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<document id="AC00E03D72ED04E363C03A4103DCDF6C" ID-DOI="10.11646/zootaxa.3341.1.1" ID-GBIF-Dataset="10b73727-8726-4994-a2ed-3d2a008678c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="213817" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460487960184" checkinUser="plazi" docAuthor="Yoshimura, Masashi &amp; Fisher, Brian L." docDate="2012" docId="03BDEF36F51F9404FFAB41CCDE108FCA" docLanguage="en" docName="zt03341p031.pdf" docOrigin="Zootaxa 3341 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3341.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Adetomyrma cassis, sp. nov." docType="treatment" docVersion="12" lastPageNumber="19" masterDocId="FF84974EF50D9416FF3C4249DF54881E" masterDocTitle="A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="19" updateTime="1698309519930" updateUser="plazi">
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<mods:title id="597F1A06049275C7B0084724EB376F8E">A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)</mods:title>
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<mods:namePart id="2C0D7194167DF0B26794861DD2F41879">Yoshimura, Masashi</mods:namePart>
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<mods:namePart id="0037CD9858CF3F6E67D3F3E47EE75E6D">Fisher, Brian L.</mods:namePart>
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<treatment id="03BDEF36F51F9404FFAB41CCDE108FCA" ID-DOI="http://doi.org/10.5281/zenodo.5874333" ID-GBIF-Taxon="119518773" ID-Zenodo-Dep="5874333" LSID="urn:lsid:plazi:treatment:03BDEF36F51F9404FFAB41CCDE108FCA" httpUri="http://treatment.plazi.org/id/03BDEF36F51F9404FFAB41CCDE108FCA" lastPageNumber="19" pageId="18" pageNumber="19">
<subSubSection id="C30E0DABF51F9404FFAB41CCDEB08B81" box="[151,484,901,927]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BAB5E20F51F9404FFAB41CCDEB08B81" blockId="18.[151,484,901,959]" box="[151,484,901,927]" pageId="18" pageNumber="19">
<heading id="D0E3E94CF51F9404FFAB41CCDEB08B81" bold="true" box="[151,484,901,927]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B9608232F51F9404FFAB41CCDEB08B81" bold="true" box="[151,484,901,927]" pageId="18" pageNumber="19">
<taxonomicName id="4C1425A3F51F9404FFAB41CCDE2B8B81" authority="Yoshimura &amp; Fisher 2012" box="[151,383,901,927]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cassis" status="sp. nov.">
<emphasis id="B9608232F51F9404FFAB41CCDE2B8B81" bold="true" box="[151,383,901,927]" italics="true" pageId="18" pageNumber="19">Adetomyrma cassis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2533F49F51F9404FEBA41CCDEB08B81" box="[390,484,901,927]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30E0DABF51F9404FFAB41EEDE9B8BA1" box="[151,463,935,959]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BAB5E20F51F9404FFAB41EEDE9B8BA1" blockId="18.[151,484,901,959]" box="[151,463,935,959]" pageId="18" pageNumber="19">
(
<figureCitation id="132F42A5F51F9404FFA341EEDFA18BA1" box="[159,245,935,959]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="18" pageNumber="19">Figs 14</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE3D41EEDE4B8BA1" box="[257,287,935,959]" captionStart="FIGURES 15 16" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1435,1278,1913]" captionTargetId="figure@6.[151,1436,1278,1931]" captionTargetPageId="6" captionText="FIGURES 15 16. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 15, A. cassis (CASENT 0163620: holotype); 16, A. venatrix (CASENT 0079480). 15, paramere with a needle-like projection on its posterodorsal portion; 16, paramere without needle-like projection on its posterodorsal portion." httpUri="https://zenodo.org/record/213823/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE1741EEDE1E8BA1" box="[299,330,935,959]" captionStart="FIGURES 29 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="18" pageNumber="19">32</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE6941EEDE278BA1" box="[341,371,935,959]" captionStart="FIGURES 38 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="18" pageNumber="19">41</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE4341EEDEC98BA1" box="[383,413,935,959]" captionStart="FIGURES 47 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="18" pageNumber="19">50</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE9541EEDE928BA1" box="[425,454,935,959]" captionStart="FIGURES 64 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="18" pageNumber="19">67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFAB41B9DE438C51" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BAB5E20F51F9404FFAB41B9DE438C51" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
<materialsCitation id="3B7C547DF51F9404FFAB41B9DE438C51" ID-GBIF-Occurrence="3446753329" collectingDate="2010-02-20" collectingDateMax="2010-02-24" collectingDateMin="2010-02-20" collectingMethod="Malaise traps" collectionCode="CASC" collectorName="B. L. Fisher &amp; al." country="Madagascar" elevation="450" latitude="-16.772667" location="Reserve Speciale Ambatovaky" longLatPrecision="13" longitude="49.2655" municipality="Toamasina" pageId="18" pageNumber="19" specimenCode="CASENT0163620, BLF24309" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9608232F51F9404FFAB41B9DE5F8C16" bold="true" box="[151,267,1008,1032]" pageId="18" pageNumber="19">
<typeStatus id="54AFE082F51F9404FFAB41B9DE528C16" box="[151,262,1008,1032]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D1295A9F51F9404FE2A41A6DE008C19" box="[278,340,1007,1031]" pageId="18" pageNumber="19" type="male">Male</specimenCount>
:
<specimenCode id="DBB2F65BF51F9404FE5F41A6DD618C19" box="[355,565,1007,1031]" pageId="18" pageNumber="19">CASENT0163620</specimenCode>
,
<specimenCode id="DBB2F65BF51F9404FD7841A6DD968C19" box="[580,706,1007,1031]" pageId="18" pageNumber="19">BLF24309</specimenCode>
:
<collectingCountry id="F3031EB0F51F9404FDED41A6DCCF8C19" box="[721,923,1007,1031]" name="Madagascar" pageId="18" pageNumber="19">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="6BCFC45AF51F9404FC9641A6DB728C19" box="[938,1062,1007,1031]" pageId="18" pageNumber="19">Toamasina</collectingMunicipality>
,
<location id="8ECB08FBF51F9404FB0A41A6DACC8C19" LSID="urn:lsid:plazi:treatment:03BDEF36F51F9404FFAB41CCDE108FCA:8ECB08FBF51F9404FB0A41A6DACC8C19" box="[1078,1432,1007,1031]" country="Madagascar" latitude="-16.772667" longLatPrecision="13" longitude="49.2655" municipality="Toamasina" name="Reserve Speciale Ambatovaky" pageId="18" pageNumber="19">Réserve Spéciale Ambatovaky</location>
,
<location id="8ECB08FBF51F9404FFAB465DDE358C32" LSID="urn:lsid:plazi:treatment:03BDEF36F51F9404FFAB41CCDE108FCA:8ECB08FBF51F9404FFAB465DDE358C32" box="[151,353,1044,1068]" country="Madagascar" latitude="-16.772667" longLatPrecision="13" longitude="49.2655" municipality="Toamasina" name="Sandrangato river" pageId="18" pageNumber="19">Sandrangato river</location>
,
<geoCoordinate id="EE2038E7F51F9404FE51465DDEA88C32" box="[365,508,1044,1068]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="9" value="-16.772667">16° 46.36´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51F9404FD36465DDDC88C32" box="[522,668,1044,1068]" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="9" value="49.2655">49° 15.93´E</geoCoordinate>
,
<quantity id="4CECF3C5F51F9404FD94465DDDA48C32" box="[680,752,1044,1068]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="19" unit="m" value="450.0">
<elevation id="0039B913F51F9404FD94465DDDA48C32" box="[680,752,1044,1068]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="19" unit="m" value="450.0">450 m</elevation>
</quantity>
alt., rainforest, 4
<collectingMethod id="52552637F51F9404FC85465DDB068C32" box="[953,1106,1044,1068]" pageId="18" pageNumber="19">Malaise traps</collectingMethod>
,
<date id="FFAA78E0F51F9404FB62465DDBA98C32" box="[1118,1277,1044,1068]" pageId="18" pageNumber="19" value="2010-02-20" valueMax="2010-02-24" valueMin="2010-02-20">
<collectingDate id="EFEE8108F51F9404FB62465DDBA98C32" box="[1118,1277,1044,1068]" pageId="18" pageNumber="19" value="2010-02-20" valueMax="2010-02-24" valueMin="2010-02-20">2024.ii.2010</collectingDate>
</date>
,
<collectorName id="26E13BF6F51F9404FA34465DDAD48C32" box="[1288,1408,1044,1068]" pageId="18" pageNumber="19">B.L.Fisher</collectorName>
<emphasis id="B9608232F51F9404FAB4465CDFE08C4E" italics="true" pageId="18" pageNumber="19">
et
<collectorName id="26E13BF6F51F9404FFAB4671DFE08C4E" box="[151,180,1080,1104]" pageId="18" pageNumber="19">al.</collectorName>
</emphasis>
[
<collectionCode id="ED05C6E5F51F9404FFFD467EDE448C51" box="[193,272,1079,1103]" pageId="18" pageNumber="19">CASC</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFFB4615DBEA8E56" pageId="18" pageNumber="19" type="description">
<paragraph id="8BAB5E20F51F9404FFFB4615DD4A8C6A" blockId="18.[151,1437,1007,2004]" box="[199,542,1116,1140]" pageId="18" pageNumber="19">
<emphasis id="B9608232F51F9404FFFB4615DEF38C6A" bold="true" box="[199,423,1116,1140]" pageId="18" pageNumber="19">Worker and queen</emphasis>
unknown.
</paragraph>
<paragraph id="8BAB5E20F51F9404FFFB46C9DD758CA2" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
<emphasis id="B9608232F51F9404FFFB46C9DEF18C86" bold="true" box="[199,421,1151,1176]" pageId="18" pageNumber="19">Male. Description.</emphasis>
Measurements:
<typeStatus id="54AFE082F51F9404FD5D4636DD938C89" box="[609,711,1151,1175]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
. HL 0.74, HW 1.08, SL 0.3, EL 0.43, WL 2.22, MnW 1.33, CI 144.8, SI 27.9, EI 58.1, MnI 123.8.
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54680DC518D6E" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
Eye well-developed and prominent, posterior margin not exceeding posterior margin of mid ocellus in fullface view. Distance between lateral ocellus and eye long, about 3× longer than diameter of lateral ocellus. Palpal formula 3,2? (three maxillary and two? labial: observed without dissection). Notaulus absent on mesoscutum (
<figureCitation id="132F42A5F51F9404FA494746DFE28D52" captionStart="FIGURES 38 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="18" pageNumber="19">Fig 41</figureCitation>
). Parapsidal line clearly impressed. Anterior margin of petiole as long as dorsal margin in lateral view. Subpetiolar process well-developed, with dense hairs ventrally.
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54735DEF28DC2" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
Left and right parameres narrowly overlapping on dorsal small part of basimere (
<figureCitation id="132F42A5F51F9404FB554735DB968D8A" box="[1129,1218,1404,1428]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="18" pageNumber="19">Figs 14</figureCitation>
,
<figureCitation id="132F42A5F51F9404FBEC4735DBBB8D8A" box="[1232,1263,1404,1428]" captionStart="FIGURES 15 16" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1435,1278,1913]" captionTargetId="figure@6.[151,1436,1278,1931]" captionTargetPageId="6" captionText="FIGURES 15 16. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 15, A. cassis (CASENT 0163620: holotype); 16, A. venatrix (CASENT 0079480). 15, paramere with a needle-like projection on its posterodorsal portion; 16, paramere without needle-like projection on its posterodorsal portion." httpUri="https://zenodo.org/record/213823/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
). Distinct, flattened projection present on each posterior portion of paramere with small denticles on distal and mesal portions of projection (
<figureCitation id="132F42A5F51F9404FE2B478DDE3A8DC2" box="[279,366,1476,1500]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="18" pageNumber="19">Figs 14</figureCitation>
,
<figureCitation id="132F42A5F51F9404FE46478DDECD8DC2" box="[378,409,1476,1500]" captionStart="FIGURES 15 16" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1435,1278,1913]" captionTargetId="figure@6.[151,1436,1278,1931]" captionTargetPageId="6" captionText="FIGURES 15 16. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 15, A. cassis (CASENT 0163620: holotype); 16, A. venatrix (CASENT 0079480). 15, paramere with a needle-like projection on its posterodorsal portion; 16, paramere without needle-like projection on its posterodorsal portion." httpUri="https://zenodo.org/record/213823/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF547A1DBEA8E56" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
Hair on compound eyes long, longer than diameter of mid ocellus (
<figureCitation id="132F42A5F51F9404FCF347A1DB4B8E1E" box="[975,1055,1512,1536]" captionStart="FIGURES 47 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="18" pageNumber="19">Fig 50</figureCitation>
). Mesofemur in dorsal view with anterior face covered in dense, subdecumbent hairs. Ventral margin of eye edged with darker pigment but without minute punctures on the area. Body color almost uniform dark brown but head darker (
<figureCitation id="132F42A5F51F9404FB5F4479DBE58E56" box="[1123,1201,1584,1608]" captionStart="FIGURES 29 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="18" pageNumber="19">Fig 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFF5441DDE568EAA" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BAB5E20F51F9404FFF5441DDE568EAA" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
<emphasis id="B9608232F51F9404FFF5441DDE1B8E72" bold="true" box="[201,335,1620,1644]" pageId="18" pageNumber="19">Etymology.</emphasis>
This species name is derived from the Latin word
<taxonomicName id="4C1425A3F51F9404FCBB441DDC9C8E72" box="[903,968,1620,1644]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cassis">cassis</taxonomicName>
(helm), and refers to the shape of its genital capsule, which looks like a Corinthian helm. The species epithet is treated as a noun in apposition, and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFF544F6DDB98EC6" box="[201,749,1727,1752]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BAB5E20F51F9404FFF544F6DDB98EC6" blockId="18.[151,1437,1007,2004]" box="[201,749,1727,1752]" pageId="18" pageNumber="19">
<emphasis id="B9608232F51F9404FFF544F6DE368EC6" bold="true" box="[201,354,1727,1752]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="F3031EB0F51F9404FE554489DD618EC6" box="[361,565,1728,1752]" name="Madagascar" pageId="18" pageNumber="19">MADAGASCAR</collectingCountry>
: as in
<figureCitation id="132F42A5F51F9404FD464489DDBD8EC6" box="[634,745,1728,1752]" captionStart="FIGURES 64 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="18" pageNumber="19">Figure 67</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFF544ADDE108FCA" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BAB5E20F51F9404FFF544ADDD498F76" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
<emphasis id="B9608232F51F9404FFF544ADDE698EE2" bold="true" box="[201,317,1764,1788]" pageId="18" pageNumber="19">Remarks.</emphasis>
<taxonomicName id="4C1425A3F51F9404FE7944ADDD748EE2" box="[325,544,1764,1788]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cassis">
<emphasis id="B9608232F51F9404FE7944ADDD748EE2" box="[325,544,1764,1788]" italics="true" pageId="18" pageNumber="19">Adetomyrma cassis</emphasis>
</taxonomicName>
is only known from a single male collected in Réserve Spéciale Ambatovaky. The male of
<taxonomicName id="4C1425A3F51F9404FE144541DEDF8F3E" box="[296,395,1800,1824]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cassis">
<emphasis id="B9608232F51F9404FE144541DE6D8F3E" box="[296,313,1800,1824]" italics="true" pageId="18" pageNumber="19">A</emphasis>
.
<emphasis id="B9608232F51F9404FE744540DEDF8F3E" box="[328,395,1801,1824]" italics="true" pageId="18" pageNumber="19">cassis</emphasis>
</taxonomicName>
is distinguished easily from other
<taxonomicName id="4C1425A3F51F9404FC284541DCF78F3E" box="[788,931,1800,1824]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F51F9404FC284541DCF78F3E" box="[788,931,1800,1824]" italics="true" pageId="18" pageNumber="19">Adetomyrma</emphasis>
</taxonomicName>
males by a distinct and flatted projection on the posterior portion of the paramere (
<figureCitation id="132F42A5F51F9404FD794565DDC78F5A" box="[581,659,1836,1860]" captionStart="FIGURES 15 16" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1435,1278,1913]" captionTargetId="figure@6.[151,1436,1278,1931]" captionTargetPageId="6" captionText="FIGURES 15 16. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 15, A. cassis (CASENT 0163620: holotype); 16, A. venatrix (CASENT 0079480). 15, paramere with a needle-like projection on its posterodorsal portion; 16, paramere without needle-like projection on its posterodorsal portion." httpUri="https://zenodo.org/record/213823/files/figure.png" pageId="18" pageNumber="19">Fig 15</figureCitation>
). This projection is not separated from the paramere by a deep notch (
<figureCitation id="132F42A5F51F9404FFA34519DFBE8F76" box="[159,234,1872,1896]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="18" pageNumber="19">Fig 14</figureCitation>
) as in
<taxonomicName id="4C1425A3F51F9404FE084519DEE08F76" box="[308,436,1872,1896]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F51F9404FE084519DE118F76" box="[308,325,1872,1896]" italics="true" pageId="18" pageNumber="19">A</emphasis>
.
<emphasis id="B9608232F51F9404FE6F4519DEE08F76" box="[339,436,1872,1896]" italics="true" pageId="18" pageNumber="19">bressleri</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F51F9404FEFF4519DD5B8F76" box="[451,527,1872,1896]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="18" pageNumber="19">Fig 13</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF5453DDE108FCA" blockId="18.[151,1437,1007,2004]" pageId="18" pageNumber="19">
This genital character observed in
<taxonomicName id="4C1425A3F51F9404FD6C453DDC7E8F92" box="[592,810,1908,1932]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cassis">
<emphasis id="B9608232F51F9404FD6C453DDC7E8F92" box="[592,810,1908,1932]" italics="true" pageId="18" pageNumber="19">Adetomyrma cassis</emphasis>
</taxonomicName>
is completely unique and sufficient to regard this male as a distinct species, although we have found neither another specimen nor sympatric distribution with other
<taxonomicName id="4C1425A3F51F9404FA6D45D1DFB18FCA" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F51F9404FA6D45D1DFB18FCA" italics="true" pageId="18" pageNumber="19">Adetomyrma</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>