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<document id="03129AA8D624D363C95567FA5A1132F2" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B22FF81AF89FD7CFD1E38D1" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera guangdongana Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="69" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="68" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="FAF815929D25D496565C0B283E242B0F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D52793B02765509F86FFAF509BC949D5">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="7B83FBA39290DCFB7B3C4FAFCFBFB686">Fabrizi, Silvia</mods:namePart>
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<mods:nameIdentifier id="0ADC4FC019F115FE662C0FB73B6EBDB0" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:namePart id="E449140500C0B59CFCCC6E1D359A1351">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="DCD7A3E5CBF28E61A4BCD8DB9CAEF8D0" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="DC7FA18C5202B8AB326D850C2BC7D57C" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="51D543B7CE90DB67ED07FA8DC3E0D2D0">Bai, Ming</mods:namePart>
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<mods:nameIdentifier id="D38317A7E28B53F6A7BCF9778097B14D" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:role id="CB639B75AF97DD0167233D9F549F9C65">
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<mods:namePart id="5CB2EDB20A810B1E1C5A893E35114C4C">Yang, Xing-Ke</mods:namePart>
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<mods:nameIdentifier id="F29B34745E3FEA095070FB5A23B960C9" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="C1881394E0C1576C2370A0C286CBBFCA">Ahrens, Dirk</mods:namePart>
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<mods:nameIdentifier id="5832CFF8154BED02531A576B396C1DCA" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:date id="A9F50F3E9451434306C59C9125041062">2021</mods:date>
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<mods:classification id="DC596CE033FDEF08C4465BE46B82B78C">journal article</mods:classification>
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<treatment id="03BD87E66B22FF81AF89FD7CFD1E38D1" ID-DOI="http://doi.org/10.5281/zenodo.4534505" ID-GBIF-Taxon="177323686" ID-Zenodo-Dep="4534505" LSID="urn:lsid:plazi:treatment:03BD87E66B22FF81AF89FD7CFD1E38D1" httpUri="http://treatment.plazi.org/id/03BD87E66B22FF81AF89FD7CFD1E38D1" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
<subSubSection id="C30E657B6B22FF86AF89FD7CFCE4387D" box="[151,826,1302,1329]" pageId="67" pageNumber="68" type="nomenclature">
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<heading id="D0E3819C6B22FF86AF89FD7CFCE4387D" bold="true" box="[151,826,1302,1329]" fontSize="11" level="2" pageId="67" pageNumber="68" reason="4">
<emphasis id="B960EAE26B22FF86AF89FD7CFCE4387D" bold="true" box="[151,826,1302,1329]" pageId="67" pageNumber="68">
<taxonomicName id="4C144D736B22FF86AF89FD7CFD32387C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,748,1302,1329]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="guangdongana" status="sp. nov.">
<emphasis id="B960EAE26B22FF86AF89FD7CFE15387D" bold="true" box="[151,459,1302,1329]" italics="true" pageId="67" pageNumber="68">Maladera guangdongana</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B22FF86ADE6FD7DFCE4387D" box="[760,826,1303,1329]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B22FF86AF89FD51FEB23819" box="[151,364,1339,1365]" pageId="67" pageNumber="68" type="description">
<paragraph id="8BAB36F06B22FF86AF89FD51FEB23819" blockId="67.[151,826,1302,1365]" box="[151,364,1339,1365]" pageId="67" pageNumber="68">
<figureCitation id="132F2A756B22FF86AF89FD51FED23819" box="[151,268,1339,1365]" captionStart="FIGURE 13" captionStartId="66.[151,250,1728,1752]" captionTargetBox="[158,1427,351,1695]" captionTargetId="figure-94@66.[151,1437,345,1705]" captionTargetPageId="66" captionText="FIGURE 13. A–D: Maladera riberai Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangdongana Ahrens, Fabrizi &amp; Liu, sp. n. A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496342" httpUri="https://zenodo.org/record/4496342/files/figure.png" pageId="67" pageNumber="68">Figures 13</figureCitation>
E–H, 91
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B22FF86AF89FDE9FEB03B61" pageId="67" pageNumber="68" type="materials_examined">
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<emphasis id="B960EAE26B22FF86AF89FDE9FE1B38D0" bold="true" box="[151,453,1410,1437]" pageId="67" pageNumber="68">Type material examined.</emphasis>
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<typeStatus id="54AF88526B22FF86AECEFDE9FDE638D1" box="[464,568,1411,1437]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
<specimenCount id="9D12FD796B22FF86AD5CFDE9FD8938D1" box="[578,599,1411,1437]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
“China—Guangdong Daqiao env.,
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,
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<collectingDate id="EFEEE9D86B22FF86AA60FDE9FF24388D" pageId="67" pageNumber="68" value="2002-05-01" valueMax="2002-05-03" valueMin="2002-05-01">1.- 3.v.2002</collectingDate>
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<collectorName id="26E153266B22FF86AE1BFDCDFE50388D" box="[261,398,1447,1473]" pageId="67" pageNumber="68">Dr. R. Fencl</collectorName>
lgt.” (
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)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6B22FF86AD11FDCDFCD538A9" ID-GBIF-Occurrence="3034556312" collectingDate="2002-05-01" collectingDateMax="2002-05-03" collectingDateMin="2002-05-01" collectionCode="CP, ZFMK" collectorName="Dr. R. Fencl" elevation="1100" latitude="24.9" location="China-Guangdong Daqiao" longLatPrecision="1249" longitude="113.01667" pageId="67" pageNumber="68" specimenCount="6" specimenCount-female="1" specimenCount-male="5" typeStatus="paratype">
<typeStatus id="54AF88526B22FF86AD11FDCDFD5E388D" box="[527,640,1447,1473]" pageId="67" pageNumber="68" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D12FD796B22FF86AD95FDCCFD12388D" box="[651,716,1446,1473]" pageId="67" pageNumber="68" type="male">5 ♂♂</specimenCount>
,
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“
<location id="8ECB602B6B22FF86AC0CFDCDFB90388D" LSID="urn:lsid:plazi:treatment:03BD87E66B22FF81AF89FD7CFD1E38D1:8ECB602B6B22FF86AC0CFDCDFB90388D" box="[786,1102,1447,1473]" latitude="24.9" longLatPrecision="1249" longitude="113.01667" name="China-Guangdong Daqiao" pageId="67" pageNumber="68">China—Guangdong Daqiao</location>
env.,
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,
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,
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</date>
,
<collectorName id="26E153266B22FF86AEBDFDA1FDEC38A9" box="[419,562,1483,1509]" pageId="67" pageNumber="68">Dr. R. Fencl</collectorName>
lgt.” (
<collectionCode id="ED05AE356B22FF86AD61FDA1FD7D38A9" box="[639,675,1483,1509]" pageId="67" pageNumber="68">CP</collectionCode>
,
<collectionCode id="ED05AE356B22FF86ADAFFDA1FCDD38A9" box="[689,771,1483,1509]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="67" pageNumber="68" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B22FF86AC07FDA1FEB53B61" ID-GBIF-Occurrence="3034555708" collectingDate="2013-04-18" collectingDateMax="2013-04-21" collectingDateMin="2013-04-18" collectionCode="ZFMK" collectorName="J. Hajek &amp; J. Ruzicka" country="China" elevation="690" latitude="24.933332" location="Nanling National Nature Reserve" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" pageId="67" pageNumber="68" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="paratype">
<specimenCount id="9D12FD796B22FF86AC07FDA1FC9838A9" box="[793,838,1483,1509]" pageId="67" pageNumber="68" type="male">1 ♂</specimenCount>
“
<collectingCountry id="F30376606B22FF86AC44FDA1FC7D38A9" box="[858,931,1483,1509]" name="China" pageId="67" pageNumber="68">China</collectingCountry>
:
<collectingRegion id="49D0F8126B22FF86ACACFDA1FBA638A9" box="[946,1144,1483,1509]" country="China" name="Guangdong" pageId="67" pageNumber="68">Guangdong prov.</collectingRegion>
<collectingMunicipality id="6BCFAC8A6B22FF86AB9FFDA1FE593B45" pageId="67" pageNumber="68">
<location id="8ECB602B6B22FF86AB9FFDA1FF2F3B45" LSID="urn:lsid:plazi:treatment:03BD87E66B22FF81AF89FD7CFD1E38D1:8ECB602B6B22FF86AB9FFDA1FF2F3B45" country="China" latitude="24.933332" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" name="Nanling National Nature Reserve" pageId="67" pageNumber="68" stateProvince="Guangdong">Nanling National Nature Reserve</location>
Dadongshan
</collectingMunicipality>
,
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<collectingDate id="EFEEE9D86B22FF86AE8CFD85FDEB3B45" box="[402,565,1519,1545]" pageId="67" pageNumber="68" value="2013-04-18" valueMax="2013-04-21" valueMin="2013-04-18">18.-21.iv.2013</collectingDate>
</date>
(border of mixed forest, at light)
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,
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,
<quantity id="4CEC9B156B22FF86ABA3FD85FADA3B45" box="[1213,1284,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="67" pageNumber="68" unit="m" value="690.0">
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<collectorName id="26E153266B22FF86AA15FD85FAB93B45" box="[1291,1383,1519,1545]" pageId="67" pageNumber="68">J. Hájek</collectorName>
&amp;
<collectorName id="26E153266B22FF86AA94FD85FF283B61" pageId="67" pageNumber="68">J. Růžicka</collectorName>
” (
<collectionCode id="ED05AE356B22FF86AE10FE79FEBC3B61" box="[270,354,1555,1581]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="67" pageNumber="68" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B22FF81AFD9FE5CFD783915" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="description">
<paragraph id="8BAB36F06B22FF86AFD9FE5CFA483B39" blockId="67.[151,1437,1410,2013]" pageId="67" pageNumber="68">
<emphasis id="B960EAE26B22FF86AFD9FE5CFE873B1C" bold="true" box="[199,345,1590,1616]" pageId="67" pageNumber="68">Description.</emphasis>
Length:
<quantity id="4CEC9B156B22FF86AEA5FE5DFDCC3B1D" box="[443,530,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="67" pageNumber="68" unit="mm" value="9.8">9.8 mm</quantity>
, length of elytra: 7.0 mm, width:
<quantity id="4CEC9B156B22FF86AC9FFE5CFC073B1D" box="[897,985,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="67" pageNumber="68" unit="mm" value="5.9">5.9 mm</quantity>
. Body oval, dark brown, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull, glabrous, except for a few small setae on the head and elytra.
</paragraph>
<paragraph id="8BAB36F06B22FF86AFD9FE15FDB03AF5" blockId="67.[151,1437,1410,2013]" pageId="67" pageNumber="68">
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a distinct blunt angle, margins moderately reflexed; anterior margin very shallowly emarginate medially; surface convex medially, weakly shiny, coarsely punctate, with a few short, erect setae; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and moderately wide, finely punctate, with two short terminal setae. Frons with fine, moderately dense punctures, glabrous except for a few setae beside eyes and frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.56. Antenna yellow, with 10 antennomeres; club with three antennomeres, little longer than remaining antennomeres combined.
<collectingRegion id="49D0F8126B22FF86AF89FFF5FF283AF5" box="[151,246,1951,1977]" country="Myanmar" name="Chin" pageId="67" pageNumber="68">Mentum</collectingRegion>
anteriorly elevated and flattened.
</paragraph>
<paragraph id="8BAB36F06B22FF81AFD9FFA9FABC3DB5" blockId="67.[151,1437,1410,2013]" lastBlockId="68.[151,1437,151,1437]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">Pronotum widest at base, lateral margins weakly and evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, anterior margin weakly convex, marginal line widely interrupted medially; surface moderately finely and densely punctate, with minute setae in punctures; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and moderately dense punctures, each bearing a single very minute seta.</paragraph>
<paragraph id="8BAB36F06B25FF81AFD9F969FB1F3CC5" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">Elytra oblong, widest at middle, striae distinctly impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures concentrated along striae, odd intervals with a few long single setae, otherwise with minute setae in punctures; epipleural margin robust, ending at the weakly convex external apical angle of elytra, epipleura densely setose, apical border with a fine rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06B25FF81AFD9F9F9FCEB3FC9" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">Ventral surface dull, thorax and metacoxa with large dense punctures, sparsely setose, metacoxa glabrous except for a few long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, with a distinct transverse row of coarse punctures each bearing a short and robust seta, some punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.66. Pygidium strongly convex, finely and densely punctate, without smooth midline, punctures with minute setae and with a few long setae along apical margin.</paragraph>
<paragraph id="8BAB36F06B25FF81AFD9FAE5FC8F3979" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">Legs moderately narrow and long; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, posterior dorsal margin not serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/ length: 1/ 3.4, dorsally sharply carinate, with two groups of spines, basal one shortly before middle, apical one at four fifths of metatibial length, basally with a few punctures, each bearing short single spines; lateral face shiny and longitudinally convex, with sparse and fine punctures, with minute setae in punctures; ventral margin serrate, with four strong equidistant spines; medial face impunctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside it with smooth subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and little longer than dorsal tibial spur. Protibia short, bidentate. Protarsal claws asymmetrical, basal lobe of inner claw widened and bluntly truncated, that if external claw with normally developed sharply pointed basal tooth.</paragraph>
<paragraph id="8BAB36F06B25FF81AFD9FC55FD783915" blockId="68.[151,1437,151,1437]" box="[199,678,1087,1113]" pageId="68" pageNumber="69">
Aedeagus:
<figureCitation id="132F2A756B25FF81AE5AFC55FE4C3915" box="[324,402,1087,1113]" captionStart="FIGURE 13" captionStartId="66.[151,250,1728,1752]" captionTargetBox="[158,1427,351,1695]" captionTargetId="figure-94@66.[151,1437,345,1705]" captionTargetPageId="66" captionText="FIGURE 13. A–D: Maladera riberai Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangdongana Ahrens, Fabrizi &amp; Liu, sp. n. A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496342" httpUri="https://zenodo.org/record/4496342/files/figure.png" pageId="68" pageNumber="69">Fig. 13</figureCitation>
E–G. Habitus:
<figureCitation id="132F2A756B25FF81AD25FC55FD7F3915" box="[571,673,1087,1113]" captionStart="FIGURE 13" captionStartId="66.[151,250,1728,1752]" captionTargetBox="[158,1427,351,1695]" captionTargetId="figure-94@66.[151,1437,345,1705]" captionTargetPageId="66" captionText="FIGURE 13. A–D: Maladera riberai Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangdongana Ahrens, Fabrizi &amp; Liu, sp. n. A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496342" httpUri="https://zenodo.org/record/4496342/files/figure.png" pageId="68" pageNumber="69">Fig. 13H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B25FF81AFD9FC08FAFA3989" pageId="68" pageNumber="69" type="diagnosis">
<paragraph id="8BAB36F06B25FF81AFD9FC08FAFA3989" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">
<emphasis id="B960EAE26B25FF81AFD9FC08FE9C3930" bold="true" box="[199,322,1122,1148]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B25FF81AE56FC09FCB83931" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[328,870,1123,1149]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="guangdongana" status="sp. nov.">
<emphasis id="B960EAE26B25FF81AE56FC09FDBB3931" box="[328,613,1123,1149]" italics="true" pageId="68" pageNumber="69">Maladera guangdongana</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B25FF81AC6FFC09FC723931" box="[881,940,1123,1149]" pageId="68" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C144D736B25FF81AB6BFC09FF0339ED" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">
<emphasis id="B960EAE26B25FF81AB6BFC09FB363931" box="[1141,1256,1123,1149]" italics="true" pageId="68" pageNumber="69">M. riberai</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B25FF81AFF8FCEDFEFF39ED" box="[230,289,1159,1185]" pageId="68" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
in shape of the aedeagus, differing by the distinctly widened base of the left paramere. From
<taxonomicName id="4C144D736B25FF81AA3DFCEDFE7C3989" authority="Kobayashi, 1991" authorityName="Kobayashi" authorityYear="1991" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="taoyuanensis">
<emphasis id="B960EAE26B25FF81AA3DFCEDFF033988" italics="true" pageId="68" pageNumber="69">M. taoyuanensis</emphasis>
Kobayashi, 1991
</taxonomicName>
from
<collectingCountry id="F30376606B25FF81AEF8FCC1FDE53989" box="[486,571,1195,1221]" name="Taiwan" pageId="68" pageNumber="69">Taiwan</collectingCountry>
, it can be distinguished by the right paramere being wider distally.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B25FF81AFD9FCA5FE6F3842" pageId="68" pageNumber="69" type="etymology">
<paragraph id="8BAB36F06B25FF81AFD9FCA5FE6F3842" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">
<emphasis id="B960EAE26B25FF81AFD9FCA5FE9039A5" bold="true" box="[199,334,1231,1257]" pageId="68" pageNumber="69">Etymology.</emphasis>
The name of the new species is derived from its occurrence in
<collectingRegion id="49D0F8126B25FF81AB02FCA5FAD739A5" box="[1052,1289,1231,1257]" country="China" name="Guangdong" pageId="68" pageNumber="69">Guangdong province</collectingRegion>
(adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B25FF81AFD9FD7CFA923835" pageId="68" pageNumber="69" type="description">
<paragraph id="8BAB36F06B25FF81AFD9FD7CFA923835" blockId="68.[151,1437,151,1437]" pageId="68" pageNumber="69">
<emphasis id="B960EAE26B25FF81AFD9FD7CFEE1387C" bold="true" box="[199,319,1302,1328]" pageId="68" pageNumber="69">Variation.</emphasis>
Length:
<quantity id="4CEC9B156B25FF81AEB6FD7CFD9F387D" box="[424,577,1302,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.05" metricValueMin="0.98" pageId="68" pageNumber="69" unit="mm" value="10.15" valueMax="10.5" valueMin="9.8">9.8–10.5 mm</quantity>
, length of elytra: 7.0–
<quantity id="4CEC9B156B25FF81AC5EFD7CFC44387D" box="[832,922,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="68" pageNumber="69" unit="mm" value="7.5">7.5 mm</quantity>
, width:
<quantity id="4CEC9B156B25FF81ACE6FD7CFB5A387D" box="[1016,1156,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.1" metricValueMin="5.9" pageId="68" pageNumber="69" unit="mm" value="6.0" valueMax="6.1" valueMin="5.9">5.9–6.1 mm</quantity>
. In one
<typeStatus id="54AF88526B25FF81ABFCFD7DFA9C387D" box="[1250,1346,1303,1329]" pageId="68" pageNumber="69" type="paratype">paratype</typeStatus>
the left paramere is as long as the right paramere, as in
<taxonomicName id="4C144D736B25FF81ADDDFD51FCE63819" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[707,824,1339,1365]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="riberai">
<emphasis id="B960EAE26B25FF81ADDDFD51FCE63819" box="[707,824,1339,1365]" italics="true" pageId="68" pageNumber="69">M. riberai</emphasis>
</taxonomicName>
, however, it is basally distictly widened, and at apex curved ventrally. Female: Eyes slightly smaller and pygidium less convex, antennal club as long as in male.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B25FF81AFD9FDE8FD1E38D1" box="[199,704,1410,1437]" pageId="68" pageNumber="69" type="distribution">
<paragraph id="8BAB36F06B25FF81AFD9FDE8FD1E38D1" blockId="68.[151,1437,151,1437]" box="[199,704,1410,1437]" pageId="68" pageNumber="69">
<emphasis id="B960EAE26B25FF81AFD9FDE8FEBF38D0" bold="true" box="[199,353,1410,1436]" pageId="68" pageNumber="69">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B25FF81AEC8FDE9FDF638D1" box="[470,552,1411,1437]" captionStart="FIGURE 91" captionStartId="347.[151,250,1019,1043]" captionTargetBox="[218,1367,187,989]" captionTargetId="figure-20@347.[213,1374,181,995]" captionTargetPageId="347" captionText="FIGURE 91. Distribution of the subgenus Omaladera: Maladera enigma sp. n., M. formosae (Brenske, 1898), M fusca (Frey, 1972), M. guangdongana sp. n., M. hongkongica (Brenske, 1898)." figureDoi="http://doi.org/10.5281/zenodo.4496562" httpUri="https://zenodo.org/record/4496562/files/figure.png" pageId="68" pageNumber="69">Fig. 91</figureCitation>
) and
<tableCitation id="C696034B6B25FF81AD78FDE9FD6438D1" box="[614,698,1411,1437]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="68" pageNumber="69">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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