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<document id="27B534EB6482242D16DEA7A41084D87D" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B76FFDCAF89FAD8FD1E3F2D" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera shaowuensis Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="24" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="02AE1A93920CC869F2540D2C4AFD0EB0">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="5590ADEC8C7A8D9F282DBF7D2A5A1146">Bai, Ming</mods:namePart>
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<treatment id="03BD87E66B76FFDCAF89FAD8FD1E3F2D" ID-DOI="http://doi.org/10.5281/zenodo.4534471" ID-GBIF-Taxon="177323604" ID-Zenodo-Dep="4534471" LSID="urn:lsid:plazi:treatment:03BD87E66B76FFDCAF89FAD8FD1E3F2D" httpUri="http://treatment.plazi.org/id/03BD87E66B76FFDCAF89FAD8FD1E3F2D" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
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<heading id="D0E3819C6B76FFD2AF89FAD8FCC33F81" bold="true" box="[151,797,690,717]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="B960EAE26B76FFD2AF89FAD8FCC33F81" bold="true" box="[151,797,690,717]" pageId="23" pageNumber="24">
<taxonomicName id="4C144D736B76FFD2AF89FAD8FD103F80" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,718,690,717]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="shaowuensis" status="sp. nov.">
<emphasis id="B960EAE26B76FFD2AF89FAD8FE733F81" bold="true" box="[151,429,690,717]" italics="true" pageId="23" pageNumber="24">Maladera shaowuensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B76FFD2ADC5FAD9FCC33F81" box="[731,797,691,717]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="C30E657B6B76FFD2AF89FABDFE8F3FBD" box="[151,337,727,753]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAB36F06B76FFD2AF89FABDFE8F3FBD" blockId="23.[151,797,690,753]" box="[151,337,727,753]" pageId="23" pageNumber="24">
<figureCitation id="132F2A756B76FFD2AF89FABDFF203FBD" box="[151,254,727,753]" captionStart="FIGURE 3" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[196,1389,193,1881]" captionTargetId="figure-19@24.[186,1402,181,1884]" captionTargetPageId="24" captionText="FIGURE 3. AE: Maladera secreta (Brenske, 1897) (holotype: Autoserica cruralis Frey, 1972); FI: M. shaowuensis Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); JM: M. euphorbiae (Burmeister, 1855) (China: Xinjiang: Ining). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496322" httpUri="https://zenodo.org/record/4496322/files/figure.png" pageId="23" pageNumber="24">Figures 3</figureCitation>
FI, 84
</paragraph>
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<subSubSection id="C30E657B6B76FFD2AF89FB75FB013E11" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BAB36F06B76FFD2AF89FB75FB013E11" blockId="23.[151,1437,798,2013]" pageId="23" pageNumber="24">
<emphasis id="B960EAE26B76FFD2AF89FB75FE613E74" bold="true" box="[151,447,798,825]" pageId="23" pageNumber="24">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B76FFD2AED8FB75FB013E11" ID-GBIF-Occurrence="3034556592" collectionCode="ZFMK" collectorName="M. Nikodym &amp; Coll. Milan Nikodym" country="China" location="Fujian" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="holotype">
<typeStatus id="54AF88526B76FFD2AED8FB75FDED3E75" box="[454,563,799,825]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
:
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<collectingCountry id="F30376606B76FFD2AD7CFB75FD763E75" box="[610,680,799,825]" name="China" pageId="23" pageNumber="24">China</collectingCountry>
prov.
<collectingRegion id="49D0F8126B76FFD2ADF0FB75FCE83E75" box="[750,822,799,825]" country="China" name="Fujian" pageId="23" pageNumber="24">Fujian</collectingRegion>
bor. occ. Shaowu env.
<quantity id="4CEC9B156B76FFD2AB26FB74FBA63E75" box="[1080,1144,798,825]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="24" unit="km" value="25.0">25km</quantity>
road Shaowu -Taining 5.- 10.7.
<collectorName id="26E153266B76FFD2AFC8FB29FEB93E11" box="[214,359,835,861]" pageId="23" pageNumber="24">M. Nikodym</collectorName>
lgt. 1991/ [ex]
<collectorName id="26E153266B76FFD2AD0AFB29FCD73E11" box="[532,777,835,861]" pageId="23" pageNumber="24">Coll. Milan Nikodym</collectorName>
, Praha/ 546
<taxonomicName id="4C144D736B76FFD2AC8AFB29FC353E11" box="[916,1003,835,861]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
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).
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</paragraph>
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<subSubSection id="C30E657B6B76FFDCAFD9FB0CFCC33C29" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAB36F06B76FFD2AFD9FB0CFA4A3EE9" blockId="23.[151,1437,798,2013]" pageId="23" pageNumber="24">
<emphasis id="B960EAE26B76FFD2AFD9FB0CFE873ECC" bold="true" box="[199,345,870,896]" pageId="23" pageNumber="24">Description.</emphasis>
Length:
<quantity id="4CEC9B156B76FFD2AEA3FB0DFDFC3ECD" box="[445,546,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="23" pageNumber="24" unit="mm" value="10.1">10.1 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B76FFD2ADF8FB0DFCE33ECD" box="[742,829,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="23" pageNumber="24" unit="mm" value="6.9">6.9 mm</quantity>
, width:
<quantity id="4CEC9B156B76FFD2AC88FB0DFC313ECD" box="[918,1007,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="23" pageNumber="24" unit="mm" value="6.2">6.2 mm</quantity>
. Body oblong, reddish brown, antenna yellowish, surface dull, anterior labroclypeus shiny, except a few short setae on head and sides of elytra, glabrous.
</paragraph>
<paragraph id="8BAB36F06B76FFD2AF89FBC5FC9A39A5" blockId="23.[151,1437,798,2013]" pageId="23" pageNumber="24">
Labroclypeus wide and subtrapezoidal, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded; anterior margin weakly emarginate medially; margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface basally dull, weakly convex medially, finely, densely punctate, with a few single erect setae on anterior third; frontoclypeal suture finely incised, angled medially; smooth area anterior to eye convex, twice as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely sparsely punctate, terminal seta present. Frons with moderately dense, irregular, fine punctures, with a few single setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.52. Antenna with ten antennomeres; club with three antennomeres and straight, little longer than remaining antennomeres combined.
<collectingRegion id="49D0F8126B76FFD2AE0AFCA5FEAD39A5" box="[276,371,1231,1257]" country="Myanmar" name="Chin" pageId="23" pageNumber="24">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B76FFD2AFD9FC99FCFA38D1" blockId="23.[151,1437,798,2013]" pageId="23" pageNumber="24">Pronotum narrow, widest at base, lateral margins weakly convex and moderately convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly convex, with fine marginal line being medially widely interrupted, base without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margin sparsely finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BAB36F06B76FFD2AFD9FDCDFABB3BAD" blockId="23.[151,1437,798,2013]" pageId="23" pageNumber="24">Elytra widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures, penultimate interval with a few single short, erect setae, otherwise with minute setae in punctures; epipleural margin robust, ending at convex external apical angle of elytra, epipleura sparsely setose; apical border of elytra broadly membraneous, with a robust rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, nearly glabrous, metasternal disc sparsely covered with longer erect setae; metacoxa with minute setae in punctures and with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.86. Pygidium strongly convex, dull, finely and densely punctate, without smooth midline, with a few long setae along apical margin.</paragraph>
<paragraph id="8BAB36F06B76FFDCAF89FE81FAE33C0D" blockId="23.[151,1437,798,2013]" lastBlockId="25.[151,1437,151,609]" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">Legs moderately long and wide; femora shiny, with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, serrated; posterior ventral margin smooth and in apical half concave, distinctly widened at apex, apex with a blunt angle, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and moderately wide, widest at middle, ratio of width/length: 1/2.3, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a serrated line beside dorsal margin and few short single spines close to dorsal margin; lateral face longitudinally convex, finely densely punctate, with minute setae in punctures; ventral margin finely serrate, with five robust equidistant setae; medial face smooth and glabrous; apex finely serrate, very shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres glabrous and with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<caption id="DF6B66786B79FFDDAF89FF1EFD853AB4" ID-DOI="http://doi.org/10.5281/zenodo.4496322" ID-Zenodo-Dep="4496322" httpUri="https://zenodo.org/record/4496322/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1908,1932]" targetBox="[196,1389,193,1881]" targetPageId="24">
<paragraph id="8BAB36F06B79FFDDAF89FF1EFD853AB4" blockId="24.[151,1436,1908,2040]" pageId="24" pageNumber="25">
<emphasis id="B960EAE26B79FFDDAF89FF1EFE913AC0" bold="true" box="[151,335,1908,1932]" pageId="24" pageNumber="25">FIGURE 3. AE:</emphasis>
<taxonomicName id="4C144D736B79FFDDAE48FF1EFD6F3AC0" authority="(Brenske, 1897)" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[342,689,1908,1932]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="secreta">
<emphasis id="B960EAE26B79FFDDAE48FF1EFDD63AC0" box="[342,520,1908,1932]" italics="true" pageId="24" pageNumber="25">Maladera secreta</emphasis>
(
<bibRefCitation id="EF854B016B79FFDDAD0BFF1EFD743AC0" author="Brenske, E." box="[533,682,1908,1932]" pageId="24" pageNumber="25" pagination="345 - 468" refId="ref331423" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske, 1897</bibRefCitation>
)
</taxonomicName>
(holotype:
<taxonomicName id="4C144D736B79FFDDAC38FF1EFBBE3AC0" authority="Frey, 1972" authorityName="G. Frey" authorityYear="1972" box="[806,1120,1908,1933]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B960EAE26B79FFDDAC38FF1EFC343AC0" box="[806,1002,1908,1932]" italics="true" pageId="24" pageNumber="25">Autoserica cruralis</emphasis>
Frey, 1972
</taxonomicName>
);
<emphasis id="B960EAE26B79FFDDAB6CFF1EFB7E3AC0" bold="true" box="[1138,1184,1908,1932]" pageId="24" pageNumber="25">FI:</emphasis>
<taxonomicName id="4C144D736B79FFDDABB8FF1FFDC93AFC" authority="Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="shaowuensis" status="sp. nov.">
<emphasis id="B960EAE26B79FFDDABB8FF1FFA963AC0" box="[1190,1352,1908,1932]" italics="true" pageId="24" pageNumber="25">M. shaowuensis</emphasis>
Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B79FFDDAD3CFFF3FD873AFC" box="[546,601,1945,1968]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26B79FFDDADC6FFF2FCCC3AFC" bold="true" box="[728,786,1944,1968]" pageId="24" pageNumber="25">JM:</emphasis>
<taxonomicName id="4C144D736B79FFDDAC07FFF3FBA83AFC" authority="(Burmeister, 1855)" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[793,1142,1944,1968]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="euphorbiae">
<emphasis id="B960EAE26B79FFDDAC07FFF3FC703AFC" box="[793,942,1944,1968]" italics="true" pageId="24" pageNumber="25">M. euphorbiae</emphasis>
(
<bibRefCitation id="EF854B016B79FFDDACA3FFF2FBAE3AFC" author="Burmeister, H." box="[957,1136,1944,1968]" pageId="24" pageNumber="25" refId="ref331559" refString="Burmeister, H. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. T. C. F. Enslin, Berlin, x + 569 pp." type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
(China: Xinjiang: Ining).
<emphasis id="B960EAE26B79FFDDAA9AFFF2FF193A98" bold="true" pageId="24" pageNumber="25">A, F, J:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26B79FFDDAD0BFFD6FDB13A98" bold="true" box="[533,623,1980,2004]" pageId="24" pageNumber="25">D, H, L:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26B79FFDDACD2FFD6FB983A98" bold="true" box="[972,1094,1980,2004]" pageId="24" pageNumber="25">B, C, G, K:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26B79FFDDAA59FFD6FA423A98" bold="true" box="[1351,1436,1980,2004]" pageId="24" pageNumber="25">E, I, M:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06B78FFDCAF89F921FCC33C29" blockId="25.[151,1437,151,609]" box="[151,797,331,357]" pageId="25" pageNumber="26">
Aedeagus:
<figureCitation id="132F2A756B78FFDCAE0AF921FE8A3C29" box="[276,340,331,357]" captionStart="FIGURE 3" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[196,1389,193,1881]" captionTargetId="figure-19@24.[186,1402,181,1884]" captionTargetPageId="24" captionText="FIGURE 3. AE: Maladera secreta (Brenske, 1897) (holotype: Autoserica cruralis Frey, 1972); FI: M. shaowuensis Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); JM: M. euphorbiae (Burmeister, 1855) (China: Xinjiang: Ining). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496322" httpUri="https://zenodo.org/record/4496322/files/figure.png" pageId="25" pageNumber="26">Fig. 3</figureCitation>
FH. Habitus:
<figureCitation id="132F2A756B78FFDCAEE5F921FD973C29" box="[507,585,331,357]" captionStart="FIGURE 3" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[196,1389,193,1881]" captionTargetId="figure-19@24.[186,1402,181,1884]" captionTargetPageId="24" captionText="FIGURE 3. AE: Maladera secreta (Brenske, 1897) (holotype: Autoserica cruralis Frey, 1972); FI: M. shaowuensis Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); JM: M. euphorbiae (Burmeister, 1855) (China: Xinjiang: Ining). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496322" httpUri="https://zenodo.org/record/4496322/files/figure.png" pageId="25" pageNumber="26">Fig. 3I</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B78FFDCAFD9F904FE913CB9" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BAB36F06B78FFDCAFD9F904FE913CB9" blockId="25.[151,1437,151,609]" pageId="25" pageNumber="26">
<emphasis id="B960EAE26B78FFDCAFD9F904FE9C3CC4" bold="true" box="[199,322,366,392]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B78FFDCAE59F905FC9A3CC5" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu, Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[327,836,367,393]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="shaowuensis" status="sp. nov.">
<emphasis id="B960EAE26B78FFDCAE59F905FD963CC5" box="[327,584,367,393]" italics="true" pageId="25" pageNumber="26">Maladera shaowuensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B78FFDCAC50F905FC563CC5" box="[846,904,367,393]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
differs from the syntopically occurring
<taxonomicName id="4C144D736B78FFDCAA23F905FEA23CE2" authority="(Brenske)" baseAuthorityName="Brenske" baseAuthorityYear="1897" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secreta">
<emphasis id="B960EAE26B78FFDCAA23F905FED93CE0" italics="true" pageId="25" pageNumber="26">Maladera secreta</emphasis>
(Brenske)
</taxonomicName>
in the shape of the aedeagus: the parameres do not possess a basal lobe, and the median apical phallobasal process is nearly completely reduced in length; the parameres are slightly shorter and more compact in the new species.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B78FFDCAFD9F995FE0E3F71" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BAB36F06B78FFDCAFD9F995FE0E3F71" blockId="25.[151,1437,151,609]" pageId="25" pageNumber="26">
<emphasis id="B960EAE26B78FFDCAFD9F995FE903F55" bold="true" box="[199,334,511,537]" pageId="25" pageNumber="26">Etymology.</emphasis>
The name (adjective in the nominative singular case) of the new species is derived from the name of its
<typeStatus id="54AF88526B78FFDCAFC9FA49FED93F71" box="[215,263,547,573]" pageId="25" pageNumber="26">type</typeStatus>
locality, Shaowu.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B78FFDCAFD9FA2CFD1E3F2D" box="[199,704,582,609]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BAB36F06B78FFDCAFD9FA2CFD1E3F2D" blockId="25.[151,1437,151,609]" box="[199,704,582,609]" pageId="25" pageNumber="26">
<emphasis id="B960EAE26B78FFDCAFD9FA2CFEBF3F2C" bold="true" box="[199,353,582,608]" pageId="25" pageNumber="26">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B78FFDCAEC8FA2DFDF63F2D" box="[470,552,583,609]" captionStart="FIGURE 84" captionStartId="343.[151,250,2012,2036]" captionTargetBox="[185,1405,1132,1983]" captionTargetId="figure-53@343.[177,1411,1127,1988]" captionTargetPageId="343" captionText="FIGURE 84. Distribution of the subgenus Aserica: Maladera secreta (Brenske, 1897), M. shaowuensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496538" httpUri="https://zenodo.org/record/4496538/files/figure.png" pageId="25" pageNumber="26">Fig. 84</figureCitation>
) and
<tableCitation id="C696034B6B78FFDCAD78FA2DFD643F2D" box="[614,698,583,609]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x records from material examined herein, o records from literature, with no material of this study)." pageId="25" pageNumber="26">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>