treatments-xml/data/03/BD/EF/03BDEF36F5069418FFAB4025DD158A41.xml

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<mods:title id="0FBCCF842301CDC5453B565BD2D0195E">A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)</mods:title>
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<taxonomicName id="4C1425A3F506941DFFAB4025DEF48A98" authority="Yoshimura &amp; Fisher 2012" box="[151,416,620,646]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bressleri" status="sp. nov.">
<emphasis id="B9608232F506941DFFAB4025DEF48A98" bold="true" box="[151,416,620,646]" italics="true" pageId="11" pageNumber="12">Adetomyrma bressleri</emphasis>
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(
<figureCitation id="132F42A5F506941DFFA340C7DFB38AB8" box="[159,231,654,678]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="11" pageNumber="12">Figs 7</figureCitation>
, 
<figureCitation id="132F42A5F506941DFFCF40C7DE578AB8" box="[243,259,654,678]" captionStart="FIGURES 9 – 10" captionStartId="5.[151,264,841,864]" captionTargetBox="[151,1435,342,814]" captionTargetId="figure@5.[151,1436,342,816]" captionTargetPageId="5" captionText="FIGURES 9 – 10. Propodeum of Adetomyrma workers in dorsal view. 9, A. bressleri (CASENT 0205995: holotype); 10, A. goblin (CASENT 0227981: holotype). 9, the propodeal spiracles are visible in dorsal view; 10, the propodeal spiracles are not visible in dorsal view." httpUri="https://zenodo.org/record/213820/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
, 
<figureCitation id="132F42A5F506941DFE3340C7DE798AB8" box="[271,301,654,678]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="11" pageNumber="12">13</figureCitation>
, 
<figureCitation id="132F42A5F506941DFE0540C7DE0C8AB8" box="[313,344,654,678]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="11" pageNumber="12">30</figureCitation>
, 
<figureCitation id="132F42A5F506941DFE5F40C7DED68AB8" box="[355,386,654,678]" captionStart="FIGURES 38 – 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 – 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="11" pageNumber="12">39</figureCitation>
, 
<figureCitation id="132F42A5F506941DFEB140C7DEFF8AB8" box="[397,427,654,678]" captionStart="FIGURES 47 – 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 – 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="11" pageNumber="12">48</figureCitation>
, 
<figureCitation id="132F42A5F506941DFE8B40C7DE818AB8" box="[439,469,654,678]" captionStart="FIGURES 56 – 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 – 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="11" pageNumber="12">57</figureCitation>
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<figureCitation id="132F42A5F506941DFEDD40C7DEAB8AB8" box="[481,511,654,678]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">65</figureCitation>
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<figureCitation id="132F42A5F506941DFD3740C7DD078AB8" box="[523,595,654,678]" captionStart="FIGURES 73 – 80" captionStartId="25.[151,264,1929,1952]" captionTargetBox="[217,1374,193,1897]" captionTargetId="figure@25.[217,1376,193,1907]" captionTargetPageId="25" captionText="FIGURES 73 – 80. Workers of Adetomyrma in lateral view and dorsal view. 73, 74, A. bressleri sp. nov. (CASENT 0205995: holotype); 75, 76, A. caputleae sp. nov. (CASENT 0004358); 77, 78, A. goblin sp. nov. (CASENT 0227981: holotype); 79, 80, A. venatrix (MCZTYPE 34786). 73, 75, 77, 79, lateral view; 74, 76, 78, 80, dorsal view." httpUri="https://zenodo.org/record/213834/files/figure.png" pageId="11" pageNumber="12">73, 74</figureCitation>
, 
<figureCitation id="132F42A5F506941DFD6340C7DD2F8AB8" box="[607,635,654,678]" captionStart="FIGURES 81 – 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 – 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="11" pageNumber="12">81</figureCitation>
)
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alt., montane rainforest, ground nest, 
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et 
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],
</materialsCitation>
<materialsCitation id="3B7C547DF506941DFFAB412FDE8A8B60" ID-GBIF-Occurrence="3446753308" box="[151,478,870,894]" collectionCode="NHMB" pageId="11" pageNumber="12" specimenCode="CASENT0205999" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
<specimenCode id="DBB2F65BF506941DFFAB412FDE328B60" box="[151,358,870,894]" pageId="11" pageNumber="12">CASENT0205999</specimenCode>
[
<collectionCode id="ED05C6E5F506941DFE4B412EDE878B60" box="[375,467,871,894]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="11" pageNumber="12">NHMB</collectionCode>
],
</materialsCitation>
with same data as 
<typeStatus id="54AFE082F506941DFDF8412FDC7E8B60" box="[708,810,870,894]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
; 
<materialsCitation id="3B7C547DF506941DFC05412FDC7E8BD8" ID-GBIF-Occurrence="3446753305" collectingDate="2011-02-23" collectingDateMax="2011-02-24" collectingDateMin="2011-02-23" collectingMethod="Malaise trap" collectionCode="CASC" collectorName="B. L. Fisher &amp; al." country="MADAGASCAR" elevation="984" latitude="-12.513833" location="Parc National Montagne d'Ambre" longLatPrecision="13" longitude="49.177834" municipality="Antsiranana" pageId="11" pageNumber="12" specimenCode="CASENT0205990, BLF26044" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D1295A9F506941DFC05412FDCDE8B60" box="[825,906,870,894]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
, 
<specimenCode id="DBB2F65BF506941DFCA5412FDB3F8B60" box="[921,1131,870,894]" pageId="11" pageNumber="12">CASENT0205990</specimenCode>
, 
<specimenCode id="DBB2F65BF506941DFB46412FDBA38B60" box="[1146,1271,870,894]" pageId="11" pageNumber="12">BLF26044</specimenCode>
: 
<collectingCountry id="F3031EB0F506941DFA39412FDF878BBC" pageId="11" pageNumber="12">MADAGASCAR</collectingCountry>
, 
<collectingMunicipality id="6BCFC45AF506941DFFE341C3DE3D8BBC" box="[223,361,906,930]" pageId="11" pageNumber="12">Antsiranana</collectingMunicipality>
, 
<location id="8ECB08FBF506941DFE4A41C2DDA18BBC" LSID="urn:lsid:plazi:treatment:03BDEF36F5069418FFAB4025DD158A41:8ECB08FBF506941DFE4A41C2DDA18BBC" box="[374,757,906,931]" country="MADAGAS- CAR" latitude="-12.513833" longLatPrecision="13" longitude="49.177834" municipality="Antsiranana" name="Parc National Montagne d'Ambre" pageId="11" pageNumber="12">Parc National Montagne d'Ambre</location>
, 
<geoCoordinate id="EE2038E7F506941DFC3D41C3DCC68BBC" box="[769,914,906,930]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="9" value="-12.513833">12° 30.83´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F506941DFC9C41C3DB678BBC" box="[928,1075,906,930]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="9" value="49.177834">49° 10.67´E</geoCoordinate>
, 
<quantity id="4CECF3C5F506941DFB0341C3DBD38BBC" box="[1087,1159,906,930]" metricMagnitude="2" metricUnit="m" metricValue="9.84" pageId="11" pageNumber="12" unit="m" value="984.0">
<elevation id="0039B913F506941DFB0341C3DBD38BBC" box="[1087,1159,906,930]" metricMagnitude="2" metricUnit="m" metricValue="9.84" pageId="11" pageNumber="12" unit="m" value="984.0">984 m</elevation>
</quantity>
alt., montane rainforest, 
<collectingMethod id="52552637F506941DFFAB41E7DE708BD8" box="[151,292,942,966]" pageId="11" pageNumber="12">Malaise trap</collectingMethod>
, 
<date id="FFAA78E0F506941DFE0C41E7DE998BD8" box="[304,461,942,966]" pageId="11" pageNumber="12" value="2011-02-23" valueMax="2011-02-24" valueMin="2011-02-23">
<collectingDate id="EFEE8108F506941DFE0C41E7DE998BD8" box="[304,461,942,966]" pageId="11" pageNumber="12" value="2011-02-23" valueMax="2011-02-24" valueMin="2011-02-23">23–24.ii.2011</collectingDate>
</date>
, 
<collectorName id="26E13BF6F506941DFEE441E6DD038BD8" box="[472,599,942,966]" pageId="11" pageNumber="12">B.L. Fisher</collectorName>
<emphasis id="B9608232F506941DFD6241F9DDC38BD8" box="[606,663,942,967]" italics="true" pageId="11" pageNumber="12">
et 
<collectorName id="26E13BF6F506941DFD4541E7DDC38BD8" box="[633,663,942,966]" pageId="11" pageNumber="12">al.</collectorName>
</emphasis>
leg. [
<collectionCode id="ED05C6E5F506941DFDE841E7DC778BD8" box="[724,803,942,966]" pageId="11" pageNumber="12">CASC</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF506941AFFF5419ADE0F8FC0" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BAB5E20F506941DFFF5419ADB728C10" blockId="11.[151,1436,726,2011]" pageId="11" pageNumber="12">
<emphasis id="B9608232F506941DFFF5419ADE928BF5" bold="true" box="[201,454,978,1003]" pageId="11" pageNumber="12">Worker. Description.</emphasis>
Measurements: 
<typeStatus id="54AFE082F506941DFDB9419BDDBF8BF4" box="[645,747,978,1002]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
. HL 0.85, HW 0.8, SL 0.45, HD 0.5, WL 1.03, PnW 0.46, MnW 0.25, PpW 0.41, PtW 0.38, CI 93.6, SI 56.9, MnI 31.8, PpI 159.8, PtI 94.1.
</paragraph>
<paragraph id="8BAB5E20F506941DFFF54652DED28C64" blockId="11.[151,1436,726,2011]" pageId="11" pageNumber="12">
HL 0.84–0.9, HW 0.73–0.8, SL 0.46–0.48, HD 0.48–0.55, WL 0.98–1.08, PnW 0.45–0.48, MnW 0.24–0.27, PpW 0.38–0.43, PtW 0.37–0.41, CI 86.7–89.5, SI 59.7–63.2, MnI 30.7–34, PpI 152.8–168.8, PtI 93.1–97.7 (4 
<typeStatus id="54AFE082F506941DFFAB462ADE568C65" box="[151,258,1123,1147]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
measured).
</paragraph>
<paragraph id="8BAB5E20F506941DFFF546CFDC7C8FA8" blockId="11.[151,1436,726,2011]" pageId="11" pageNumber="12">
Head subquadrate in full-face view: longer than wide, sides slightly convex, wider around midlength of head capsule, weakly converging anteriorly and posteriorly, posterior margin concave (
<figureCitation id="132F42A5F506941DFB1F46E3DB248CDC" box="[1059,1136,1194,1218]" captionStart="FIGURES 81 – 84" captionStartId="26.[151,264,1265,1288]" captionTargetBox="[151,1435,179,1240]" captionTargetId="figure@26.[151,1436,179,1257]" captionTargetPageId="26" captionText="FIGURES 81 – 84. Head of Adetomyrma workers in full-face view. 81, A. bressleri sp. nov. (CASENT 0205995: holotype); 82, A. caputleae sp. nov. (CASENT 0004358); 83, A. goblin sp. nov. (CASENT 0227981: holotype); 84, A. venatrix (MCZTYPE 34786)." httpUri="https://zenodo.org/record/213835/files/figure.png" pageId="11" pageNumber="12">Fig 81</figureCitation>
). Clypeal principal surface deflected ventrally. Anterior margin of clypeus broadly convex, furnished with row of about 26 small, specialized, conical setae. Frontal carinae in full-face view short and low, expanded laterally as small frontal lobes about 1.5× wider than length of antennal insertions. Mandible subfalcate, without distinct basal and masticatory margins, inner margin with two apical teeth and five basal denticles (
<figureCitation id="132F42A5F506941DFDCF4773DC668D4C" box="[755,818,1338,1362]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="11" pageNumber="12">Fig 7</figureCitation>
): one small denticle on base of subapical tooth (second tooth from apex) (
<figureCitation id="132F42A5F506941DFE594717DEF78D68" box="[357,419,1374,1398]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="11" pageNumber="12">Fig 7</figureCitation>
); out of five denticles, second and fifth denticles from apical-most one larger than others; second denticle largest (longest); third denticle small and adjacent with second one, and division between them unclear on its base; basal-most denticle sometimes divided into two adjacent small, low denticles (observed in CASENT0205999). Palpal formula 3,3 (three maxillary and three labial). Antennal scape shorter than head length. Pedicel (second antennal segment) shorter than combined length of next three (third to fifth) antennal segments. Antenna gradually broadened from third segment and not forming distinct club. Dorsal outline of mesosoma in lateral view somewhat arched, mesonotum raised from pronotal and propodeal dorsum, metanotal groove often clearly impressed. Pronotum in dorsal view longer than broad, with convex sides. Mesonotum in dorsal view short, slightly wider than long. Metapleuron fully fused with propodeum, but divided by shallow furrow. Dorsal face of propodeum almost as wide as pronotum, less than 1.5× longer than wide, sides strongly converging toward mesonotum (
<figureCitation id="132F42A5F506941DFE17448FDE288EC0" box="[299,380,1734,1758]" captionStart="FIGURES 73 – 80" captionStartId="25.[151,264,1929,1952]" captionTargetBox="[217,1374,193,1897]" captionTargetId="figure@25.[217,1376,193,1907]" captionTargetPageId="25" captionText="FIGURES 73 – 80. Workers of Adetomyrma in lateral view and dorsal view. 73, 74, A. bressleri sp. nov. (CASENT 0205995: holotype); 75, 76, A. caputleae sp. nov. (CASENT 0004358); 77, 78, A. goblin sp. nov. (CASENT 0227981: holotype); 79, 80, A. venatrix (MCZTYPE 34786). 73, 75, 77, 79, lateral view; 74, 76, 78, 80, dorsal view." httpUri="https://zenodo.org/record/213834/files/figure.png" pageId="11" pageNumber="12">Fig 74</figureCitation>
). Propodeal dorsal margin in lateral view less than 2× the length of the declivitous margin, rounding gently into the latter. Propodeal spiracle large, located close to propodeal dorsal margin in lateral view, fully visible in dorsal view. Subpetiolar process distinctly developed, shaped like an irregular axe blade. Shallow and sparse punctures covering head, mesosoma, and abdomen. Clypeus with two groups of hairs; mid clypeal hairs directed dorsally, and anterior clypeal hairs deflected anteriorly. Mid clypeal hairs consisting of one long hair and one or two shorter hairs around the long one. With clypeus in full-face view, anterior clypeal hairs consisting of one pair of long hairs and a single shorter hair laterally (
<figureCitation id="132F42A5F506941DFDE045D7DC4E8FA8" box="[732,794,1950,1974]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="11" pageNumber="12">Fig 7</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F506941DFFF5458BDED98FC4" blockId="11.[151,1436,726,2011]" box="[201,397,1986,2011]" pageId="11" pageNumber="12">
<emphasis id="B9608232F506941DFFF5458BDE438FC5" bold="true" box="[201,279,1986,2011]" pageId="11" pageNumber="12">Queen</emphasis>
unknown.
</paragraph>
<paragraph id="8BAB5E20F501941AFFF542D1DA1488CA" blockId="12.[151,1437,151,391]" pageId="12" pageNumber="13">
<emphasis id="B9608232F501941AFFF542D1DEF188AE" bold="true" box="[201,421,151,176]" pageId="12" pageNumber="13">Male. Description.</emphasis>
Measurements: HL 0.82–0.86, HW 1.18–1.26, SL 0.27–0.3, EL 0.44–0.51, WL 2.22–2.35, MnW 1.55–1.69, CI 140.4–147.2, SI 22.9–24.4, EI 54.1–59.3, MnI 129.3–140.6 (5 individuals measured).
</paragraph>
<paragraph id="8BAB5E20F501941AFFF542A9DED08999" blockId="12.[151,1437,151,391]" pageId="12" pageNumber="13">
Eye well-developed and prominent, posterior margin not exceeding posterior margin of mid ocellus in fullface view. Distance between lateral ocellus and eye long, 3× longer than diameter of lateral ocellus. Palpal formula 3,3 (three maxillary and three labial). Notaulus absent on mesoscutum (
<figureCitation id="132F42A5F501941AFC83436EDB598921" box="[959,1037,295,319]" captionStart="FIGURES 38 – 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 – 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="12" pageNumber="13">Fig 39</figureCitation>
). Parapsidal line clearly impressed. Anterior margin of petiole longer than dorsal margin in lateral view. Subpetiolar process well-developed, with dense hairs ventrally.
</paragraph>
<caption id="DF6B0EA8F501941AFFAB47CFDD8B8E06" httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1435,447,1381]" targetPageId="12">
<paragraph id="8BAB5E20F501941AFFAB47CFDD8B8E06" blockId="12.[151,1436,1414,1560]" pageId="12" pageNumber="13">
<emphasis id="B9608232F501941AFFAB47CFDE028D82" bold="true" box="[151,342,1414,1437]" pageId="12" pageNumber="13">FIGURES 29–37.</emphasis>
Males of 
<taxonomicName id="4C1425A3F501941AFEFE47CEDD178D83" box="[450,579,1415,1437]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F501941AFEFE47CEDD178D83" box="[450,579,1415,1437]" italics="true" pageId="12" pageNumber="13">Adetomyrma</emphasis>
</taxonomicName>
in lateral view. 29, 
<taxonomicName id="4C1425A3F501941AFC2B47C1DCE18D83" authority="Yoshimura &amp; Fisher 2012" box="[791,949,1416,1437]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aureocuprea" status="sp. nov.">
<emphasis id="B9608232F501941AFC2B47C1DCE18D83" box="[791,949,1416,1437]" italics="true" pageId="12" pageNumber="13">A. aureocuprea</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFC8247CEDB458D82" bold="true" box="[958,1041,1415,1436]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFC8247CEDB458D82" box="[958,1041,1415,1436]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0227991: holotype); 30, 
<taxonomicName id="4C1425A3F501941AFABA47C1DFBB8DA2" authority="Yoshimura &amp; Fisher 2012" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bressleri" status="sp. nov.">
<emphasis id="B9608232F501941AFABA47C1DFBB8DA2" italics="true" pageId="12" pageNumber="13">A. bressleri</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFE3F47EFDE358DA5" bold="true" box="[259,353,1446,1467]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFE3F47EFDE358DA5" box="[259,353,1446,1467]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0008693); 31, 
<taxonomicName id="4C1425A3F501941AFDAE47EFDC4B8DA2" authority="Yoshimura &amp; Fisher 2012" box="[658,799,1446,1468]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="caputleae" status="sp. nov.">
<emphasis id="B9608232F501941AFDAE47EFDC4B8DA2" box="[658,799,1446,1468]" italics="true" pageId="12" pageNumber="13">A. caputleae</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFC0F47EFDCC58DA5" bold="true" box="[819,913,1446,1467]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFC0F47EFDCC58DA5" box="[819,913,1446,1467]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0079552); 32, 
<taxonomicName id="4C1425A3F501941AFBFE47EFDA7D8DA2" authority="Yoshimura &amp; Fisher 2012" box="[1218,1321,1446,1468]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cassis" status="sp. nov.">
<emphasis id="B9608232F501941AFBFE47EFDA7D8DA2" box="[1218,1321,1446,1468]" italics="true" pageId="12" pageNumber="13">A. cassis</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFA0147EFDACF8DA5" bold="true" box="[1341,1435,1446,1467]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFA0147EFDACF8DA5" box="[1341,1435,1446,1467]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0163620: holotype); 33, 
<taxonomicName id="4C1425A3F501941AFD24478CDD868DC4" authority="Yoshimura &amp; Fisher 2012" box="[536,722,1476,1498]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="caudapinniger" status="sp. nov.">
<emphasis id="B9608232F501941AFD24478CDD868DC4" box="[536,722,1476,1498]" italics="true" pageId="12" pageNumber="13">A. caudapinniger</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFDDD478CDC6F8DC4" bold="true" box="[737,827,1477,1498]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFDDD478CDC6F8DC4" box="[737,827,1477,1498]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0244415: holotype); 34, 
<taxonomicName id="4C1425A3F501941AFBF1478CDA668DC4" authority="Yoshimura &amp; Fisher 2012" box="[1229,1330,1476,1498]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cilium" status="sp. nov.">
<emphasis id="B9608232F501941AFBF1478CDA668DC4" box="[1229,1330,1476,1498]" italics="true" pageId="12" pageNumber="13">A. cilium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2533F49F501941AFA7E478CDACF8DC4" box="[1346,1435,1477,1498]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B9608232F501941AFA7E478CDACF8DC4" bold="true" box="[1346,1435,1477,1498]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
(CASENT0007808: holotype); 35, 
<taxonomicName id="4C1425A3F501941AFD1947ADDDFE8DE7" authority="Yoshimura &amp; Fisher 2012" box="[549,682,1507,1529]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="clarivida" status="sp. nov.">
<emphasis id="B9608232F501941AFD1947ADDDFE8DE7" box="[549,682,1507,1529]" italics="true" pageId="12" pageNumber="13">A. clarivida</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFD8247AADC488DE6" bold="true" box="[702,796,1507,1528]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFD8247AADC488DE6" box="[702,796,1507,1528]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0064259: holotype); 36, 
<taxonomicName id="4C1425A3F501941AFB8347ADDA7E8DE7" authority="Yoshimura &amp; Fisher 2012" box="[1215,1322,1507,1529]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="goblin" status="sp. nov.">
<emphasis id="B9608232F501941AFB8347ADDA7E8DE7" box="[1215,1322,1507,1529]" italics="true" pageId="12" pageNumber="13">A. goblin</emphasis>
</taxonomicName>
<emphasis id="B9608232F501941AFA0147AADACF8DE6" bold="true" box="[1341,1435,1507,1528]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A2533F49F501941AFA0147AADACF8DE6" box="[1341,1435,1507,1528]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0084070); 37, 
<taxonomicName id="4C1425A3F501941AFEAA444BDD518E06" box="[406,517,1538,1560]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F501941AFEAA444BDD518E06" box="[406,517,1538,1560]" italics="true" pageId="12" pageNumber="13">A. venatrix</emphasis>
</taxonomicName>
(CASENT0151606).
</paragraph>
</caption>
<paragraph id="8BAB5E20F501941AFFF54417DE0F8F50" blockId="12.[151,1436,1630,2014]" pageId="12" pageNumber="13">
Left and right parameres broadly overlapping along almost entire length of basimere (
<figureCitation id="132F42A5F501941AFB9C4417DBBA8E68" box="[1184,1262,1630,1654]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="12" pageNumber="13">Fig 13</figureCitation>
). Distinct, flattened, and needle-like projection present on each posterodorsal portion of paramere (
<figureCitation id="132F42A5F501941AFB7044C8DBCD8E87" box="[1100,1177,1665,1689]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="12" pageNumber="13">Fig 13</figureCitation>
). Basal ring reduced to about 0.33× of parameral maximum height in lateral view. Basal projection on cuspis extraordinarily well developed, as large as digitus, and replacing the usual position of cuspis; cuspis also well developed and elongate, and surrounding digitus. Aedeagus in lateral view as in 
<figureCitation id="132F42A5F501941AFDD744A7DC088F18" box="[747,860,1774,1798]" captionStart="FIGURES 56 – 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 – 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="12" pageNumber="13">Figure 57</figureCitation>
; mid portion distinctly widened and distal portion extremely narrowed; ventral projection vestigial and inconspicuous; its apical portion narrowed but hard, and curved as a hook.
</paragraph>
<paragraph id="8BAB5E20F501941AFFF54510DE0F8FC0" blockId="12.[151,1436,1630,2014]" pageId="12" pageNumber="13">
Hair on compound eyes long, almost as long as diameter of mid ocellus (
<figureCitation id="132F42A5F501941AFB304510DB088F6F" box="[1036,1116,1881,1905]" captionStart="FIGURES 47 – 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 – 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="12" pageNumber="13">Fig 48</figureCitation>
). Mesofemur in dorsal view with anterior face covered in dense subdecumbent hairs. Ventral margin of eye edged with darker pigment but without minute punctures on the area. Body yellow with bicolored head, brown on its dorsal face and yellow on its ventral face (
<figureCitation id="132F42A5F501941AFE3C458FDE1A8FC0" box="[256,334,1990,2014]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="12" pageNumber="13">Fig 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF500941BFFF542D1DD538902" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BAB5E20F500941BFFF542D1DD538902" blockId="13.[151,1437,151,644]" pageId="13" pageNumber="14">
<emphasis id="B9608232F500941BFFF542D1DE1B88AE" bold="true" box="[201,335,152,176]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific epithet is a patronymic referring to Dr. Barry Lee Bressler, retired physicist, former adjunct professor of physics at Virginia Polytechnic Institute and State University, and amateur naturalist, in recognition of his interest in myrmecology and his support for research on ants. The species epithet is treated as a noun in apposition, and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF500941BFFF5436EDDB98921" box="[201,749,295,320]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BAB5E20F500941BFFF5436EDDB98921" blockId="13.[151,1437,151,644]" box="[201,749,295,320]" pageId="13" pageNumber="14">
<emphasis id="B9608232F500941BFFF5436EDE36895E" bold="true" box="[201,354,295,320]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingCountry id="F3031EB0F500941BFE55436EDD618921" box="[361,565,295,319]" name="Madagascar" pageId="13" pageNumber="14">MADAGASCAR</collectingCountry>
: as in 
<figureCitation id="132F42A5F500941BFD46436EDDBD8921" box="[634,745,295,319]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="13" pageNumber="14">Figure 65</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF500941BFFF54302DB2F8A9A" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BAB5E20F500941BFFF54302DB2F8A9A" blockId="13.[151,1437,151,644]" pageId="13" pageNumber="14">
<emphasis id="B9608232F500941BFFF54302DD7A897A" bold="true" box="[201,558,331,356]" pageId="13" pageNumber="14">Additional material examined</emphasis>
: in addition to the 
<typeStatus id="54AFE082F500941BFC3F4304DC67897B" box="[771,819,333,357]" pageId="13" pageNumber="14">type</typeStatus>
material, specimens from the following localities were examined in this study: 
<collectingCountry id="F3031EB0F500941BFE984326DD3B8999" box="[420,623,367,391]" name="Madagascar" pageId="13" pageNumber="14">MADAGASCAR</collectingCountry>
. Sakalava Beach, vegetated beach dunes, 
<geoCoordinate id="EE2038E7F500941BFB7A4326DB818999" box="[1094,1237,367,391]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-12.262834">12° 15.77´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFBDE4326DA278999" box="[1250,1395,367,391]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="49.3975">49° 23.85´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFABC4326DFF989B2" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="10.0">10 m</quantity>
alt., across sandy trail in dwarf littoral forest; 
<quantity id="4CECF3C5F500941BFD8B43DCDDA589B2" box="[695,753,404,428]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="km" value="7.0">7 km</quantity>
N Joffreville, camp 2 of Fisher, 
<geoCoordinate id="EE2038E7F500941BFB5C43DDDB9989B2" box="[1120,1229,404,428]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-12.333333">12° 20´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFBE643DDDA1C89B2" box="[1242,1352,404,428]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="49.25">49° 15´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFA6943DDDAC889B2" box="[1365,1436,404,428]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="13" pageNumber="14" unit="m" value="360.0">360 m</quantity>
alt., in dry forest; Parc National Montagne d'Ambre, 
<geoCoordinate id="EE2038E7F500941BFDCB43FEDCDD89D1" box="[759,905,439,463]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-12.513833">12° 30.83´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFCA443FEDB7889D1" box="[920,1068,439,463]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="49.177834">49° 10.67´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFB0643FEDBD789D1" box="[1082,1155,439,463]" metricMagnitude="2" metricUnit="m" metricValue="9.84" pageId="13" pageNumber="14" unit="m" value="984.0">984 m</quantity>
alt., montane rainforest; Parc National Montagne d'Ambre, first campsite, 
<geoCoordinate id="EE2038E7F500941BFDF54395DC0D89EA" box="[713,857,476,500]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-12.5145">12° 30.87´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFC5A4395DCAD89EA" box="[870,1017,476,500]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="49.18133">49° 10.88´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFB3A4395DB1989EA" box="[1030,1101,476,500]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="13" pageNumber="14" unit="m" value="960.0">960 m</quantity>
alt., rainforest; Parc National Montagne d'Ambre, lemur trail, 
<geoCoordinate id="EE2038E7F500941BFEC343B6DD3E8A09" box="[511,618,511,535]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-12.516666">12° 31´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFD4A43B6DDB58A09" box="[630,737,511,535]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="49.183334">49° 11´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFDD143B6DC678A09" box="[749,819,511,535]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="13" pageNumber="14" unit="m" value="975.0">975 m</quantity>
alt., rainforest; Parc National Montagne d'Ambre, Petit Lac road, 
<geoCoordinate id="EE2038E7F500941BFE3B406DDEC08A22" box="[263,404,548,572]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-12.520333">12° 31.22´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFE9C406DDD648A22" box="[416,560,548,572]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="49.179165">49° 10.75´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFD00406DDDDB8A22" box="[572,655,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="13" pageNumber="14" unit="m" value="1125.0">1125 m</quantity>
alt., rainforest; Parc National Montagne d'Ambre, 
<quantity id="4CECF3C5F500941BFBFA406DDA758A22" box="[1222,1313,548,572]" metricMagnitude="4" metricUnit="m" metricValue="1.22" pageId="13" pageNumber="14" unit="km" value="12.2">12.2 km</quantity>
211° SSW Joffreville, 
<geoCoordinate id="EE2038E7F500941BFE2B400EDEF08A41" box="[279,420,583,607]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-12.5963335">12° 35.78´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFE8C400EDD678A41" box="[432,563,583,607]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="49.1595">49° 9.57´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFD01400EDDC68A41" box="[573,658,583,607]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="14" unit="m" value="1300.0">1300 m</quantity>
alt., montane rainforest; Andasibe National Park, botanic garden near entrance, west of 
<collectionCode id="ED05C6E5F500941BFE604025DEE98A9A" box="[348,445,620,644]" pageId="13" pageNumber="14">ANGAP</collectionCode>
office, 
<geoCoordinate id="EE2038E7F500941BFD2F4025DDF58A9A" box="[531,673,620,644]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="9" value="-18.926332">18° 55.58´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F500941BFD914025DC6B8A9A" box="[685,831,620,644]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="9" value="48.407833">48° 24.47´E</geoCoordinate>
, 
<quantity id="4CECF3C5F500941BFC764025DCCB8A9A" box="[842,927,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="13" pageNumber="14" unit="m" value="1025.0">1025 m</quantity>
alt., tropical forest.
</paragraph>
</subSubSection>
<caption id="DF6B0EA8F500941BFFAB47C6DD8B8E3E" httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1432,685,1400]" targetPageId="13">
<paragraph id="8BAB5E20F500941BFFAB47C6DD8B8E3E" blockId="13.[151,1436,1423,1569]" pageId="13" pageNumber="14">
<emphasis id="B9608232F500941BFFAB47C6DE028DBB" bold="true" box="[151,342,1423,1446]" pageId="13" pageNumber="14">FIGURES 38–46.</emphasis>
Males of 
<taxonomicName id="4C1425A3F500941BFEFF47D9DD108DB8" box="[451,580,1424,1446]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F500941BFEFF47D9DD108DB8" box="[451,580,1424,1446]" italics="true" pageId="13" pageNumber="14">Adetomyrma</emphasis>
</taxonomicName>
in dorsal view. 38, 
<taxonomicName id="4C1425A3F500941BFC2B47D8DCE18DB8" authority="Yoshimura &amp; Fisher 2012" box="[791,949,1425,1446]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aureocuprea" status="sp. nov.">
<emphasis id="B9608232F500941BFC2B47D8DCE18DB8" box="[791,949,1425,1446]" italics="true" pageId="13" pageNumber="14">A. aureocuprea</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFC8147D9DB448DBB" bold="true" box="[957,1040,1424,1445]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFC8147D9DB448DBB" box="[957,1040,1424,1445]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0227991: holotype); 39, 
<taxonomicName id="4C1425A3F500941BFABA47D8DFBB8DDB" authority="Yoshimura &amp; Fisher 2012" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bressleri" status="sp. nov.">
<emphasis id="B9608232F500941BFABA47D8DFBB8DDB" italics="true" pageId="13" pageNumber="14">A. bressleri</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFE3F47E6DE358DDA" bold="true" box="[259,353,1455,1476]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFE3F47E6DE358DDA" box="[259,353,1455,1476]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0008693); 40, 
<taxonomicName id="4C1425A3F500941BFDAE47E6DC4B8DDB" authority="Yoshimura &amp; Fisher 2012" box="[658,799,1455,1477]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="caputleae" status="sp. nov.">
<emphasis id="B9608232F500941BFDAE47E6DC4B8DDB" box="[658,799,1455,1477]" italics="true" pageId="13" pageNumber="14">A. caputleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2533F49F500941BFC0F47E6DCC58DDA" box="[819,913,1455,1476]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B9608232F500941BFC0F47E6DCC58DDA" bold="true" box="[819,913,1455,1476]" pageId="13" pageNumber="14">sp. nov.</emphasis>
</taxonomicNameLabel>
(CASENT0079552); 41, 
<taxonomicName id="4C1425A3F500941BFBFE47E6DA7D8DDB" authority="Yoshimura &amp; Fisher 2012" box="[1218,1321,1455,1477]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cassis" status="sp. nov.">
<emphasis id="B9608232F500941BFBFE47E6DA7D8DDB" box="[1218,1321,1455,1477]" italics="true" pageId="13" pageNumber="14">A. cassis</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFA0147E6DACF8DDA" bold="true" box="[1341,1435,1455,1476]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFA0147E6DACF8DDA" box="[1341,1435,1455,1476]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0163620: holotype); 42, 
<taxonomicName id="4C1425A3F500941BFD244787DD868DFD" authority="Yoshimura &amp; Fisher 2012" box="[536,722,1485,1507]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="caudapinniger" status="sp. nov.">
<emphasis id="B9608232F500941BFD244787DD868DFD" box="[536,722,1485,1507]" italics="true" pageId="13" pageNumber="14">A. caudapinniger</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFDDD4787DC6F8DFD" bold="true" box="[737,827,1486,1507]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFDDD4787DC6F8DFD" box="[737,827,1486,1507]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0244415: holotype); 43, 
<taxonomicName id="4C1425A3F500941BFBF14787DA668DFD" authority="Yoshimura &amp; Fisher 2012" box="[1229,1330,1485,1507]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cilium" status="sp. nov.">
<emphasis id="B9608232F500941BFBF14787DA668DFD" box="[1229,1330,1485,1507]" italics="true" pageId="13" pageNumber="14">A. cilium</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFA7E4787DACF8DFD" bold="true" box="[1346,1435,1486,1507]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFA7E4787DACF8DFD" box="[1346,1435,1486,1507]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0007808: holotype); 44, 
<taxonomicName id="4C1425A3F500941BFD1947A5DDFE8E1C" authority="Yoshimura &amp; Fisher 2012" box="[549,682,1516,1538]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="clarivida" status="sp. nov.">
<emphasis id="B9608232F500941BFD1947A5DDFE8E1C" box="[549,682,1516,1538]" italics="true" pageId="13" pageNumber="14">A. clarivida</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFD8247A5DC488E1F" bold="true" box="[702,796,1516,1537]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFD8247A5DC488E1F" box="[702,796,1516,1537]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0064259: holotype); 45, 
<taxonomicName id="4C1425A3F500941BFB8347A5DA7E8E1C" authority="Yoshimura &amp; Fisher 2012" box="[1215,1322,1516,1538]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="goblin" status="sp. nov.">
<emphasis id="B9608232F500941BFB8347A5DA7E8E1C" box="[1215,1322,1516,1538]" italics="true" pageId="13" pageNumber="14">A. goblin</emphasis>
</taxonomicName>
<emphasis id="B9608232F500941BFA0147A5DACF8E1F" bold="true" box="[1341,1435,1516,1537]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A2533F49F500941BFA0147A5DACF8E1F" box="[1341,1435,1516,1537]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CASENT0084070); 46, 
<taxonomicName id="4C1425A3F500941BFEAA4442DD518E3E" box="[406,517,1547,1568]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F500941BFEAA4442DD518E3E" box="[406,517,1547,1568]" italics="true" pageId="13" pageNumber="14">A. venatrix</emphasis>
</taxonomicName>
(CASENT0151606).
</paragraph>
</caption>
<subSubSection id="C30E0DABF5009418FFF54401DB868A22" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BAB5E20F500941BFFF54401DB098F0A" blockId="13.[151,1436,1607,2028]" pageId="13" pageNumber="14">
<emphasis id="B9608232F500941BFFF54401DE698E7E" bold="true" box="[201,317,1608,1632]" pageId="13" pageNumber="14">Remarks.</emphasis>
Workers of 
<taxonomicName id="4C1425A3F500941BFEF44401DDEB8E7E" box="[456,703,1608,1632]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F500941BFEF44401DDEB8E7E" box="[456,703,1608,1632]" italics="true" pageId="13" pageNumber="14">Adetomyrma bressleri</emphasis>
</taxonomicName>
are clearly distinguished from other genus members by a combination of a single shorter hair lateral to the longest one on the anterior clypeal margin in full-face view, the presence of a small denticle on the basal portion of the mandibular subapical tooth, and a large propodeal spiracle that is located posterodorsally and fully visible in dorsal view. 
<taxonomicName id="4C1425A3F500941BFC3644FDDCDE8ED2" box="[778,906,1716,1740]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F500941BFC3644FDDCDE8ED2" box="[778,906,1716,1740]" italics="true" pageId="13" pageNumber="14">A. bressleri</emphasis>
</taxonomicName>
is easily distinguished from other 
<taxonomicName id="4C1425A3F500941BFA3144FDDAC88ED2" box="[1293,1436,1716,1740]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F500941BFA3144FDDAC88ED2" box="[1293,1436,1716,1740]" italics="true" pageId="13" pageNumber="14">Adetomyrma</emphasis>
</taxonomicName>
males by its giant yellow body, parameres overlapping broadly on almost the whole length of the basimere, and a flattened, needle-like projection on the posterodorsal portion of the paramere (
<figureCitation id="132F42A5F500941BFB3F44B5DB048F0A" box="[1027,1104,1788,1812]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1025,1048]" captionTargetBox="[151,1435,369,1003]" captionTargetId="figure@6.[151,1436,369,1004]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Terminal abdominal segments of Adetomyrma males in oblique dorsal view. 13, A. bressleri (CASENT 0008693); 14, A. cassis (CASENT 0163620: holotype). 13, paramere is broadly overlapping dorsally, and with a deep notch separating between a needle-like projection and paramere; 14, paramere is not or narrowly overlapping dorsally, and without a deep notch between a needle-like projection and the paramere." httpUri="https://zenodo.org/record/213822/files/figure.png" pageId="13" pageNumber="14">Fig 13</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F5009418FFF54556DE8188E9" blockId="13.[151,1436,1607,2028]" lastBlockId="14.[151,1436,151,607]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
The males of 
<taxonomicName id="4C1425A3F500941BFE5D4569DEB48F26" box="[353,480,1824,1848]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F500941BFE5D4569DEB48F26" box="[353,480,1824,1848]" italics="true" pageId="13" pageNumber="14">A. bressleri</emphasis>
</taxonomicName>
(= mg03: 
<bibRefCitation id="EF8523D1F500941BFD694556DC2C8F29" author="Yoshimura &amp; Fisher" box="[597,888,1823,1847]" firstAuthor="Yoshimura" pageId="13" pageNumber="14" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">Yoshimura &amp; Fisher 2012</bibRefCitation>
) have been collected numerous times by Malaise traps at the higher elevations of Montagne d'Ambre (
<quantity id="4CECF3C5F500941BFDDA450DDC268F42" box="[742,882,1860,1884]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="13" pageNumber="14" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">900–1300m</quantity>
). The first collection date from Malaise traps was in 2001. Since then, three expeditions to Montagne d'Ambre have been organized to find the female castes, but not until 2011 was the worker caste finally found. To find the worker caste, Malaise traps were set up and monitored for the collection of males. Once males were collected, logs and branches on the ground around the trap were inspected. On the last day of the trip in 2011, workers were found in a log near a trap that had collected males. As the log was pulled apart for worker ants (BLF26111), the nearby Malaise trap (BLF26044), which was moved directly above the log, captured males leaving the log. These collections did not yield a visible queen but included 
<specimenCount id="9D1295A9F5039418FFAB4296DE7688E9" box="[151,290,223,247]" pageId="14" pageNumber="15" type="worker">291 workers</specimenCount>
and 
<specimenCount id="9D1295A9F5039418FE644296DE8588E9" box="[344,465,223,247]" pageId="14" pageNumber="15" type="larva">814 larvae</specimenCount>
.
</paragraph>
<paragraph id="8BAB5E20F5039418FFF5434DDA2F8A09" blockId="14.[151,1436,151,607]" pageId="14" pageNumber="15">
Although the males of 
<taxonomicName id="4C1425A3F5039418FEFA434DDD7F8902" box="[454,555,260,284]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="goblin">
<emphasis id="B9608232F5039418FEFA434DDD7F8902" box="[454,555,260,284]" italics="true" pageId="14" pageNumber="15">A. goblin</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F5039418FD06434DDDD08902" box="[570,644,260,284]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="14" pageNumber="15">Fig 71</figureCitation>
) and 
<taxonomicName id="4C1425A3F5039418FDFD434DDC388902" box="[705,876,260,284]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F5039418FDFD434DDC388902" box="[705,876,260,284]" italics="true" pageId="14" pageNumber="15">A. aureocuprea</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F5039418FC46434DDC908902" box="[890,964,260,284]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="14" pageNumber="15">Fig 64</figureCitation>
) were collected from either the same location or localities close to where 
<taxonomicName id="4C1425A3F5039418FEC64361DD2D895E" box="[506,633,296,320]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F5039418FEC64361DD2D895E" box="[506,633,296,320]" italics="true" pageId="14" pageNumber="15">A. bressleri</emphasis>
</taxonomicName>
was found (
<figureCitation id="132F42A5F5039418FC3F436EDC1B8921" box="[771,847,295,319]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="14" pageNumber="15">Fig 65</figureCitation>
), we differentiate the workers of the former two species from the latter with confidence. The association between males and workers in 
<taxonomicName id="4C1425A3F5039418FB784305DBFF897A" box="[1092,1195,332,356]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="goblin">
<emphasis id="B9608232F5039418FB784305DBFF897A" box="[1092,1195,332,356]" italics="true" pageId="14" pageNumber="15">A. goblin</emphasis>
</taxonomicName>
has been clarified by colony series material. The morphological similarities of the males of 
<taxonomicName id="4C1425A3F5039418FC9A4339DB058996" box="[934,1105,368,392]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F5039418FC9A4339DB058996" box="[934,1105,368,392]" italics="true" pageId="14" pageNumber="15">A. aureocuprea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1425A3F5039418FBB44339DA568996" box="[1160,1282,368,392]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F5039418FBB44339DA568996" box="[1160,1282,368,392]" italics="true" pageId="14" pageNumber="15">A. venatrix</emphasis>
</taxonomicName>
strongly suggest that the workers of 
<taxonomicName id="4C1425A3F5039418FE9143DDDD0E89B2" box="[429,602,404,428]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F5039418FE9143DDDD0E89B2" box="[429,602,404,428]" italics="true" pageId="14" pageNumber="15">A. aureocuprea</emphasis>
</taxonomicName>
share several important characters with that of 
<taxonomicName id="4C1425A3F5039418FB4043DDDBAC89B2" box="[1148,1272,404,428]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F5039418FB4043DDDBAC89B2" box="[1148,1272,404,428]" italics="true" pageId="14" pageNumber="15">A. venatrix</emphasis>
</taxonomicName>
, including the lack of a basal denticle on the base of the subapical mandible tooth (as in 
<figureCitation id="132F42A5F5039418FCF243FEDB5D89D1" box="[974,1033,439,463]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="14" pageNumber="15">Fig 8</figureCitation>
), and two shorter hairs lateral to the longest hair on the anterior margin of the clypeus in full-face view (
<figureCitation id="132F42A5F5039418FCBE4395DCEA89EA" box="[898,958,476,500]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="14" pageNumber="15">Fig 8</figureCitation>
). In contrast, the worker of 
<taxonomicName id="4C1425A3F5039418FBCE4395DA2589EA" box="[1266,1393,476,500]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F5039418FBCE4395DA2589EA" box="[1266,1393,476,500]" italics="true" pageId="14" pageNumber="15">A. bressleri</emphasis>
</taxonomicName>
has a basal denticle (
<figureCitation id="132F42A5F5039418FE6D43B6DEDA8A09" box="[337,398,511,535]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="14" pageNumber="15">Fig 7</figureCitation>
), and only a single hair lateral to the longest anterior clypeal hair in full-face view (
<figureCitation id="132F42A5F5039418FA1343B6DA398A09" box="[1327,1389,511,535]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[151,1435,1241,1709]" captionTargetId="figure@4.[151,1436,1241,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Clypeus and mandible of Adetomyrma workers in oblique full-face view. 7, A. bressleri (CASENT 0205995: holotype); 8, A. caputleae (CASENT 0227994: paratype). 7, only a single hair is present lateral of the longest anterior clypeal hair; 8, two or more hairs are present lateral of the longest anterior clypeal hair. 7, a denticle is present on the base of the subapical tooth; 8, no denticle is present on the base of the subapical tooth." httpUri="https://zenodo.org/record/213819/files/figure.png" pageId="14" pageNumber="15">Fig 7</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F5039418FFF5406DDB868A22" blockId="14.[151,1436,151,607]" box="[201,1234,548,572]" pageId="14" pageNumber="15">This new species corresponds to the following species code below used in previous studies:</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF5039418FFF5400EDD158A41" box="[201,577,583,607]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BAB5E20F5039418FFF5400EDD158A41" blockId="14.[151,1436,151,607]" box="[201,577,583,607]" pageId="14" pageNumber="15">
mg03: 
<bibRefCitation id="EF8523D1F5039418FE2B400EDD6A8A41" author="Yoshimura &amp; Fisher" box="[279,574,583,607]" firstAuthor="Yoshimura" pageId="14" pageNumber="15" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">Yoshimura &amp; Fisher 2012</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>