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<document id="CD43002ED7D5F90CF69B71F01D4CEC07" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BD4FF72AF89FA2CFD103E75" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera guanxianensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="184" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="182" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="A991CC4F01A088E00D7B68D7AA1D0E51">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="50BFB40275DA0135649488EA9F7A19CD">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="234424D64B7674CEE8463D8925A82C54">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="18F782DA161D0D4A1F4DC7BAD0707687">Bai, Ming</mods:namePart>
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<mods:namePart id="F19AA022FA39CD2A5B0B9EFBB212AD48">Ahrens, Dirk</mods:namePart>
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<treatment id="03BD87E66BD4FF72AF89FA2CFD103E75" ID-DOI="http://doi.org/10.5281/zenodo.4534658" ID-GBIF-Taxon="177323551" ID-Zenodo-Dep="4534658" LSID="urn:lsid:plazi:treatment:03BD87E66BD4FF72AF89FA2CFD103E75" httpUri="http://treatment.plazi.org/id/03BD87E66BD4FF72AF89FA2CFD103E75" lastPageId="183" lastPageNumber="184" pageId="181" pageNumber="182">
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<heading id="D0E3819C6BD4FF70AF89FA2CFCEA3F2D" bold="true" box="[151,820,582,609]" fontSize="11" level="2" pageId="181" pageNumber="182" reason="4">
<emphasis id="B960EAE26BD4FF70AF89FA2CFCEA3F2D" bold="true" box="[151,820,582,609]" pageId="181" pageNumber="182">
<taxonomicName id="4C144D736BD4FF70AF89FA2CFD3B3F2C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,741,582,609]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="guanxianensis" status="sp. nov.">
<emphasis id="B960EAE26BD4FF70AF89FA2CFE1A3F2D" bold="true" box="[151,452,582,609]" italics="true" pageId="181" pageNumber="182">Maladera guanxianensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD4FF70ADECFA2DFCEA3F2D" box="[754,820,583,609]" pageId="181" pageNumber="182" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD4FF70AF89FA01FEA33FC9" box="[151,381,619,645]" pageId="181" pageNumber="182" type="description">
<paragraph id="8BAB36F06BD4FF70AF89FA01FEA33FC9" blockId="181.[151,820,582,645]" box="[151,381,619,645]" pageId="181" pageNumber="182">
<figureCitation id="132F2A756BD4FF70AF89FA01FED33FC9" box="[151,269,619,645]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="181" pageNumber="182">Figures 41</figureCitation>
A–D, 108
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD4FF70AF89FAD9FBBB39A5" pageId="181" pageNumber="182" type="materials_examined">
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<emphasis id="B960EAE26BD4FF70AF89FAD9FE653F80" bold="true" box="[151,443,690,717]" pageId="181" pageNumber="182">Type material examined.</emphasis>
<typeStatus id="54AF88526BD4FF70AEDEFAD9FDF33F81" box="[448,557,691,717]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
:
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“China-Sichuan pr. Guanxian
<date id="FFAA10306BD4FF70AC84FAD9FC233F81" box="[922,1021,691,717]" pageId="181" pageNumber="182" value="1992-07-08">8.7.1992</date>
J. Schneider/ 552
<taxonomicName id="4C144D736BD4FF70ABDAFAD9FAC53F81" box="[1220,1307,691,717]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
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).
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:
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“China-Sichuan pr. Guanxian
<date id="FFAA10306BD4FF70AC0FFABDFC553FBD" box="[785,907,727,753]" pageId="181" pageNumber="182" value="1992-07-08">8.VII.1992</date>
R. Dunda lgt.” (
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),
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“China-Sichuan pr. Guanxian
<date id="FFAA10306BD4FF70AE30FA91FE4F3E59" box="[302,401,763,789]" pageId="181" pageNumber="182" value="1992-07-08">8.7.1992</date>
J. Schneider lgt.” (
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),
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“
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(W-Hubei) Daba Shan riv. Bank
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SE Muyuping
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/
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(slopes with decid. For./ leaves)
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Wrase [18]” (
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),
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“
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Centr.
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Volong 150 SV Chendu 9.-10.7.94 Benes lg.” (
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),
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,
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“
<collectingCountry id="F30376606BD4FF70ACC9FB29FBC13E11" box="[983,1055,835,861]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
, N
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, 4.-7.VII. Wuling Shan,
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Zhangjiajie, ~
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Jaroslav Turna leg., 2003” (
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),
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“
<collectingCountry id="F30376606BD4FF70AB14FB0DFB8C3ECD" box="[1034,1106,871,897]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
, W
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, 10.VII. Road Badong-Jianshi
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, ~
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~
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W Zhashuping Jaroslav Turna leg., 2003” (
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),
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,
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“
<collectingCountry id="F30376606BD4FF70AA4EFBE1FA463EE9" box="[1360,1432,907,933]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
,
<collectingRegion id="49D0F8126BD4FF70AF89FBC5FF2A3E85" box="[151,244,943,969]" country="China" name="Sichuan" pageId="181" pageNumber="182">Sichuan</collectingRegion>
, 2002, Wenchuan, 19.viii., h~
<quantity id="4CEC9B156BD4FF70AD5BFBC4FD0B3E85" box="[581,725,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="181" pageNumber="182" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500-1600m</quantity>
alt., leg. S. Murzin, I. Shokhin” (
<collectionCode id="ED05AE356BD4FF70AB4EFBC5FBA93E85" box="[1104,1143,943,969]" pageId="181" pageNumber="182">CP</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AB95FBC5FB683E85" box="[1163,1206,943,969]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“
<collectingCountry id="F30376606BD4FF70ABD6FBC5FACE3E85" box="[1224,1296,943,969]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
, NW
<collectingRegion id="49D0F8126BD4FF70AA4FFBC5FF0E3EA1" country="China" name="Hunan" pageId="181" pageNumber="182">Hunan prov.</collectingRegion>
Wulingshan,
<quantity id="4CEC9B156BD4FF70AE74FBB9FE743EA1" box="[362,426,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="181" pageNumber="182" unit="m" value="800.0">800m</quantity>
Tianzishan Nat. Res.
<date id="FFAA10306BD4FF70AD81FBB9FC913EA1" box="[671,847,979,1005]" pageId="181" pageNumber="182" value="1997-06-16" valueMax="1997-06-18" valueMin="1997-06-16">16-18 Jun 1997</date>
, Bolm lgt.” (
<collectionCode id="ED05AE356BD4FF70ACC1FBB9FBD83EA1" box="[991,1030,979,1005]" pageId="181" pageNumber="182">CP</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AB04FBB9FB9A3EA1" box="[1050,1092,979,1005]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“
<collectingCountry id="F30376606BD4FF70AB4AFBB9FB423EA1" box="[1108,1180,979,1005]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
,
<collectingRegion id="49D0F8126BD4FF70ABB9FBB9FADF3EA1" box="[1191,1281,979,1005]" country="China" name="Sichuan" pageId="181" pageNumber="182">Sichuan</collectingRegion>
27.VI.-2.VII. Jiulonggou near Dayi 1995 (=
<collectingRegion id="49D0F8126BD4FF70AEFCFB9DFDBF395D" box="[482,609,1015,1041]" country="China" name="Chongqing" pageId="181" pageNumber="182">Chongqing</collectingRegion>
Jiulong Valley) cca
<quantity id="4CEC9B156BD4FF70AC5DFB9DFC5D395D" box="[835,899,1015,1041]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="181" pageNumber="182" unit="km" value="60.0">60km</quantity>
W of Chengdu
<geoCoordinate id="EE2050376BD4FF70AB2DFB9DFB4D395D" box="[1075,1171,1015,1041]" degrees="31" direction="north" minutes="00" orientation="latitude" pageId="181" pageNumber="182" precision="925" value="31.0">31°00’N</geoCoordinate>
<geoCoordinate id="EE2050376BD4FF70AB84FB9DFADB395D" box="[1178,1285,1015,1041]" degrees="103" direction="east" minutes="30" orientation="longitude" pageId="181" pageNumber="182" precision="925" value="103.5">103°30’E</geoCoordinate>
M. Tryzna &amp; O. Safranek lgt.” (
<collectionCode id="ED05AE356BD4FF70AE7AFC71FE553979" box="[356,395,1051,1077]" pageId="181" pageNumber="182">CP</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AE80FC71FE143979" box="[414,458,1051,1077]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D12FD796BD4FF70AEC8FC71FDCB3979" box="[470,533,1051,1077]" pageId="181" pageNumber="182" type="female">2 ♀♀</specimenCount>
“
<collectingCountry id="F30376606BD4FF70AD38FC71FDB13979" box="[550,623,1051,1077]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
: Near Kwanhsien Alt.
<quantity id="4CEC9B156BD4FF70AC72FC70FC213979" box="[876,1023,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.2954" metricValueMax="1.524" metricValueMin="1.0668" pageId="181" pageNumber="182" unit="ft" value="4250.0" valueMax="5000.0" valueMin="3500.0">3500-5000ft.</quantity>
<date id="FFAA10306BD4FF70AB1AFC70FBB23979" box="[1028,1132,1050,1077]" pageId="181" pageNumber="182" value="1935-07">VII-1935</date>
D.C. Graham” (
<collectionCode id="ED05AE356BD4FF70AA3BFC71FAA23979" box="[1317,1404,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AA90FC71FF723915" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Mt. Omei, Kiating Szechuan,
<collectingCountry id="F30376606BD4FF70AD0FFC55FD8A3915" box="[529,596,1087,1113]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
VII-1-32” (
<collectionCode id="ED05AE356BD4FF70ADC5FC55FCEC3915" box="[731,818,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AC5BFC55FCB13915" box="[837,879,1087,1113]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Szech.
<collectingCountry id="F30376606BD4FF70ACCBFC55FBC63915" box="[981,1048,1087,1113]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
DC Graham/ nr Weichow 7.28-30 ‘33
<quantity id="4CEC9B156BD4FF70AFDAFC08FE863931" box="[196,344,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.2098" metricValueMax="2.71272" metricValueMin="1.70688" pageId="181" pageNumber="182" unit="ft" value="7250.0" valueMax="8900.0" valueMin="5600.0">5600-8900ft.</quantity>
” (
<collectionCode id="ED05AE356BD4FF70AE6FFC09FE163931" box="[369,456,1123,1149]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AEC5FC09FDDB3931" box="[475,517,1123,1149]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Szechwan
<collectingCountry id="F30376606BD4FF70AD8FFC09FD0A3931" box="[657,724,1123,1149]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
DC Graham/ nr Kuanshien
<date id="FFAA10306BD4FF70AB11FC09FB5B3931" box="[1039,1157,1123,1149]" pageId="181" pageNumber="182" value="1933-08">Aug. 1933</date>
<quantity id="4CEC9B156BD4FF70AB93FC09FAFF3931" box="[1165,1313,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="9.144" metricValueMax="12.192" metricValueMin="6.096" pageId="181" pageNumber="182" unit="ft" value="3000.0" valueMax="4000.0" valueMin="2000.0">2000-4000ft.</quantity>
” (
<collectionCode id="ED05AE356BD4FF70AA27FC09FA4E3931" box="[1337,1424,1123,1149]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AF89FCEDFF1F39ED" box="[151,193,1159,1185]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Wen Chuan Shien
<quantity id="4CEC9B156BD4FF70AEBAFCEDFE3B39ED" box="[420,485,1159,1185]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="181" pageNumber="182" unit="mi" value="30.0">30 mi</quantity>
NNW Kuanshien
<quantity id="4CEC9B156BD4FF70ADAAFCEDFCDA39ED" box="[692,772,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="181" pageNumber="182" unit="ft" value="4000.0">4000ft.</quantity>
<date id="FFAA10306BD4FF70AC14FCEDFC4A39ED" box="[778,916,1159,1185]" pageId="181" pageNumber="182" value="1933-07-19">VII-19 1933</date>
/ Szechwan
<collectingCountry id="F30376606BD4FF70AB02FCEDFB8139ED" box="[1052,1119,1159,1185]" name="China" pageId="181" pageNumber="182">China</collectingCountry>
DC Graham” (
<collectionCode id="ED05AE356BD4FF70AA17FCEDFABE39ED" box="[1289,1376,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AA6CFCEDFA4239ED" box="[1394,1436,1159,1185]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Hupe Tan-che-chan
<quantity id="4CEC9B156BD4FF70AE96FCC0FDCA3989" box="[392,532,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="181" pageNumber="182" unit="m" value="1200.0" valueMax="1500.0" valueMin="900.0">900-1500 m</quantity>
” (
<collectionCode id="ED05AE356BD4FF70AD30FCC1FD5A3989" box="[558,644,1195,1221]" pageId="181" pageNumber="182">ZMHB</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AD87FCC1FD1A3989" box="[665,708,1195,1221]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Museum Paris Mou-Pin A. David 1870” (
<collectionCode id="ED05AE356BD4FF70ABACFCC1FAD33989" box="[1202,1293,1195,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="181" pageNumber="182">MNHN</collectionCode>
),
<specimenCount id="9D12FD796BD4FF70AA3FFCC1FA933989" box="[1313,1357,1195,1221]" pageId="181" pageNumber="182" type="male">1 ♂</specimenCount>
“Siao-Lou Lou-Chan Chasseurs indigenes/ Museum Paris 1952 Coll. R. Oberthur” (
<collectionCode id="ED05AE356BD4FF70ACE3FCA5FB8739A5" box="[1021,1113,1231,1257]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="181" pageNumber="182">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD4FF72AFD9FD7CFD793F55" lastPageId="183" lastPageNumber="184" pageId="181" pageNumber="182" type="description">
<paragraph id="8BAB36F06BD4FF70AFD9FD7CFB593819" blockId="181.[151,1437,1302,2013]" pageId="181" pageNumber="182">
<emphasis id="B960EAE26BD4FF70AFD9FD7CFE87387C" bold="true" box="[199,345,1302,1328]" pageId="181" pageNumber="182">Description.</emphasis>
Length: 11.0 mm, length of elytra:
<quantity id="4CEC9B156BD4FF70ADFDFD7DFCE4387D" box="[739,826,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="181" pageNumber="182" unit="mm" value="8.1">8.1 mm</quantity>
, width:
<quantity id="4CEC9B156BD4FF70AC8CFD7CFC35387D" box="[914,1003,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="181" pageNumber="182" unit="mm" value="6.5">6.5 mm</quantity>
. Body wide, oval, dark brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
</paragraph>
<paragraph id="8BAB36F06BD4FF70AFD9FD35FC273BD5" blockId="181.[151,1437,1302,2013]" pageId="181" pageNumber="182">
Labroclypeus wide, lateral margins weakly convex and moderately convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface weakly shiny, widely dull at base, surface evenly convex, densely and coarsely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes three times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, with a short terminal seta. Eyes small, ratio diameter/ interocular width: 0.5. Frons with fine and moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126BD4FF70AEA8FE15FDCB3BD5" box="[438,533,1663,1689]" country="Myanmar" name="Chin" pageId="181" pageNumber="182">Mentum</collectingRegion>
convexly elevated and flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BD4FF70AFD9FEC9FB1C3A65" blockId="181.[151,1437,1302,2013]" pageId="181" pageNumber="182">Pronotum widest at base, lateral margins weakly convex and weakly convergent anteriorly, anterior angles sharp and distinctly produced; anterior margin convex, with widely interrupted marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum.</paragraph>
<paragraph id="8BAB36F06BD4FF70AFD9FF59FF2A3AF5" blockId="181.[151,1437,1302,2013]" pageId="181" pageNumber="182">Elytra widest at posterior third, striae finely impressed, finely punctate, intervals flat, finely and densely punctate, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06BD4FF73AFD9FFA9FED13C51" blockId="181.[151,1437,1302,2013]" lastBlockId="182.[151,1436,151,285]" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.45. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium strongly convex, coarsely and densely punctate, on apical margin with a few long setae.</paragraph>
<caption id="DF6B66786BD7FF73AF89FFFAFB463ABC" ID-DOI="http://doi.org/10.5281/zenodo.4496416" ID-Zenodo-Dep="4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="182" pageNumber="183" startId="182.[151,250,1936,1960]" targetBox="[258,1331,322,1901]" targetPageId="182">
<paragraph id="8BAB36F06BD7FF73AF89FFFAFB463ABC" blockId="182.[151,1436,1936,2032]" pageId="182" pageNumber="183">
<emphasis id="B960EAE26BD7FF73AF89FFFAFEBD3AE4" bold="true" box="[151,355,1936,1960]" pageId="182" pageNumber="183">FIGURE 41. A–D:</emphasis>
<taxonomicName id="4C144D736BD7FF73AE73FFFAFC803AE4" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[365,862,1936,1960]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="guanxianensis" status="sp. nov.">
<emphasis id="B960EAE26BD7FF73AE73FFFAFDB43AE4" box="[365,618,1936,1960]" italics="true" pageId="182" pageNumber="183">Maladera guanxianensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD7FF73AC74FFFBFC7A3AE4" box="[874,932,1937,1960]" pageId="182" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26BD7FF73AB38FFFAFB813AE4" bold="true" box="[1062,1119,1936,1960]" pageId="182" pageNumber="183">E–H:</emphasis>
<taxonomicName id="4C144D736BD7FF73AB76FFFBFEF93A80" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="guangzhaishanica" status="sp. nov.">
<emphasis id="B960EAE26BD7FF73AB76FFFBFA9B3AE4" box="[1128,1349,1936,1960]" italics="true" pageId="182" pageNumber="183">M. guangzhaishanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD7FF73AE2CFFDFFEB73A80" box="[306,361,1973,1996]" pageId="182" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26BD7FF73AEF6FFDEFDC93A80" bold="true" box="[488,535,1972,1996]" pageId="182" pageNumber="183">I–L:</emphasis>
<taxonomicName id="4C144D736BD7FF73AD01FFDFFC163A80" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[543,968,1972,1996]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="shiruguanensis" status="sp. nov.">
<emphasis id="B960EAE26BD7FF73AD01FFDFFD053A80" box="[543,731,1972,1996]" italics="true" pageId="182" pageNumber="183">M. shiruguanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD7FF73ACCDFFDFFBD43A80" box="[979,1034,1973,1996]" pageId="182" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
(holotype).
<emphasis id="B960EAE26BD7FF73AB96FFDEFB063A80" bold="true" box="[1160,1240,1972,1996]" pageId="182" pageNumber="183">A, E, I:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26BD7FF73AE05FFB2FEA93ABC" bold="true" box="[283,375,2008,2032]" pageId="182" pageNumber="183">C, G, K:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26BD7FF73AD6BFFB2FD133ABC" bold="true" box="[629,717,2008,2032]" pageId="182" pageNumber="183">D, H, L:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06BD6FF72AFD9F8FDFAAA3CB9" blockId="183.[151,1437,151,825]" pageId="183" pageNumber="184">Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae absent; posterior ventral margin almost straight, not widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia moderately long and wide, flattened, ratio width/length: 1/3.4, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at two thirds of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with a very superficial fine punctures; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical.</paragraph>
<paragraph id="8BAB36F06BD6FF72AFD9F995FD793F55" blockId="183.[151,1437,151,825]" box="[199,679,511,537]" pageId="183" pageNumber="184">
Aedeagus:
<figureCitation id="132F2A756BD6FF72AE5AF995FE4D3F55" box="[324,403,511,537]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="183" pageNumber="184">Fig. 41</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BD6FF72AD23F995FD7D3F55" box="[573,675,511,537]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="183" pageNumber="184">Fig. 41D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AFD9FA48FA433F2D" pageId="183" pageNumber="184" type="diagnosis">
<paragraph id="8BAB36F06BD6FF72AFD9FA48FA433F2D" blockId="183.[151,1437,151,825]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AFD9FA48FE9C3F70" bold="true" box="[199,322,546,572]" pageId="183" pageNumber="184">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BD6FF72AE51FA49FC523F71" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[335,908,547,573]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="guanxiamensis" status="sp. nov.">
<emphasis id="B960EAE26BD6FF72AE51FA49FDAF3F70" box="[335,625,547,573]" italics="true" pageId="183" pageNumber="184">Maladera guanxiamensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD6FF72AC83FA49FC3E3F71" box="[925,992,547,573]" pageId="183" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
differs from all other species of the
<taxonomicName id="4C144D736BD6FF72AF89FA2DFE423F2D" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[151,412,583,609]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="granuligera">
<emphasis id="B960EAE26BD6FF72AF89FA2DFE423F2D" box="[151,412,583,609]" italics="true" pageId="183" pageNumber="184">Maladera granuligera</emphasis>
</taxonomicName>
group in having the apical apophysis of the phallobase dorsal rather than lateral.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AFD9FA01FEED3FE6" pageId="183" pageNumber="184" type="etymology">
<paragraph id="8BAB36F06BD6FF72AFD9FA01FEED3FE6" blockId="183.[151,1437,151,825]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AFD9FA01FE903FC9" bold="true" box="[199,334,619,645]" pageId="183" pageNumber="184">Etymology.</emphasis>
The name of the new species is derived from its
<typeStatus id="54AF88526BD6FF72AC68FA01FC783FC9" box="[886,934,619,645]" pageId="183" pageNumber="184">type</typeStatus>
locality, Guanxian (adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AFD9FAD8FC8C3E59" pageId="183" pageNumber="184" type="description">
<paragraph id="8BAB36F06BD6FF72AFD9FAD8FC8C3E59" blockId="183.[151,1437,151,825]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AFD9FAD8FEE13F80" bold="true" box="[199,319,690,716]" pageId="183" pageNumber="184">Variation.</emphasis>
Length: 10.1–11.0 mm, length of elytra:
<quantity id="4CEC9B156BD6FF72AC3DFAD9FC713F81" box="[803,943,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="8.1" metricValueMin="7.1" pageId="183" pageNumber="184" unit="mm" value="7.6" valueMax="8.1" valueMin="7.1">7.1–8.1 mm</quantity>
, width:
<quantity id="4CEC9B156BD6FF72AB11FAD8FB423F81" box="[1039,1180,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="183" pageNumber="184" unit="mm" value="6.35" valueMax="6.5" valueMin="6.2">6.2–6.5 mm</quantity>
. Female: eyes smaller than in male, ratio of diameter/interocular width: 0.4; antennal club with three antennomeres, distinctly shorter than remaining antennomeres combined; pygidium weakly convex.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AFD9FB74FD103E75" box="[199,718,798,825]" pageId="183" pageNumber="184" type="distribution">
<paragraph id="8BAB36F06BD6FF72AFD9FB74FD103E75" blockId="183.[151,1437,151,825]" box="[199,718,798,825]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AFD9FB74FEBF3E74" bold="true" box="[199,353,798,824]" pageId="183" pageNumber="184">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BD6FF72AEC8FB75FDE83E75" box="[470,566,799,825]" captionStart="FIGURE 108" captionStartId="355.[151,250,1963,1987]" captionTargetBox="[214,1370,1124,1931]" captionTargetId="figure-60@355.[208,1379,1118,1940]" captionTargetPageId="355" captionText="FIGURE 108. Distribution of the Maladera granuligera group: Maladera baoxingensis sp. n., M. guangzhaishanica sp. n., M. guanxianensis sp. n., M. lushanensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496610" httpUri="https://zenodo.org/record/4496610/files/figure.png" pageId="183" pageNumber="184">Fig. 108</figureCitation>
) and
<tableCitation id="C696034B6BD6FF72AD6AFB75FD163E75" box="[628,712,799,825]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="183" pageNumber="184">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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