9402C6FC82FA13" box="[640,803,1463,1489]" pageId="16" pageNumber="17">Sahamaloatra</collectingCounty>
,
<locationDeviation id="9A1A764AFFE0FF965A2002C6FB12FA13" box="[820,1203,1463,1490]" location="Ranomafana" pageId="16" pageNumber="17">
<quantity id="4CEFC28DFFE0FF965A2002C6FC2AFA13" box="[820,907,1463,1489]" metricMagnitude="3" metricUnit="m" metricValue="6.6" pageId="16" pageNumber="17" unit="km" value="6.6">6.6 km</quantity>
310° NW Ranomafana
</locationDeviation>
,
<date id="FFA949A8FFE0FF965DD102C6FAD0FA13" box="[1221,1393,1463,1490]" pageId="16" pageNumber="17" value="2003-03-31">
<collectingDate id="EFEDB040FFE0FF965DD102C6FAD0FA13" box="[1221,1393,1463,1490]" pageId="16" pageNumber="17" value="2003-03-31">31 Mar. 2003</collectingDate>
</date>
,
<geoCoordinate id="EE2309AFFFE0FF9659A902AFFE92FA3A" box="[189,307,1502,1528]" degrees="21" direction="south" minutes="14.2" orientation="latitude" pageId="16" pageNumber="17" precision="92" value="-21.236666">21º14.2S</geoCoordinate>
,
<geoCoordinate id="EE2309AFFFE0FF96582B02AFFE16FA3A" box="[319,439,1502,1528]" degrees="47" direction="east" minutes="23.8" orientation="longitude" pageId="16" pageNumber="17" precision="92" value="47.396667">47º23.8E</geoCoordinate>
,
<quantity id="4CEFC28DFFE0FF9658D002AFFDBFFA3A" box="[452,542,1502,1529]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="17" unit="m" value="1150.0">
<elevation id="003A885BFFE0FF9658D002AFFDBFFA3A" box="[452,542,1502,1529]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="17" unit="m" value="1150.0">1150 m</elevation>
</quantity>
,
<collectorName id="26E20ABEFFE0FF965B3E02AFFDD2FA3A" box="[554,627,1502,1528]" pageId="16" pageNumber="17">Fisher</collectorName>
<emphasis id="B963B37AFFE0FF965B6F02AEFD14FA3A" box="[635,693,1502,1528]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, BLF# (Brian L. Fisher); deposited at
<collectionCode id="ED06F7ADFFE0FF965D6E02AFFB6BFA3A" box="[1146,1226,1502,1528]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="16" pageNumber="17">CASC</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BA86F68FFE0FF9659FE0152FEC6F9FF" blockId="16.[189,1399,1571,1751]" box="[234,359,1571,1597]" pageId="16" pageNumber="17">
<materialsCitation id="3B7F6535FFE0FF9659FE0152FEC6F9FF" ID-GBIF-Occurrence="4433204303" box="[234,359,1571,1597]" collectingDate="2003-03-31" collectionCode="SAMC" collectorName="Fisher" county="Sahamaloatra" elevation="1150" latitude="-21.236666" location="P. N. Ranomafana" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" pageId="16" pageNumber="17" specimenCode="CASENT0317785" specimenCount="4" specimenCount-worker="4" typeStatus="paratype">
<emphasis id="B963B37AFFE0FF9659FE0152FEC6F9FF" bold="true" box="[234,359,1571,1597]" pageId="16" pageNumber="17">
<typeStatus id="54ACD1CAFFE0FF9659FE0152FEC6F9FF" box="[234,359,1571,1597]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A90138FD33F915" blockId="16.[189,1399,1571,1751]" pageId="16" pageNumber="17">
<specimenCount id="9D11A4E1FFE0FF9659A90138FE99F9A1" box="[189,312,1609,1635]" pageId="16" pageNumber="17" type="worker">4 workers</specimenCount>
, pinned, #1 and #2 same collection and locality data as holotype. #1:
<specimenCode id="DBB1C713FFE0FF965D820138FAD3F9A6" box="[1174,1394,1609,1636]" pageId="16" pageNumber="17">CASENT0317785</specimenCode>
, deposited at
<collectionCode id="ED06F7ADFFE0FF9658480101FE12F948" box="[348,435,1648,1674]" collectionName="South Africa, Cape Town, Iziko Museum of Capetown (formerly South African Museum)" pageId="16" pageNumber="17">SAMC</collectionCode>
.
<materialsCitation id="3B7F6535FFE0FF9658D70101FC42F948" ID-GBIF-Occurrence="3761959310" box="[451,995,1648,1674]" collectingCountry="Madagascar" collectingDate="2003-03-31" collectionCode="MHNG" collectorName="Fisher" country="Madagascar" county="Sahamaloatra" elevation="1150" latitude="-21.236666" location="P. N. Ranomafana" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" pageId="16" pageNumber="17" specimenCode="CASENT0317786" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">
#2:
<specimenCode id="DBB1C713FFE0FF9658E20101FD70F948" box="[502,721,1648,1674]" pageId="16" pageNumber="17">CASENT0317786</specimenCode>
, deposited at
<collectionCode id="ED06F7ADFFE0FF965A960101FC41F948" box="[898,992,1648,1674]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="16" pageNumber="17">MHNG</collectionCode>
.
</materialsCitation>
<materialsCitation id="3B7F6535FFE0FF965AE40101FAE8F972" ID-GBIF-Occurrence="3761959323" collectingDate="2003-03-31" collectionCode="MCZC" collectorName="Fisher" county="Sahamaloatra" elevation="1150" latitude="-21.236666" location="P. N. Ranomafana" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" pageId="16" pageNumber="17" specimenCode="CASENT0317787" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
#3 and #4: BLF08628, ex dead twig above ground, remaining data same as holotype. #3:
<specimenCode id="DBB1C713FFE0FF965A7B01E7FBEBF972" box="[879,1098,1686,1712]" pageId="16" pageNumber="17">CASENT0317787</specimenCode>
, deposited at
<collectionCode id="ED06F7ADFFE0FF965DFA01E7FAE4F972" box="[1262,1349,1686,1712]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="16" pageNumber="17">MCZC</collectionCode>
.
</materialsCitation>
<materialsCitation id="3B7F6535FFE0FF965C4501E7FD33F915" ID-GBIF-Occurrence="3761959334" collectingDate="2003-03-31" collectionCode="UCDC" collectorName="Fisher" county="Sahamaloatra" elevation="1150" latitude="-21.236666" location="P. N. Ranomafana" longLatPrecision="126" longitude="47.396667" municipality="Ranomafana" pageId="16" pageNumber="17" specimenCode="CASENT0317788" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
#4:
<specimenCode id="DBB1C713FFE0FF9659A901CCFE38F915" box="[189,409,1725,1751]" pageId="16" pageNumber="17">CASENT0317788</specimenCode>
, deposited at
<collectionCode id="ED06F7ADFFE0FF965B2D01CCFD2FF915" box="[569,654,1725,1751]" collectionName="USA, California, Davis, University of California, R.M. Bohart Museum of Entomology" pageId="16" pageNumber="17">UCDC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE0FF9659A90073FE03F8B6" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BA86F68FFE0FF9659A90073FEFCF8D9" blockId="16.[189,349,1793,1820]" box="[189,349,1793,1820]" pageId="16" pageNumber="17">
<heading id="D0E0D804FFE0FF9659A90073FEFCF8D9" bold="true" box="[189,349,1793,1820]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B963B37AFFE0FF9659A90073FEFCF8D9" bold="true" box="[189,349,1793,1820]" pageId="16" pageNumber="17">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A90045FE03F8B6" blockId="16.[189,1399,1843,1909]" pageId="16" pageNumber="17">
<collectingCountry id="F3002FF8FFE0FF9659A90045FE33F88C" box="[189,402,1844,1870]" name="Madagascar" pageId="16" pageNumber="17">MADAGASCAR</collectingCountry>
:
<collectingRegion id="49D3A18AFFE0FF9658890042FD99F88F" box="[413,568,1843,1869]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="17">Fianarantsoa</collectingRegion>
:
<location id="8EC839B3FFE0FF965B560045FCB0F88F" LSID="urn:lsid:plazi:treatment:03BEDE7EFFE0FF955B3A0501FB22FC92:8EC839B3FFE0FF965B560045FCB0F88F" box="[578,785,1843,1870]" country="Madagascar" latitude="-21.23667" longLatPrecision="1" longitude="47.39667" name="P. N. Ranomafana" pageId="16" pageNumber="17" stateProvince="Fianarantsoa">P.N. Ranomafana</location>
,
<location id="8EC839B3FFE0FF965A090042FBABF88F" LSID="urn:lsid:plazi:treatment:03BEDE7EFFE0FF955B3A0501FB22FC92:8EC839B3FFE0FF965A090042FBABF88F" box="[797,1034,1843,1869]" country="Madagascar" latitude="-21.23667" longLatPrecision="1" longitude="47.39667" name="Sahamaloatra River" pageId="16" pageNumber="17" stateProvince="Fianarantsoa">Sahamaloatra River</location>
:
<geoCoordinate id="EE2309AFFFE0FF965D010042FB2EF88F" box="[1045,1167,1843,1869]" degrees="21.23667" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-21.23667">-21.23667</geoCoordinate>
,
<geoCoordinate id="EE2309AFFFE0FF965D8F0042FAADF88F" box="[1179,1292,1843,1869]" degrees="47.39667" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="47.39667">47.39667</geoCoordinate>
,
<quantity id="4CEFC28DFFE0FF965C030042FAD0F88C" box="[1303,1393,1843,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="17" unit="m" value="1150.0">
<elevation id="003A885BFFE0FF965C030042FAD0F88C" box="[1303,1393,1843,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="17" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, montane rainforest.
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE0FF9659A900EEFC80F829" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BA86F68FFE0FF9659A900EEFE58F87B" blockId="16.[189,505,1951,1977]" box="[189,505,1951,1977]" pageId="16" pageNumber="17">
<heading id="D0E0D804FFE0FF9659A900EEFE58F87B" bold="true" box="[189,505,1951,1977]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B963B37AFFE0FF9659A900EEFE58F87B" bold="true" box="[189,505,1951,1977]" pageId="16" pageNumber="17">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE0FF9659A900A0FC80F829" blockId="16.[189,801,2001,2027]" box="[189,801,2001,2027]" pageId="16" pageNumber="17">
(
<collectionCode id="ED06F7ADFFE0FF9659D100A0FEB7F829" box="[197,278,2001,2027]" pageId="16" pageNumber="17">CASC</collectionCode>
,
<collectionCode id="ED06F7ADFFE0FF96583600A0FED9F829" box="[290,376,2001,2027]" pageId="16" pageNumber="17">PSWC</collectionCode>
): refer to Supplementary Material 1.
</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE1FF9559A9067AFDFBFD8C" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
<paragraph id="8BA86F68FFE1FF9759A9067AFEBEFEE7" blockId="17.[189,287,267,293]" box="[189,287,267,293]" pageId="17" pageNumber="18">
<heading id="D0E0D804FFE1FF9759A9067AFEBEFEE7" bold="true" box="[189,287,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B963B37AFFE1FF9759A9067AFEBEFEE7" bold="true" box="[189,287,267,293]" pageId="17" pageNumber="18">Worker</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759FE064AFE03FE97" blockId="17.[189,1399,315,525]" box="[234,418,315,341]" pageId="17" pageNumber="18">
<emphasis id="B963B37AFFE1FF9759FE064AFE03FE97" bold="true" box="[234,418,315,341]" pageId="17" pageNumber="18">Measurements</emphasis>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A90611FCB1FDCE" blockId="17.[189,1399,315,525]" pageId="17" pageNumber="18">
(n = 12) [
<typeStatus id="54ACD1CAFFE1FF975831062EFE33FEB8" box="[293,402,351,378]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
] HW 0.72
<emphasis id="B963B37AFFE1FF975B1C0611FDB3FEBB" box="[520,530,352,377]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.88 [0.83]; HL 0.69
<emphasis id="B963B37AFFE1FF975BEB0611FCA8FEBB" box="[767,777,352,377]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.80 [0.78]; EL 0.15
<emphasis id="B963B37AFFE1FF975AE60611FC5DFEBB" box="[1010,1020,352,377]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.19 [0.17]; SL 0.54
<emphasis id="B963B37AFFE1FF975DF00611FB4FFEBB" box="[1252,1262,352,377]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.64 [0.61]; WL 0.79
<emphasis id="B963B37AFFE1FF97583206F4FE91FE5C" box="[294,304,389,414]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.91 [0.90]; SPL 0.05
<emphasis id="B963B37AFFE1FF975B2706F4FD9CFE5C" box="[563,573,389,414]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.08 [0.06]; PTH 0.17
<emphasis id="B963B37AFFE1FF975A5206F4FCF1FE5C" box="[838,848,389,414]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.20 [0.20]; PTL 0.22
<emphasis id="B963B37AFFE1FF975D4006F4FBFFFE5C" box="[1108,1118,389,414]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.26 [0.26]; PTW 0.29
<emphasis id="B963B37AFFE1FF975C7906F4FAD6FE5C" box="[1389,1399,389,414]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.35 [0.35]; PPL 0.13
<emphasis id="B963B37AFFE1FF9758D406DBFE6BFE01" box="[448,458,426,451]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.18 [0.16]; PPW 0.23
<emphasis id="B963B37AFFE1FF975BC206DBFD41FE01" box="[726,736,426,451]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.28 [0.25]; LHT 0.56
<emphasis id="B963B37AFFE1FF975AFE06DBFC55FE01" box="[1002,1012,426,451]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.67 [0.66]; CI 1.02
<emphasis id="B963B37AFFE1FF975DF606DBFB4DFE01" box="[1250,1260,426,451]" italics="true" pageId="17" pageNumber="18">-</emphasis>
1.11 [1.06]; OI 0.20
<emphasis id="B963B37AFFE1FF97580306BFFE80FE25" box="[279,289,462,487]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.25 [0.21]; SI 0.73
<emphasis id="B963B37AFFE1FF975B1206BFFDB1FE25" box="[518,528,462,487]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.83 [0.77]; SPI 0.06
<emphasis id="B963B37AFFE1FF975A1206BFFCB1FE25" box="[774,784,462,487]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.10 [0.07]; PTHI 0.71
<emphasis id="B963B37AFFE1FF975D0806BFFB87FE25" box="[1052,1062,462,487]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.81 [0.79]; PTWI 1.18
<emphasis id="B963B37AFFE1FF975C2D06BFFAE2FE25" box="[1337,1347,462,487]" italics="true" pageId="17" pageNumber="18">-</emphasis>
1.47 [1.36]; PPI 1.56
<emphasis id="B963B37AFFE1FF97586C0682FE23FDCE" box="[376,386,499,524]" italics="true" pageId="17" pageNumber="18">-</emphasis>
1.97 [1.59]; LBI 1.30
<emphasis id="B963B37AFFE1FF975B690682FD26FDCE" box="[637,647,499,524]" italics="true" pageId="17" pageNumber="18">-</emphasis>
1.47 [1.38].
</paragraph>
<paragraph id="8BA86F68FFE1FF9759FE0544FEDAFD8D" blockId="17.[189,837,565,628]" box="[234,379,565,591]" pageId="17" pageNumber="18">
<emphasis id="B963B37AFFE1FF9759FE0544FEDAFD8D" bold="true" box="[234,379,565,591]" pageId="17" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A9052BFCE4FDB6" blockId="17.[189,837,565,628]" box="[189,837,602,628]" pageId="17" pageNumber="18">
(
<figureCitation id="132C73EDFFE1FF9759D2052BFE9EFDB6" box="[198,319,602,628]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[213,1378,276,1881]" captionTargetId="figure@18.[206,1382,265,1890]" captionTargetPageId="18" captionText="Fig. 6. Crematogaster alafara sp. nov., worker, queen and distribution. A-C. Worker (CASENT0492837). A. Full-face. B. Profile. C. Dorsal. D. Species distribution. E-G. Queen (CASENT0050441). E. Profile. F. Full-face. G. Dorsal." figureDoi="http://doi.org/10.5281/zenodo.3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="17" pageNumber="18">
Fig. 6A
<emphasis id="B963B37AFFE1FF975834052BFE8BFDB1" box="[288,298,602,627]" italics="true" pageId="17" pageNumber="18">-</emphasis>
C
</figureCitation>
) Small size (HW 0.72
<emphasis id="B963B37AFFE1FF975B50052BFDEFFDB1" box="[580,590,602,627]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.88, WL 0.79
<emphasis id="B963B37AFFE1FF975BE3052BFCA0FDB1" box="[759,769,602,627]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.91).
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A905EDFAABFCE6" blockId="17.[189,1399,668,805]" pageId="17" pageNumber="18">
Masticatory margin of mandibles with four teeth; head shape quadrate, as long, or slightly longer than wide (CI 1.02
<emphasis id="B963B37AFFE1FF97587C05B3FED3FD19" box="[360,370,706,731]" italics="true" pageId="17" pageNumber="18">-</emphasis>
1.11); posterior margin of head in full-face view laterally rounded; occipital carinae distinct; antennal scapes reaching or slightly surpassing head margin; midline of eyes situated slightly above midline of head in full-face view; eyes moderately large (OI 0.20
<emphasis id="B963B37AFFE1FF975D13047AFBB0FCE6" box="[1031,1041,779,804]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.25) and protruding.
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A9043FFD8BFBAA" blockId="17.[189,1399,845,1128]" pageId="17" pageNumber="18">
Pronotum laterally angular; promesonotal suture indistinct, promesonotum forming one plane; mesonotum usually without posterior face; laterally mesonotum carinate, sometimes ending in small denticles, but not greatly set off from propodeum; metanotal groove shallow, laterally constricted and bordered by carinae; propodeal spines short, spiniform (SPI 0.06
<emphasis id="B963B37AFFE1FF975ACD04CDFC42FC17" box="[985,995,956,981]" italics="true" pageId="17" pageNumber="18">-</emphasis>
0.10) and straight; dorsal face of propodeum distinct; posterior face of propodeum gently sloping; petiole in dorsal view greatly flared and convex, dorsolaterally rounded to carinate, without posterolateral tubercules or denticles; subpetiolar process usually a small, angular dent; postpetiole distinctly bilobed, with a narrow median impression; subpostpetiolar process absent.
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A903E0FD85FA45" blockId="17.[189,1399,1169,1415]" pageId="17" pageNumber="18">Head sculpture aciculate to faintly reticulate; sculpture otherwise mostly reticulate; face usually with &lt;six erect, longer setae, and regular appressed to decumbent pubescence; erect pilosity on promesonotum usually sparse, &lt;six long setae; otherwise promesonotum dorsally with sparse appressed to subdecumbent pubescence; petiole and postpetiole usually with a pair of short, erect setae posterolaterally, and sparse suberect, decumbent or appressed pubescence; abdominal tergites and sternites four to seven usually with moderately abundant, short, erect pilosity, and regular appressed to decumbent pubescence throughout. Color yellow-brown to brown.</paragraph>
<paragraph id="8BA86F68FFE1FF9759A902C2FEAFFA0F" blockId="17.[189,270,1459,1485]" box="[189,270,1459,1485]" pageId="17" pageNumber="18">
<heading id="D0E0D804FFE1FF9759A902C2FEAFFA0F" bold="true" box="[189,270,1459,1485]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B963B37AFFE1FF9759A902C2FEAFFA0F" bold="true" box="[189,270,1459,1485]" pageId="17" pageNumber="18">Queen</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759FE0292FE03FA3F" blockId="17.[189,1398,1507,1643]" box="[234,418,1507,1533]" pageId="17" pageNumber="18">
<emphasis id="B963B37AFFE1FF9759FE0292FE03FA3F" bold="true" box="[234,418,1507,1533]" pageId="17" pageNumber="18">Measurements</emphasis>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A90179FD1FF9A9" blockId="17.[189,1398,1507,1643]" pageId="17" pageNumber="18">(n = 1) HW 1.01, HL 1.01, EL 0.33, SL 0.64, MSNW 0.76, MSNL 0.87, WL 1.68, SPL 0.00, PTH 0.29, PTL 0.33, PTW 0.39, PPL 0.23, PPW 0.39, LHT 0.75, CI 0.99, OI 0.32, SI 0.63, MSNI 0.87, SPI 0.00, PTHI 0.87, PTWI 1.18, PPI 1.68, LBI 2.24.</paragraph>
<paragraph id="8BA86F68FFE1FF9759FE01E2FEDAF96F" blockId="17.[189,1245,1683,1747]" box="[234,379,1683,1709]" pageId="17" pageNumber="18">
<emphasis id="B963B37AFFE1FF9759FE01E2FEDAF96F" bold="true" box="[234,379,1683,1709]" pageId="17" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A901C9FB7CF910" blockId="17.[189,1245,1683,1747]" box="[189,1245,1720,1747]" pageId="17" pageNumber="18">
(
<figureCitation id="132C73EDFFE1FF9759D201C9FE9CF910" box="[198,317,1720,1746]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[213,1378,276,1881]" captionTargetId="figure@18.[206,1382,265,1890]" captionTargetPageId="18" captionText="Fig. 6. Crematogaster alafara sp. nov., worker, queen and distribution. A-C. Worker (CASENT0492837). A. Full-face. B. Profile. C. Dorsal. D. Species distribution. E-G. Queen (CASENT0050441). E. Profile. F. Full-face. G. Dorsal." figureDoi="http://doi.org/10.5281/zenodo.3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="17" pageNumber="18">
Fig. 6E
<emphasis id="B963B37AFFE1FF97580901C8FE86F910" box="[285,295,1721,1746]" italics="true" pageId="17" pageNumber="18">-</emphasis>
G
</figureCitation>
) Small queen (HW 1.01, WL 1.68). With worker characters, except as follows.
</paragraph>
<paragraph id="8BA86F68FFE1FF9759A9018AFE47F89C" blockId="17.[189,1399,1787,1886]" pageId="17" pageNumber="18">Masticatory margin of mandibles with five teeth; antennal scapes reaching level of lateral ocelli; eyes large (OI 0.33), situated below midline of head in full-face view; head elongate-rectangular (CI 0.99), posterior margin straight.</paragraph>
<paragraph id="8BA86F68FFE1FF9759A900F9FD1EF829" blockId="17.[189,1398,1927,2027]" pageId="17" pageNumber="18">Mesoscutum in dorsal view longer than wide (MSNI 0.87); dorsal face of propodeum short; propodeal tubercules or denticles absent; petiole moderately flared, postpetiole with complete impression; subpetiolar process present as minute tooth.</paragraph>
<caption id="DF683FE0FFE2FF9459A900FBFE6EF829" ID-DOI="http://doi.org/10.5281/zenodo.3825363" ID-Zenodo-Dep="3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1930,1956]" targetBox="[213,1378,276,1881]" targetPageId="18">
<paragraph id="8BA86F68FFE2FF9459A900FBFE6EF829" blockId="18.[189,1399,1930,2027]" pageId="18" pageNumber="19">
<emphasis id="B963B37AFFE2FF9459A900FBFEA6F866" bold="true" box="[189,263,1930,1956]" pageId="18" pageNumber="19">Fig. 6.</emphasis>
<taxonomicName id="4C1714EBFFE2FF94581800FAFDB4F866" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[268,533,1930,1957]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE2FF94581800FAFDB4F866" box="[268,533,1930,1957]" italics="true" pageId="18" pageNumber="19">Crematogaster alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE2FF945B0E00FAFDD4F867" box="[538,629,1931,1957]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, worker, queen and distribution.
<emphasis id="B963B37AFFE2FF945AFC00FBFB82F866" bold="true" box="[1000,1059,1930,1956]" pageId="18" pageNumber="19">A-C.</emphasis>
Worker (CASENT0492837).
<emphasis id="B963B37AFFE2FF9459A900DCFF7BF805" bold="true" box="[189,218,1965,1991]" pageId="18" pageNumber="19">A.</emphasis>
Full-face.
<emphasis id="B963B37AFFE2FF94584C00DFFED2F80A" bold="true" box="[344,371,1966,1992]" pageId="18" pageNumber="19">B.</emphasis>
Profile.
<emphasis id="B963B37AFFE2FF9458C200DCFE52F805" bold="true" box="[470,499,1965,1991]" pageId="18" pageNumber="19">C.</emphasis>
Dorsal.
<emphasis id="B963B37AFFE2FF945B4200DFFDD2F80A" bold="true" box="[598,627,1966,1992]" pageId="18" pageNumber="19">D.</emphasis>
Species distribution.
<emphasis id="B963B37AFFE2FF945A7B00DCFC0BF805" bold="true" box="[879,938,1965,1991]" pageId="18" pageNumber="19">E-G.</emphasis>
Queen (CASENT0050441).
<emphasis id="B963B37AFFE2FF945DEB00DFFABBF80A" bold="true" box="[1279,1306,1966,1992]" pageId="18" pageNumber="19">E.</emphasis>
Profile.
<emphasis id="B963B37AFFE2FF9459A900A0FF70F829" bold="true" box="[189,209,2001,2027]" pageId="18" pageNumber="19">F.</emphasis>
Full-face.
<emphasis id="B963B37AFFE2FF94584700A1FED3F828" bold="true" box="[339,370,2000,2026]" pageId="18" pageNumber="19">G.</emphasis>
Dorsal.
</paragraph>
</caption>
<paragraph id="8BA86F68FFE3FF9559A9067AFDC8FE1B" blockId="19.[189,1399,267,473]" pageId="19" pageNumber="20">Sculpture aciculate to reticulate throughout; face with six longer erect setae (but n = 1) and regular short suberect to decumbent pubescence; mesonotum with regular short, erect to suberect pubescence, interspersed with abundant, longer, erect setae; petiole and postpetiole with fairly dense, short, suberect to decumbent pilosity, and postpetiole with additional longer dorso-posterior setae; abdominal tergites and sternites four to seven with abundant, short, erect pilosity and dense appressed to decumbent pubescence. Color yellowish brown.</paragraph>
<paragraph id="8BA86F68FFE3FF9559A90573FF5CFDDE" blockId="19.[189,253,514,540]" box="[189,253,514,540]" pageId="19" pageNumber="20">
<heading id="D0E0D804FFE3FF9559A90573FF5CFDDE" bold="true" box="[189,253,514,540]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B963B37AFFE3FF9559A90573FF5CFDDE" bold="true" box="[189,253,514,540]" pageId="19" pageNumber="20">Male</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE3FF9559A90545FDFBFD8C" blockId="19.[189,602,564,591]" box="[189,602,564,591]" pageId="19" pageNumber="20">Males of this species are unknown.</paragraph>
</subSubSection>
<subSubSection id="C30D3CE3FFE3FF9559A90507FB22FC92" pageId="19" pageNumber="20" type="multiple">
<paragraph id="8BA86F68FFE3FF9559A90507FE4EFD52" blockId="19.[189,495,630,657]" box="[189,495,630,657]" pageId="19" pageNumber="20">
<heading id="D0E0D804FFE3FF9559A90507FE4EFD52" bold="true" box="[189,495,630,657]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B963B37AFFE3FF9559A90507FE4EFD52" bold="true" box="[189,495,630,657]" pageId="19" pageNumber="20">Distribution and biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA86F68FFE3FF9559A905D7FB22FC92" blockId="19.[189,1399,678,849]" pageId="19" pageNumber="20">
<collectingCountry id="F3002FF8FFE3FF9559A905D7FE33FD02" box="[189,402,678,704]" name="Madagascar" pageId="19" pageNumber="20">MADAGASCAR</collectingCountry>
.
<taxonomicName id="4C1714EBFFE3FF95588905D7FD08FD02" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[413,681,678,704]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alafara" status="sp. nov.">
<emphasis id="B963B37AFFE3FF95588905D7FD08FD02" box="[413,681,678,704]" italics="true" pageId="19" pageNumber="20">Crematogaster alafara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE3FF955BA505D7FCADFD02" box="[689,780,678,704]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
has only been collected at a few rainforest localities in central, northern and western Madagascar (
<figureCitation id="132C73EDFFE3FF955BC205BBFC90FD27" box="[726,817,714,741]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[213,1378,276,1881]" captionTargetId="figure@18.[206,1382,265,1890]" captionTargetPageId="18" captionText="Fig. 6. Crematogaster alafara sp. nov., worker, queen and distribution. A-C. Worker (CASENT0492837). A. Full-face. B. Profile. C. Dorsal. D. Species distribution. E-G. Queen (CASENT0050441). E. Profile. F. Full-face. G. Dorsal." figureDoi="http://doi.org/10.5281/zenodo.3825363" httpUri="https://zenodo.org/record/3825363/files/figure.png" pageId="19" pageNumber="20">Fig. 6D</figureCitation>
) ranging from
<quantity id="4CEFC28DFFE3FF955AF605BBFB8DFD26" box="[994,1068,714,740]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="m" value="600.0">600 m</quantity>
to
<quantity id="4CEFC28DFFE3FF955D4505BBFB0BFD26" box="[1105,1194,714,741]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="m" value="1350.0">1350 m</quantity>
in elevation. The species is known to co-occur with
<taxonomicName id="4C1714EBFFE3FF955B76059FFD40FCCA" authorityName="Forel" authorityYear="1891" box="[610,737,750,776]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="sewellii">
<emphasis id="B963B37AFFE3FF955B76059FFD40FCCA" box="[610,737,750,776]" italics="true" pageId="19" pageNumber="20">C. sewellii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1714EBFFE3FF955BE6059FFCCEFCCA" authorityName="Blaimer &amp; Fisher" authorityYear="2013" baseAuthorityName="Dalla Torre" baseAuthorityYear="1893" box="[754,879,750,776]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dentata" status="stat. nov.">
<emphasis id="B963B37AFFE3FF955BE6059FFCCEFCCA" box="[754,879,750,776]" italics="true" pageId="19" pageNumber="20">C. dentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE3FF955A6C059EFC45FCCB" box="[888,996,751,777]" pageId="19" pageNumber="20">stat. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1714EBFFE3FF955D36059FFB3DFCCA" authority="Blaimer &amp; Fisher, 2013" authorityName="Blaimer &amp; Fisher" authorityYear="2013" box="[1058,1180,750,776]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mafybe" status="sp. nov.">
<emphasis id="B963B37AFFE3FF955D36059FFB3DFCCA" box="[1058,1180,750,776]" italics="true" pageId="19" pageNumber="20">C. mafybe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2500E01FFE3FF955DB1059EFB5FFCCB" box="[1189,1278,751,777]" pageId="19" pageNumber="20" rank="species">sp. nov</taxonomicNameLabel>
.. The few collections have been made from dead twigs above ground at the lower elevation range, and from rotten logs at higher altitudes. Otherwise nothing is known of the biology of this species.
</paragraph>
</subSubSection>
</treatment>
</document>