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<document id="56987485D05F5EBB9857D5769AFBF028" ID-DOI="10.11646/zootaxa.3341.1.1" ID-GBIF-Dataset="10b73727-8726-4994-a2ed-3d2a008678c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="213817" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460487960184" checkinUser="plazi" docAuthor="Yoshimura, Masashi &amp; Fisher, Brian L." docDate="2012" docId="03BDEF36F51D9404FFAB42DEDD148B58" docLanguage="en" docName="zt03341p031.pdf" docOrigin="Zootaxa 3341 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3341.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Adetomyrma caputleae, sp. nov." docType="treatment" docVersion="12" lastPageNumber="19" masterDocId="FF84974EF50D9416FF3C4249DF54881E" masterDocTitle="A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="17" updateTime="1698309519930" updateUser="plazi">
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<mods:title id="36D4E33D14818D1B126391139EC82FBD">A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)</mods:title>
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<mods:namePart id="85ABD7DDFFA39878046D11F3C93120AE">Yoshimura, Masashi</mods:namePart>
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<treatment id="03BDEF36F51D9404FFAB42DEDD148B58" ID-DOI="http://doi.org/10.5281/zenodo.5874327" ID-GBIF-Taxon="119518768" ID-Zenodo-Dep="5874327" LSID="urn:lsid:plazi:treatment:03BDEF36F51D9404FFAB42DEDD148B58" httpUri="http://treatment.plazi.org/id/03BDEF36F51D9404FFAB42DEDD148B58" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C30E0DABF51D9406FFAB42DEDD4588AF" box="[151,529,151,177]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BAB5E20F51D9406FFAB42DEDD4588AF" blockId="16.[151,867,151,210]" box="[151,529,151,177]" pageId="16" pageNumber="17">
<heading id="D0E3E94CF51D9406FFAB42DEDD4588AF" bold="true" box="[151,529,151,177]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B9608232F51D9406FFAB42DEDD4588AF" bold="true" box="[151,529,151,177]" pageId="16" pageNumber="17">
<taxonomicName id="4C1425A3F51D9406FFAB42DEDEF988AF" authority="Yoshimura &amp; Fisher 2012" box="[151,429,151,177]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="caputleae" status="sp. nov.">
<emphasis id="B9608232F51D9406FFAB42DEDEF988AF" bold="true" box="[151,429,151,177]" italics="true" pageId="16" pageNumber="17">Adetomyrma caputleae</emphasis>
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<taxonomicNameLabel id="A2533F49F51D9406FE8842DEDD4588AF" box="[436,529,151,177]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BAB5E20F51D9406FFAB42F3DC3788CC" blockId="16.[151,867,151,210]" box="[151,867,186,210]" pageId="16" pageNumber="17">
(
<figureCitation id="132F42A5F51D9406FFA342F3DFB388CC" box="[159,231,186,210]" captionStart="FIGURES 7 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
,
<figureCitation id="132F42A5F51D9406FFCF42F3DE4488CC" box="[243,272,186,210]" captionStart="FIGURES 11 12" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1435,1126,1768]" captionTargetId="figure@5.[151,1436,1126,1771]" captionTargetPageId="5" captionText="FIGURES 11 12. Metatarsus of Adetomyrma workers in posterior view. 11, A. caputleae (CASENT 0227994: paratype); 12, A. venatrix (CASENT 0489808). 11, the fifth tarsomere distinctly wider compared to fourth; 12, the fifth tarsomere almost the same width as fourth." httpUri="https://zenodo.org/record/213821/files/figure.png" pageId="16" pageNumber="17">11</figureCitation>
,
<figureCitation id="132F42A5F51D9406FE2042F3DE6E88CC" box="[284,314,186,210]" captionStart="FIGURES 21 22" captionStartId="8.[151,257,695,716]" captionTargetBox="[163,1427,163,684]" captionTargetId="figure@8.[163,1428,163,701]" captionTargetPageId="8" captionText="FIGURES 21 22. Head of Adetomyrma males in oblique dorsal view. 21, A. caputleae (CASENT 0227999: paratype); 22, A. venatrix (CASENT 0151606). 21, lateral ocellus is distant from the eye; 22, lateral ocellus is close to the eye." httpUri="https://zenodo.org/record/213826/files/figure.png" pageId="16" pageNumber="17">21</figureCitation>
,
<figureCitation id="132F42A5F51D9406FE7A42F3DE3088CC" box="[326,356,186,210]" captionStart="FIGURES 29 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="16" pageNumber="17">31</figureCitation>
,
<figureCitation id="132F42A5F51D9406FE4C42F3DEDA88CC" box="[368,398,186,210]" captionStart="FIGURES 38 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="16" pageNumber="17">40</figureCitation>
,
<figureCitation id="132F42A5F51D9406FEA642F3DEEC88CC" box="[410,440,186,210]" captionStart="FIGURES 47 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="16" pageNumber="17">49</figureCitation>
,
<figureCitation id="132F42A5F51D9406FEF842F3DEB788CC" box="[452,483,186,210]" captionStart="FIGURES 56 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="16" pageNumber="17">58</figureCitation>
,
<figureCitation id="132F42A5F51D9406FED242F3DD5888CC" box="[494,524,186,210]" captionStart="FIGURES 64 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="16" pageNumber="17">66</figureCitation>
,
<figureCitation id="132F42A5F51D9406FD2442F3DD3488CC" box="[536,608,186,210]" captionStart="FIGURES 73 80" captionStartId="25.[151,264,1929,1952]" captionTargetBox="[217,1374,193,1897]" captionTargetId="figure@25.[217,1376,193,1907]" captionTargetPageId="25" captionText="FIGURES 73 80. Workers of Adetomyrma in lateral view and dorsal view. 73, 74, A. bressleri sp. nov. (CASENT 0205995: holotype); 75, 76, A. caputleae sp. nov. (CASENT 0004358); 77, 78, A. goblin sp. nov. (CASENT 0227981: holotype); 79, 80, A. venatrix (MCZTYPE 34786). 73, 75, 77, 79, lateral view; 74, 76, 78, 80, dorsal view." httpUri="https://zenodo.org/record/213834/files/figure.png" pageId="16" pageNumber="17">75, 76</figureCitation>
,
<figureCitation id="132F42A5F51D9406FD5042F3DDDF88CC" box="[620,651,186,210]" captionStart="FIGURES 81 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="16" pageNumber="17">82</figureCitation>
,
<figureCitation id="132F42A5F51D9406FDAA42F3DC5C88CC" box="[662,776,186,210]" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="16" pageNumber="17">85, 87, 89</figureCitation>
,
<figureCitation id="132F42A5F51D9406FC2842F3DC0F88CC" box="[788,859,186,210]" captionStart="FIGURES 91 94" captionStartId="29.[151,264,1742,1765]" captionTargetBox="[151,1435,824,1711]" captionTargetId="figure@29.[151,1436,824,1726]" captionTargetPageId="29" captionText="FIGURES 91 94. Colonies of Adetomyrma. 91, 92, A. caputleae (collection BLF 02531, ex rotten log); 93, 94, A. goblin (collection BLF 14240, ex rotten log, soil). 91, larvae, naked pupae, and workers; 92, pupae, workers, and a large ergatoid queen; 93, elongate larvae feeding between sclerites of a Tenebrionidae larva; 94 two workers grooming a larva. 91, 92, photos by Brian Fisher; 93, 94, photos by Alex Wild." httpUri="https://zenodo.org/record/213837/files/figure.png" pageId="16" pageNumber="17">91, 92</figureCitation>
)
</paragraph>
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<materialsCitation id="3B7C547DF51D9406FFAB434ADD9A897C" ID-GBIF-Occurrence="3446753336" collectedFrom="rotten log" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CASC" collectorName="Fisher &amp; Griswold &amp; al." country="Madagascar" elevation="1300m" latitude="-18.473333" location="Andranomay" longLatPrecision="127" longitude="47.96" municipality="Antananarivo" pageId="16" pageNumber="17" specimenCode="CASENT0227992, BLF02537" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
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<typeStatus id="54AFE082F51D9406FFAB434ADE528905" box="[151,262,259,283]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D1295A9F51D9406FE28434BDE388904" box="[276,364,258,282]" pageId="16" pageNumber="17" type="worker">Worker</specimenCount>
:
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<specimenCode id="DBB2F65BF51D9406FD6B434BDD808904" box="[599,724,258,282]" pageId="16" pageNumber="17">BLF02537</specimenCode>
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<collectingCountry id="F3031EB0F51D9406FDDD434BDCFF8904" box="[737,939,258,282]" name="Madagascar" pageId="16" pageNumber="17">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="6BCFC45AF51D9406FC84434BDB078904" box="[952,1107,258,282]" pageId="16" pageNumber="17">Antananarivo</collectingMunicipality>
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<locationDeviation id="9A194702F51D9406FB5D434BDBAF8904" box="[1121,1275,258,282]" pageId="16" pageNumber="17">
<quantity id="4CECF3C5F51D9406FB5D434BDBC88904" box="[1121,1180,258,282]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="km" value="3.0">3 km</quantity>
41° NE
</locationDeviation>
<location id="8ECB08FBF51D9406FA39434BDACC8904" LSID="urn:lsid:plazi:treatment:03BDEF36F51D9404FFAB42DEDD148B58:8ECB08FBF51D9406FA39434BDACC8904" box="[1285,1432,258,282]" country="Madagascar" latitude="-18.473333" longLatPrecision="127" longitude="47.96" municipality="Antananarivo" name="Andranomay" pageId="16" pageNumber="17">Andranomay</location>
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<locationDeviation id="9A194702F51D9406FFAB436FDE318920" bearing="147.0" box="[151,357,294,318]" distance="11500.0" pageId="16" pageNumber="17">
<quantity id="4CECF3C5F51D9406FFAB436FDFA68920" box="[151,242,294,318]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="17" unit="km" value="11.5">11.5 km</quantity>
147° SSE
</locationDeviation>
<location id="8ECB08FBF51D9406FE50436FDEA78920" LSID="urn:lsid:plazi:treatment:03BDEF36F51D9404FFAB42DEDD148B58:8ECB08FBF51D9406FE50436FDEA78920" box="[364,499,294,318]" country="Madagascar" latitude="-18.473333" longLatPrecision="127" longitude="47.96" municipality="Antananarivo" name="Anjozorobe" pageId="16" pageNumber="17">Anjozorobe</location>
,
<geoCoordinate id="EE2038E7F51D9406FEC2436FDD2B8920" box="[510,639,294,318]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="92" value="-18.473333">18° 28.4´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51D9406FDB7436FDC5A8920" box="[651,782,294,318]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="92" value="47.96">47° 57.6´E</geoCoordinate>
,
<elevation id="0039B913F51D9406FC26436FDCDD8920" box="[794,905,294,318]" pageId="16" pageNumber="17">1300malt</elevation>
., montane rainforest, ex
<collectedFrom id="38081DDDF51D9406FA30436EDAC28920" box="[1292,1430,294,318]" pageId="16" pageNumber="17">rotten log</collectedFrom>
,
<date id="FFAA78E0F51D9406FFAB4303DE61897C" box="[151,309,330,354]" pageId="16" pageNumber="17" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="EFEE8108F51D9406FFAB4303DE61897C" box="[151,309,330,354]" pageId="16" pageNumber="17" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">513.xii.2000</collectingDate>
</date>
,
<collectorName id="26E13BF6F51D9406FE7C4303DEDC897C" box="[320,392,330,354]" pageId="16" pageNumber="17">Fisher</collectorName>
,
<collectorName id="26E13BF6F51D9406FEAF4303DEAF897C" box="[403,507,330,354]" pageId="16" pageNumber="17">Griswold</collectorName>
<emphasis id="B9608232F51D9406FD3E4305DD6F897D" box="[514,571,331,355]" italics="true" pageId="16" pageNumber="17">
et
<collectorName id="26E13BF6F51D9406FD224302DD6F897D" box="[542,571,331,355]" pageId="16" pageNumber="17">al.</collectorName>
</emphasis>
leg. [
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</paragraph>
<paragraph id="8BAB5E20F51D9406FFF54326DD168A08" blockId="16.[151,1437,258,2010]" pageId="16" pageNumber="17">
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<emphasis id="B9608232F51D9406FFF54326DE1C8999" bold="true" box="[201,328,367,391]" pageId="16" pageNumber="17">
<typeStatus id="54AFE082F51D9406FFF54326DE178999" box="[201,323,367,391]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
.
</emphasis>
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:
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with same data as
<typeStatus id="54AFE082F51D9406FEE543B7DD6B8A08" box="[473,575,510,534]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51D9407FFF5406ADCBD8BF2" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BAB5E20F51D9406FFF5406ADB608A40" blockId="16.[151,1437,258,2010]" pageId="16" pageNumber="17">
<emphasis id="B9608232F51D9406FFF5406ADE928A25" bold="true" box="[201,454,546,571]" pageId="16" pageNumber="17">Worker. Description.</emphasis>
Measurements:
<typeStatus id="54AFE082F51D9406FDB9406BDDBF8A24" box="[645,747,546,570]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
. HL 0.72, HW 0.7, SL 0.44, HD 0.5, WL 1.01, PnW 0.47, MnW 0.26, PpW 0.36, PtW 0.39, CI 97.4, SI 63.1, MnI 36.5, PpI 142.7, PtI 106.3.
</paragraph>
<paragraph id="8BAB5E20F51D9406FFF54022DE3F8AD4" blockId="16.[151,1437,258,2010]" pageId="16" pageNumber="17">HL 0.570.72, HW 0.530.7, SL 0.350.45, HD 0.360.5, WL 0.811.05, PnW 0.360.45, MnW 0.20.26, PpW 0.30.37, PtW 0.270.4, CI 91.997.9, SI 60.468.9, MnI 37.238.8, PpI 140.7149.5, PtI 91.3107 (10 individuals measured).</paragraph>
<paragraph id="8BAB5E20F51D9406FFF5409FDB528DFC" blockId="16.[151,1437,258,2010]" pageId="16" pageNumber="17">
Head quadrate in full-face view, almost as long as wide; sides slightly convex, widening near the mandibular insertions, converging posteriorly; posterior margin almost flat (
<figureCitation id="132F42A5F51D9406FC4D40B3DC948B0C" box="[881,960,762,786]" captionStart="FIGURES 81 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="16" pageNumber="17">Fig 82</figureCitation>
). Clypeal principal surface deflected ventrally. Anterior margin of clypeus broadly convex with small, shallow central concavity, and furnished with a row of about 26 small, specialized conical setae. Frontal carinae in full-face view short and low, expanded laterally as small frontal lobes covering more than about 2× the length of antennal insertions. Mandible subfalcate, without distinct basal and masticatory margins (
<figureCitation id="132F42A5F51D9406FD0941C3DD278BBC" box="[565,627,906,930]" captionStart="FIGURES 7 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="16" pageNumber="17">Fig 8</figureCitation>
): inner margin with two apical teeth and three basal denticles; no denticle on base of subapical tooth (second tooth from apex); apical-most denticle slightly larger than other two; sometimes basal-most (third from apical-most one) denticle divided into two adjacent small, low denticles (CASENT004363). Palpal formula 3,3 (three maxillary and three labial). Antennal scape shorter than head length. Pedicel (the second antennal segment) shorter than combined length of next three (third to fifth) antennal segments. Antenna gradually broadened from third segment and not forming a distinct club. Dorsal outline of mesosoma in lateral view flattened, mesonotum not raised from pronotal and propodeal dorsum. Pronotum in dorsal view longer than broad, with convex sides. Mesonotum in dorsal view short, 2× as wide as long. Metapleuron fully fused with propodeum, division of the two plates not distinguishable in lateral view. Dorsal face of propodeum narrower than pronotum, about 1.5× longer than wide, with subparallel sides converging slightly toward mesonotum (
<figureCitation id="132F42A5F51D9406FA034687DAC48CF8" box="[1343,1424,1230,1254]" captionStart="FIGURES 73 80" captionStartId="25.[151,264,1929,1952]" captionTargetBox="[217,1374,193,1897]" captionTargetId="figure@25.[217,1376,193,1907]" captionTargetPageId="25" captionText="FIGURES 73 80. Workers of Adetomyrma in lateral view and dorsal view. 73, 74, A. bressleri sp. nov. (CASENT 0205995: holotype); 75, 76, A. caputleae sp. nov. (CASENT 0004358); 77, 78, A. goblin sp. nov. (CASENT 0227981: holotype); 79, 80, A. venatrix (MCZTYPE 34786). 73, 75, 77, 79, lateral view; 74, 76, 78, 80, dorsal view." httpUri="https://zenodo.org/record/213834/files/figure.png" pageId="16" pageNumber="17">Fig 76</figureCitation>
). Propodeum in lateral view, its dorsal margin 2× or less the length of its declivity, and rounding gently into the latter. Propodeal spiracle medium sized, distant from propodeal dorsal margin in lateral view, usually not visible in dorsal view. Subpetiolar process distinctly developed, shaped like an irregular axe blade. Shallow and dense punctures covering dorsal head, mesosoma, abdomen, and ventral head. Clypeus with two groups of hairs; mid clypeal hairs directed dorsally, and anterior clypeal hairs deflected anteriorly. Mid clypeal hairs consisting of one long hair and one to several shorter hairs around the long one. With head in full-face view, anterior clypeal hairs consisting of one pair of long hairs and two (sometimes three) shorter hairs laterally (
<figureCitation id="132F42A5F51D9406FC864783DCAC8DFC" box="[954,1016,1482,1506]" captionStart="FIGURES 7 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="16" pageNumber="17">Fig 8</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F51D9406FFF547A7DCCA8E50" blockId="16.[151,1437,258,2010]" pageId="16" pageNumber="17">
<emphasis id="B9608232F51D9406FFF547A7DEEA8E19" bold="true" box="[201,446,1518,1543]" pageId="16" pageNumber="17">Queen. Description.</emphasis>
Measurements: HL 0.921.07, HW 1.051.18, SL 0.530.56, EL 0.070.07, WL 1.41.48, MnW 0.550.68, PpW 0.740.78, PtW 0.810.93, CI 108.1114.6, SI 47.151, EI 6.97.2, MnI 51.157.7, PpI 114.2134.4, PtI 110126.1 (3 individuals measured).
</paragraph>
<paragraph id="8BAB5E20F51D9407FFF54413DBC4897A" blockId="16.[151,1437,258,2010]" lastBlockId="17.[151,1436,151,2012]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Wingless. Head quadrate in full-face view (
<figureCitation id="132F42A5F51D9406FD804413DC5F8E6C" box="[700,779,1626,1650]" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="16" pageNumber="17">Fig 89</figureCitation>
), almost as long as wide; sides slightly convex; posterior margin slightly concave. Eye small, consisting of about seven ommatidia. Mid ocellus vestigial in a small concavity. Lateral ocelli absent. Clypeal principal surface not strongly deflected. Anterior margin of clypeus with central concavity, and furnished with a row of about 17 small, specialized, conical setae. Frontal lobes separated by midposterior portion of clypeus. Mandible subfalcate, without distinct basal and masticatory margins, inner margin with two apical teeth and two low and dull basal denticles, no denticle on base of subapical tooth (second tooth from apex). Antennal scape 0.5× as long as combined length of pedicel and flagellum. First flagellomere as long as wide. Distal antennal segments not forming distinct club. In lateral view, mesosomal dorsum somewhat flattened (
<figureCitation id="132F42A5F51D9406FFA24533DFB98F8C" box="[158,237,1914,1938]" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="16" pageNumber="17">Fig 85</figureCitation>
); pronotum occupying entire anterior and about 0.33× of dorsal margins; mesonotum reduced in size and only occupying about 0.33× of mesosomal dorsal margin; metanotum not visible; propodeum occupying about 0.33× of remaining dorsal and posterior margins. With mesosoma in dorsal view, division between mesoscutum and mesoscutellum often unclear (
<figureCitation id="132F42A5F51C9407FD2A42DEDD3788B1" box="[534,611,151,175]" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="17" pageNumber="18">Fig 87</figureCitation>
); axillae not distinct; metanotum reduced into narrow tip; and propodeum distinctly widened posteriorly. Propodeal spiracle in lateral view located close to propodeal declivity in lateral view, distance from declivity margin as long as diameter of spiracle. Petiole stouter than that in conspecific worker (
<figureCitation id="132F42A5F51C9407FFA3434DDFB98902" box="[159,237,260,284]" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="17" pageNumber="18">Fig 87</figureCitation>
), distinctly wider than long, width approximately same as that of propodeum in dorsal view, anterodorsal portion more protruding anteriorly in lateral view. When metasoma seen in dorsal view, little difference in width present between posterior portion of petiole and anterior portion of abdominal segment III.
</paragraph>
<paragraph id="8BAB5E20F51C9407FFF54326DC8989D1" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
Clypeus with long hairs; division of mid clypeal hairs and anterior clypeal hairs unclear. With head in full-face view, anterior portion of clypeus with one pair of long hairs and two (sometimes three) shorter hairs laterally (
<figureCitation id="132F42A5F51C9407FA4943DDDFE389D1" captionStart="FIGURES 85 90" captionStartId="28.[151,264,1944,1967]" captionTargetBox="[151,1435,371,1911]" captionTargetId="figure@28.[151,1436,357,1919]" captionTargetPageId="28" captionText="FIGURES 85 90. Queens of Adetomyrma species. 85, 87, 89, A. caputleae sp. nov. (CASENT 0004350); 86, 88, 90, A. goblin sp. nov. (CASENT 0007401); 85, 86, lateral view; 87, 88, dorsal view; 89, 90, head in full-face view." httpUri="https://zenodo.org/record/213836/files/figure.png" pageId="17" pageNumber="18">Fig 89</figureCitation>
). Body surface with denser appressed hairs than in conspecific workers.
</paragraph>
<paragraph id="8BAB5E20F51C9407FFF54395DA548A09" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="B9608232F51C9407FFF54395DEF289EA" bold="true" box="[201,422,475,500]" pageId="17" pageNumber="18">Male. Description.</emphasis>
Measurements: HL 0.640.73, HW 0.911.1, SL 0.20.26, EL 0.390.44, WL 1.842.03, MnW 1.131.27, CI 143151.2, SI 2124.3, EI 60.467, MnI 114.8124.1 (5 individuals measured).
</paragraph>
<paragraph id="8BAB5E20F51C9407FFF5406CDDE98AD2" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
Eye well developed and prominent, posterior margin not exceeding posterior margin of mid ocellus in full-face view. Distance between lateral ocellus and eye longer than 1.5× diameter of lateral ocellus (
<figureCitation id="132F42A5F51C9407FBAA400EDBB78A41" box="[1174,1251,583,607]" captionStart="FIGURES 21 22" captionStartId="8.[151,257,695,716]" captionTargetBox="[163,1427,163,684]" captionTargetId="figure@8.[163,1428,163,701]" captionTargetPageId="8" captionText="FIGURES 21 22. Head of Adetomyrma males in oblique dorsal view. 21, A. caputleae (CASENT 0227999: paratype); 22, A. venatrix (CASENT 0151606). 21, lateral ocellus is distant from the eye; 22, lateral ocellus is close to the eye." httpUri="https://zenodo.org/record/213826/files/figure.png" pageId="17" pageNumber="18">Fig 21</figureCitation>
). Palpal formula 3,3 (three maxillary and three labial). Notaulus absent on mesoscutum. Parapsidal line clearly impressed (
<figureCitation id="132F42A5F51C9407FA7D4025DAC48A9A" box="[1345,1424,620,644]" captionStart="FIGURES 38 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="17" pageNumber="18">Fig 40</figureCitation>
). Anterior margin of petiole as long as or shorter than dorsal margin in lateral view. Subpetiolar process well developed, usually with more than five hairs ventrally.
</paragraph>
<paragraph id="8BAB5E20F51C9407FFF5409EDAC18B9E" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
Left and right parameres not overlapping or narrowly overlapping on dorsal small part of basimere. No distinct projection or lobe present on posterodorsal portion of paramere. Basal ring not reduced, covering whole anterior margin of paramere in lateral view. Basal projection on cuspis clear but not extraordinarily well developed. Aedeagus in lateral view as in
<figureCitation id="132F42A5F51C9407FE9A410DDD418B42" box="[422,533,836,860]" captionStart="FIGURES 56 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="17" pageNumber="18">Figure 58</figureCitation>
; narrowing from base of ventral projection; apical margin relatively sharp with a small, broadly triangular projection on posteroventral portion; posteroventral margin of ventral projection convex.
</paragraph>
<paragraph id="8BAB5E20F51C9407FFF541C5DCBD8BF2" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
Hair on compound eyes short, 0.33× to 0.5× diameter of mid ocellus. Mesofemur in dorsal view, its anterior face with dense decumbent hairs, and sometimes one or two longer suberect hairs. Ventral margin of eye not edged with darker pigment or punctures. Body color uniform dark brown (
<figureCitation id="132F42A5F51C9407FCB2419DDC8F8BF2" box="[910,987,980,1004]" captionStart="FIGURES 29 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="17" pageNumber="18">Fig 31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51C9407FFF541B1DA098C2A" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BAB5E20F51C9407FFF541B1DA098C2A" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="B9608232F51C9407FFF541B1DE1B8C0E" bold="true" box="[201,335,1016,1040]" pageId="17" pageNumber="18">Etymology.</emphasis>
This species name is derived from the Latin words caput (head) and lea (lioness), referring to the stout, thick head of the new species. The species epithet is treated as a noun in apposition, and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51C9407FFF54676DDB98C49" box="[201,749,1087,1112]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BAB5E20F51C9407FFF54676DDB98C49" blockId="17.[151,1436,151,2012]" box="[201,749,1087,1112]" pageId="17" pageNumber="18">
<emphasis id="B9608232F51C9407FFF54676DE368C46" bold="true" box="[201,354,1087,1112]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="F3031EB0F51C9407FE554676DD618C49" box="[361,565,1087,1111]" name="Madagascar" pageId="17" pageNumber="18">MADAGASCAR</collectingCountry>
: as in
<figureCitation id="132F42A5F51C9407FD464676DDBD8C49" box="[634,745,1087,1111]" captionStart="FIGURES 64 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="17" pageNumber="18">Figure 66</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51C9407FFF5462ADC788EFE" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BAB5E20F51C9407FFF5462ADC788EFE" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="B9608232F51C9407FFF5462ADD7A8C62" bold="true" box="[201,558,1123,1148]" pageId="17" pageNumber="18">Additional material examined</emphasis>
: in addition to the
<typeStatus id="54AFE082F51C9407FC3F462CDC678C63" box="[771,819,1125,1149]" pageId="17" pageNumber="18">type</typeStatus>
material, specimens from the following localities were examined in this study:
<collectingCountry id="F3031EB0F51C9407FE9D46C1DD388CBE" box="[417,620,1160,1184]" name="Madagascar" pageId="17" pageNumber="18">MADAGASCAR</collectingCountry>
. Analamanga Region, district of Ankazobe, Ambohitantely,
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NE of Ankazobe,
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,
<geoCoordinate id="EE2038E7F51C9407FE9346E5DD6B8CDA" box="[431,575,1196,1220]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.2815">47° 16.89´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD7646E5DDC58CDA" box="[586,657,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="17" pageNumber="18" unit="m" value="701.0">701 m</quantity>
alt., sclerophyll forest;
<quantity id="4CECF3C5F51C9407FCA546E5DC858CDA" box="[921,977,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
41° NE Andranomay,
<quantity id="4CECF3C5F51C9407FBF346E5DA7D8CDA" box="[1231,1321,1196,1220]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="18" unit="km" value="11.5">11.5 km</quantity>
147° SSE Anjozorobe,
<geoCoordinate id="EE2038E7F51C9407FE164699DEFA8CF6" box="[298,430,1232,1256]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="92" value="-18.473333">18° 28.4´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FE804699DD158CF6" box="[444,577,1232,1256]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="92" value="47.96">47° 57.6´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD724699DDF18CF6" box="[590,677,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="18" unit="m" value="1300.0">1300 m</quantity>
alt., montane rainforest; Ankokoy Forest,
<quantity id="4CECF3C5F51C9407FBB64699DB908CF6" box="[1162,1220,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
E Ibity, malaise in Uapaca forest,
<geoCoordinate id="EE2038E7F51C9407FE7D46BDDE968D12" box="[321,450,1268,1292]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-20.0675">20° 4.05´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FEF346BDDD358D12" box="[463,609,1268,1292]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="46.9995">46° 59.97´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD5146BDDD968D12" box="[621,706,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="m" value="1700.0">1700 m</quantity>
alt., Uapaca forest; Amoron'i Mania Region, district of Ambositra, Italaviana Uapaca forest,
<quantity id="4CECF3C5F51C9407FED84751DD668D2E" box="[484,562,1304,1328]" metricMagnitude="5" metricUnit="m" metricValue="1.35" pageId="17" pageNumber="18" unit="km" value="135.0">135km</quantity>
SE of Antsirabe,
<geoCoordinate id="EE2038E7F51C9407FDC14751DC2B8D2E" box="[765,895,1304,1328]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="92" value="-20.173334">20° 10.4´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FCB14751DB468D2E" box="[909,1042,1304,1328]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.086">47° 5.16´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FB224751DB208D2E" box="[1054,1140,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="17" pageNumber="18" unit="m" value="1359.0">1359 m</quantity>
alt., Uapaca forest; Forêt d'Atsirakambiaty,
<quantity id="4CECF3C5F51C9407FE584775DEE78D4A" box="[356,435,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="17" pageNumber="18" unit="km" value="7.6">7.6 km</quantity>
285° WNW Itremo,
<geoCoordinate id="EE2038E7F51C9407FD994775DC7C8D4A" box="[677,808,1340,1364]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="92" value="-20.593334">20° 35.6´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FC0A4775DCE88D4A" box="[822,956,1340,1364]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="92" value="46.56333">46° 33.8´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FCF54775DB4B8D4A" box="[969,1055,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="18" unit="m" value="1550.0">1550 m</quantity>
alt., montane rainforest;
<quantity id="4CECF3C5F51C9407FA034775DAC88D4A" box="[1343,1436,1340,1364]" metricMagnitude="4" metricUnit="m" metricValue="2.7399999999999998" pageId="17" pageNumber="18" unit="km" value="27.4">27.4 km</quantity>
SSW Ambositra,
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,
<geoCoordinate id="EE2038E7F51C9407FED64729DD398D66" box="[490,621,1376,1400]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="92" value="47.18667">47° 11.2´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD454729DD9A8D66" box="[633,718,1376,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="18" unit="m" value="1600.0">1600 m</quantity>
alt., disturbed montane rainforest; Parc naturel communautaire,
<quantity id="4CECF3C5F51C9407FFAB47CDDFA78D82" box="[151,243,1412,1436]" metricMagnitude="4" metricUnit="m" metricValue="2.68" pageId="17" pageNumber="18" unit="km" value="26.8">26.8 km</quantity>
SW Ambositra,
<geoCoordinate id="EE2038E7F51C9407FE8C47CDDD658D82" box="[432,561,1412,1436]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="92" value="-20.775">20° 46.5´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FD0147CDDD998D82" box="[573,717,1412,1436]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.183666">47° 11.02´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FDE547CDDC7B8D82" box="[729,815,1412,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.755" pageId="17" pageNumber="18" unit="m" value="1755.0">1755 m</quantity>
alt., disturbed montane rainforest; Miandritsara Forest,
<quantity id="4CECF3C5F51C9407FFAB47E1DF8B8DDE" box="[151,223,1448,1472]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="km" value="40.0">40 km</quantity>
S of Ambositra,
<geoCoordinate id="EE2038E7F51C9407FE9F47E1DD608DDE" box="[419,564,1448,1472]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-20.792667">20° 47.56´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FD7D47E1DD808DDE" box="[577,724,1448,1472]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.175667">47° 10.54´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FDDD47E1DC7D8DDE" box="[737,809,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="8.22" pageId="17" pageNumber="18" unit="m" value="822.0">822 m</quantity>
alt., rainforest; Vohiparara broken bridge, Fianarantsoa Prov.,
<geoCoordinate id="EE2038E7F51C9407FFDD4785DE218DFA" box="[225,373,1484,1508]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.226166">21° 13.57´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FEB84785DD4D8DFA" box="[388,537,1484,1508]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.369835">47° 22.19´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD144785DD298DFA" box="[552,637,1484,1508]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="17" pageNumber="18" unit="m" value="1110.0">1110 m</quantity>
alt., high altitude rainforest; radio tower, Ranomafana National Park, Fianarantsoa Prov.,
<geoCoordinate id="EE2038E7F51C9407FE4947B9DD508E16" box="[373,516,1520,1544]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.250834">21° 15.05´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FD2D47B9DDF78E16" box="[529,675,1520,1544]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.407166">47° 24.43´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FD9247B9DC578E16" box="[686,771,1520,1544]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="17" pageNumber="18" unit="m" value="1130.0">1130 m</quantity>
alt., forest edge, mixed tropical forest, open area; Fitovavy Fitovinany Region, district of Ifanadiana,
<quantity id="4CECF3C5F51C9407FD52445DDDE18E32" box="[622,693,1556,1580]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="km" value="12.0">12 km</quantity>
W of Ranomafana,
<geoCoordinate id="EE2038E7F51C9407FCAB445DDB728E32" box="[919,1062,1556,1580]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.250834">21° 15.05´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FB0F445DDB908E32" box="[1075,1220,1556,1580]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.407166">47° 24.43´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FBEC445DDA708E32" box="[1232,1316,1556,1580]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="17" pageNumber="18" unit="m" value="1127.0">1127 m</quantity>
alt., forest edge, open area; JIRAMA water works near river, Ranomafana National Park, Fianarantsoa Prov.,
<geoCoordinate id="EE2038E7F51C9407FA384471DAC28E4E" box="[1284,1430,1592,1616]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.2485">21° 14.91´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FFAB4415DE788E6A" box="[151,300,1628,1652]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.452168">47° 27.13´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FE054415DED68E6A" box="[313,386,1628,1652]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="17" pageNumber="18" unit="m" value="690.0">690 m</quantity>
alt., open area near stream; Belle Vue trail, Ranomafana National Park, Fianarantsoa Prov.,
<geoCoordinate id="EE2038E7F51C9407FFAB44C9DE7C8E86" box="[151,296,1664,1688]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.2665">21° 15.99´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FE0B44C9DE9E8E86" box="[311,458,1664,1688]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.420166">47° 25.21´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FEE444C9DD7A8E86" box="[472,558,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="17" pageNumber="18" unit="m" value="1020.0">1020 m</quantity>
alt., mixed tropical forest; Foret d'Ambalagoavy Nord,
<collectingCountry id="F3031EB0F51C9407FB9744C9DBA98E86" box="[1195,1277,1664,1688]" name="Republic of the Congo" pageId="17" pageNumber="18">Ikongo</collectingCountry>
, Ambatombe,
<geoCoordinate id="EE2038E7F51C9407FFAB44EDDE718EA2" box="[151,293,1700,1724]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-21.8275">21° 49.65´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FE0E44EDDE978EA2" box="[306,451,1700,1724]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="47.338833">47° 20.33´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FEF244EDDD418EA2" box="[462,533,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="17" pageNumber="18" unit="m" value="625.0">625 m</quantity>
alt.; Parc National d'Andringitra, Plateau d'Andohariana,
<quantity id="4CECF3C5F51C9407FBA744EDDBA38EA2" box="[1179,1271,1700,1724]" metricMagnitude="4" metricUnit="m" metricValue="3.59" pageId="17" pageNumber="18" unit="km" value="35.9">35.9 km</quantity>
205° Ambalavao,
<geoCoordinate id="EE2038E7F51C9407FFF14481DE1A8EFE" box="[205,334,1736,1760]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="-22.152334">22° 9.14´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51C9407FE664481DEB88EFE" box="[346,492,1736,1760]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="46.899166">46° 53.95´E</geoCoordinate>
,
<quantity id="4CECF3C5F51C9407FECB4481DD188EFE" box="[503,588,1736,1760]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="2000.0">2000 m</quantity>
alt., ericoid thicket.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51C9404FFF544A5DBDE8A38" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BAB5E20F51C9407FFF544A5DEE98FC2" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="B9608232F51C9407FFF544A5DE698F1A" bold="true" box="[201,317,1772,1796]" pageId="17" pageNumber="18">Remarks.</emphasis>
The worker of
<taxonomicName id="4C1425A3F51C9407FED744A5DDBA8F1A" box="[491,750,1772,1796]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="caputleae">
<emphasis id="B9608232F51C9407FED744A5DDBA8F1A" box="[491,750,1772,1796]" italics="true" pageId="17" pageNumber="18">Adetomyrma caputleae</emphasis>
</taxonomicName>
is easily distinguished from that of
<taxonomicName id="4C1425A3F51C9407FB4344A5DBAB8F1A" box="[1151,1279,1772,1796]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F51C9407FB4344A5DBAB8F1A" box="[1151,1279,1772,1796]" italics="true" pageId="17" pageNumber="18">A. bressleri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1425A3F51C9407FA0944A5DAC88F1A" box="[1333,1436,1772,1796]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="goblin">
<emphasis id="B9608232F51C9407FA0944A5DAC88F1A" box="[1333,1436,1772,1796]" italics="true" pageId="17" pageNumber="18">A. goblin</emphasis>
</taxonomicName>
by the lack of a small denticle on the basal portion of the subapical tooth and by having more than one hair lateral to the longest anterior clypeal hairs (
<figureCitation id="132F42A5F51C9407FD09457DDD278F52" box="[565,627,1844,1868]" captionStart="FIGURES 7 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="17" pageNumber="18">Fig 8</figureCitation>
), and from
<taxonomicName id="4C1425A3F51C9407FDC5457DDC218F52" box="[761,885,1844,1868]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F51C9407FDC5457DDC218F52" box="[761,885,1844,1868]" italics="true" pageId="17" pageNumber="18">A. venatrix</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F51C9407FCB8457DDC848F52" box="[900,976,1844,1868]" captionStart="FIGURES 81 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="17" pageNumber="18">Fig 84</figureCitation>
) by the flattened posterior margin of the head in full-face view (
<figureCitation id="132F42A5F51C9407FEA04511DEBC8F6E" box="[412,488,1880,1904]" captionStart="FIGURES 81 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="17" pageNumber="18">Fig 82</figureCitation>
) and the swollen ventral margin of the head in lateral view. In the male, this species is distinguished from other
<taxonomicName id="4C1425A3F51C9407FEE84535DD378F8A" box="[468,611,1916,1940]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F51C9407FEE84535DD378F8A" box="[468,611,1916,1940]" italics="true" pageId="17" pageNumber="18">Adetomyrma</emphasis>
</taxonomicName>
species by the combination of a smaller lateral ocellus distant from the eye, the relatively longer dorsal margin than anterior margin of the petiole, the paramere without projection or lobe, and the brown body color.
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF542DEDECA89D1" blockId="18.[151,1436,151,463]" pageId="18" pageNumber="19">
Geographical variation in male characters can be observed in body size, wing color, head shape, and the development of the subpetiolar process and its hairs. The smallest specimens are often similar to males of
<taxonomicName id="4C1425A3F51F9404FAB842F5DFA788E6" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F51F9404FAB842F5DAC188CA" box="[1412,1429,188,212]" italics="true" pageId="18" pageNumber="19">A</emphasis>
.
<emphasis id="B9608232F51F9404FFAB42A8DFA788E6" box="[151,243,225,248]" italics="true" pageId="18" pageNumber="19">venatrix</emphasis>
</taxonomicName>
; however, the diagnostic characters above still separate
<taxonomicName id="4C1425A3F51F9404FC4042A9DB5F88E6" box="[892,1035,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="caputleae">
<emphasis id="B9608232F51F9404FC4042A9DCD988E6" box="[892,909,224,248]" italics="true" pageId="18" pageNumber="19">A</emphasis>
.
<emphasis id="B9608232F51F9404FCA242A9DB5F88E6" box="[926,1035,224,248]" italics="true" pageId="18" pageNumber="19">caputleae</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1425A3F51F9404FB6842A9DB8588E6" box="[1108,1233,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F51F9404FB6842A9DB3188E6" box="[1108,1125,224,248]" italics="true" pageId="18" pageNumber="19">A</emphasis>
.
<emphasis id="B9608232F51F9404FB4A42A8DB8588E6" box="[1142,1233,225,248]" italics="true" pageId="18" pageNumber="19">venatrix</emphasis>
</taxonomicName>
. Specimens from Forêt d'Atsirakambiaty (
<geoCoordinate id="EE2038E7F51F9404FE92434DDD628902" box="[430,566,260,284]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="92" value="-20.593334">20° 35.6´S</geoCoordinate>
,
<geoCoordinate id="EE2038E7F51F9404FD79434DDD9A8902" box="[581,718,260,284]" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="92" value="46.56333">46° 33.8´E</geoCoordinate>
,
<quantity id="4CECF3C5F51F9404FDE1434DDC618902" box="[733,821,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="19" unit="m" value="1550.0">1550 m</quantity>
alt.) show larger morphological differences from the others in wing color and head shape. The Forêt d'Atsirakambiaty specimens were separately listed as
<emphasis id="B9608232F51F9404FA214361DA7A895E" box="[1309,1326,296,320]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. mg05 in
<bibRefCitation id="EF8523D1F51F9404FFAB4305DE82897A" author="Yoshimura &amp; Fisher" box="[151,470,332,356]" firstAuthor="Yoshimura" pageId="18" pageNumber="19" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">Yoshimura &amp; Fisher (2012)</bibRefCitation>
. However, our detailed examination revealed that the apparent differences appear in only one parapatric population. Moreover, the morphological differences are not very consistent, which are observed even within members of a single colony. Therefore these morphological differences should be regarded as intraspecific variations.
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54047DBDE8A38" blockId="18.[201,1162,526,550]" box="[201,1162,526,550]" pageId="18" pageNumber="19">This new species corresponds to the following species codes used in previous studies:</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF51F9404FFF5401FDD148B58" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BAB5E20F51F9404FFF5401FDE9C8A70" blockId="18.[201,685,598,838]" box="[201,456,598,623]" pageId="18" pageNumber="19">
sp.2a:
<bibRefCitation id="EF8523D1F51F9404FE2E401FDE9C8A70" author="Saux" box="[274,456,598,623]" etAl="et al." firstAuthor="Saux" pageId="18" pageNumber="19" pagination="457" refId="ref20495" refString="Saux, C., Fisher, B. L. &amp; Spicer, G. S. (2004) Dracula ant phylogeny as inferred by nuclear 28 S rDNA sequences and implications for ant systematics (Hymenoptera: Formicidae: Amblyoponinae). Molecular Phylogenetics and Evolution, 33, 457 - 468." type="journal article" year="2004">
Saux
<emphasis id="B9608232F51F9404FE6D4011DEDD8A70" box="[337,393,598,623]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2004
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54033DDF98A8C" blockId="18.[201,685,598,838]" box="[201,685,634,659]" pageId="18" pageNumber="19">
sp.2b:
<bibRefCitation id="EF8523D1F51F9404FE284033DE998A8C" author="Saux" box="[276,461,634,659]" etAl="et al." firstAuthor="Saux" pageId="18" pageNumber="19" pagination="457" refId="ref20495" refString="Saux, C., Fisher, B. L. &amp; Spicer, G. S. (2004) Dracula ant phylogeny as inferred by nuclear 28 S rDNA sequences and implications for ant systematics (Hymenoptera: Formicidae: Amblyoponinae). Molecular Phylogenetics and Evolution, 33, 457 - 468." type="journal article" year="2004">
Saux
<emphasis id="B9608232F51F9404FE6E4035DEDE8A8D" box="[338,394,635,659]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="EF8523D1F51F9404FEE44032DDF98A8C" author="Moreau" box="[472,685,634,659]" etAl="et al." firstAuthor="Moreau" pageId="18" pageNumber="19" pagination="101" refId="ref20389" refString="Moreau, C. S., Bell, C. D., Vila, R., Archibald, S. B. &amp; Pierce, N. E. (2006) Phylogeny of the ants: Diversification in the age of angiosperms. Science, 312, 101 - 104." type="journal article" year="2006">
Moreau
<emphasis id="B9608232F51F9404FD0A4035DD3A8A8D" box="[566,622,635,659]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2006
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF540D7DEA38AA8" blockId="18.[201,685,598,838]" box="[201,503,670,695]" pageId="18" pageNumber="19">
MAD02:
<bibRefCitation id="EF8523D1F51F9404FE0940D7DEA38AA8" author="Brady" box="[309,503,670,695]" etAl="et al." firstAuthor="Brady" pageId="18" pageNumber="19" pagination="18172 - 18177" refId="ref20148" refString="Brady, S. G., Schultz, T. R. &amp; Fisher, B. L. (2006) Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences of the United States of America, 103, 18172 - 18177." type="journal article" year="2006">
Brady
<emphasis id="B9608232F51F9404FEBC40E9DEED8AA8" box="[384,441,670,695]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2006
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF5408BDD738AC4" blockId="18.[201,685,598,838]" box="[201,551,706,731]" pageId="18" pageNumber="19">
sp.Ma-02:
<bibRefCitation id="EF8523D1F51F9404FE7E408BDD738AC4" author="Ouellette" box="[322,551,706,731]" etAl="et al." firstAuthor="Ouellette" pageId="18" pageNumber="19" pagination="359" refId="ref20444" refString="Ouellette, G. D., Fisher, B. L. &amp; Girman, D. J. (2006) Molecular systematics of basal subfamilies of ants using 28 S rRNA (Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution, 40, 359 - 369." type="journal article" year="2006">
Ouellette
<emphasis id="B9608232F51F9404FE8C408DDEBC8AC5" box="[432,488,707,731]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2006
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF540AFDD738AE0" blockId="18.[201,685,598,838]" box="[201,551,742,767]" pageId="18" pageNumber="19">
sp.Ma-04:
<bibRefCitation id="EF8523D1F51F9404FE7E40AFDD738AE0" author="Ouellette" box="[322,551,742,767]" etAl="et al." firstAuthor="Ouellette" pageId="18" pageNumber="19" pagination="359" refId="ref20444" refString="Ouellette, G. D., Fisher, B. L. &amp; Girman, D. J. (2006) Molecular systematics of basal subfamilies of ants using 28 S rRNA (Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution, 40, 359 - 369." type="journal article" year="2006">
Ouellette
<emphasis id="B9608232F51F9404FE8C40A1DEBC8AE0" box="[432,488,742,767]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
<quantity id="4CECF3C5F51F9404FED340AFDD738AE0" box="[495,551,742,766]" metricMagnitude="-3" metricUnit="kg" metricValue="2.006" pageId="18" pageNumber="19" unit="mg" value="2006.0">2006</quantity>
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54143DD148B3C" blockId="18.[201,685,598,838]" box="[201,576,778,802]" pageId="18" pageNumber="19">
mg02:
<bibRefCitation id="EF8523D1F51F9404FE2B4143DD148B3C" author="Yoshimura &amp; Fisher" box="[279,576,778,802]" firstAuthor="Yoshimura" pageId="18" pageNumber="19" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">
Yoshimura &amp; Fisher,
<quantity id="4CECF3C5F51F9404FD344143DD148B3C" box="[520,576,778,802]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0119999999999996" pageId="18" pageNumber="19" unit="mg" value="2012.0">2012</quantity>
</bibRefCitation>
</paragraph>
<paragraph id="8BAB5E20F51F9404FFF54167DD148B58" blockId="18.[201,685,598,838]" box="[201,576,814,838]" pageId="18" pageNumber="19">
mg05:
<bibRefCitation id="EF8523D1F51F9404FE2B4167DD148B58" author="Yoshimura &amp; Fisher" box="[279,576,814,838]" firstAuthor="Yoshimura" pageId="18" pageNumber="19" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">Yoshimura &amp; Fisher, 2012</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>