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<document id="192219B4DEE6BB883E64F0F953339827" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66A22FE80AF89FD7CFD103E75" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera parva" docType="treatment" docVersion="10" lastPageNumber="326" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="324" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="9B7CF02EA21AE2F63B024B3BC10EC15D">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="0736AB9C80A22DAF60D67EAC89ADA1BB">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="677711B1912EB48C236EB0407C3694D8">Bai, Ming</mods:namePart>
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<mods:namePart id="3B2743D30CADC01A47675275808274D1">Ahrens, Dirk</mods:namePart>
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<taxonomicName id="4C144D736A22FE86AF89FD7CFDD7387C" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[151,521,1302,1329]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B960EAE26A22FE86AF89FD7CFDD7387C" bold="true" box="[151,521,1302,1329]" pageId="323" pageNumber="324">
<emphasis id="B960EAE26A22FE86AF89FD7CFE86387D" bold="true" box="[151,344,1302,1329]" italics="true" pageId="323" pageNumber="324">Maladera parva</emphasis>
(
<bibRefCitation id="EF854B016A22FE86AE79FD7CFDDF387C" author="Moser, J." box="[359,513,1302,1328]" pageId="323" pageNumber="324" pagination="325 - 343" refId="ref334156" refString="Moser, J. (1908) Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe Entomologique de Belgique, 52, 325 - 343. https: // doi. org / 10.5962 / bhl. part. 5012" type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</emphasis>
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<figureCitation id="132F2A756A22FE86AF89FD51FED03819" box="[151,270,1339,1365]" captionStart="FIGURE 77" captionStartId="324.[151,250,1598,1622]" captionTargetBox="[161,1422,358,1563]" captionTargetId="figure-93@324.[151,1436,345,1575]" captionTargetPageId="324" captionText="FIGURE 77. A–D: Maladera parva (Moser, 1908) (Paralectotype: Central Tonkin, Chiem Hoa); E–H: M. punctulata (Frey, 1972). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496522" httpUri="https://zenodo.org/record/4496522/files/figure.png" pageId="323" pageNumber="324">Figures 77</figureCitation>
A–D, 130
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A22FE86AF89FD15FCEB3899" pageId="323" pageNumber="324" type="reference_group">
<paragraph id="8BAB36F06A22FE86AF89FD15FD3F38DB" blockId="323.[151,821,1407,1493]" box="[151,737,1407,1431]" pageId="323" pageNumber="324">
<treatmentCitationGroup id="AB0411DE6A22FE86AF89FD15FD3F38DB" box="[151,737,1407,1431]" pageId="323" pageNumber="324">
<treatmentCitation id="0AB510E16A22FE86AF89FD15FDF638DB" author="Moser, J." box="[151,552,1407,1431]" page="326" pageId="323" pageNumber="324" year="1908">
<taxonomicName id="4C144D736A22FE86AF89FD15FDF638DB" authority="Moser, 1908: 326" authorityName="Moser" authorityPageNumber="326" authorityYear="1908" box="[151,552,1407,1431]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B960EAE26A22FE86AF89FD15FEB238DB" box="[151,364,1407,1431]" italics="true" pageId="323" pageNumber="324">Cephaloserica parva</emphasis>
<bibRefCitation id="EF854B016A22FE86AE6DFDEAFDF638DB" author="Moser, J." box="[371,552,1407,1431]" pageId="323" pageNumber="324" pagination="325 - 343" refId="ref334156" refString="Moser, J. (1908) Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe Entomologique de Belgique, 52, 325 - 343. https: // doi. org / 10.5962 / bhl. part. 5012" type="journal article" year="1908">Moser, 1908: 326</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16A22FE86AD2FFD15FD0338DB" author="Ahrens, D." box="[561,733,1407,1431]" page="71" pageId="323" pageNumber="324" year="2002">
<bibRefCitation id="EF854B016A22FE86AD2FFD15FD0338DB" author="Ahrens, D." box="[561,733,1407,1431]" pageId="323" pageNumber="324" pagination="53 - 91" refId="ref329519" refString="Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91." type="journal article" year="2002">Ahrens 2002: 71</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BAB36F06A22FE86AF89FDF4FD5B38FA" blockId="323.[151,821,1407,1493]" box="[151,645,1438,1462]" pageId="323" pageNumber="324">
<treatmentCitationGroup id="AB0411DE6A22FE86AF89FDF4FD5B38FA" box="[151,645,1438,1462]" pageId="323" pageNumber="324">
<taxonomicName id="4C144D736A22FE86AF89FDF4FDD038FA" authority=": Ahrens 2006 b: 237" authorityName="Ahrens" authorityPageNumber="237" authorityYear="2006" box="[151,526,1438,1462]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B960EAE26A22FE86AF89FDF4FEE238F9" box="[151,316,1438,1462]" italics="true" pageId="323" pageNumber="324">Maladera parva</emphasis>
:
<treatmentCitation id="0AB510E16A22FE86AE57FDF4FDD038FA" author="Ahrens, D." box="[329,526,1438,1462]" page="237" pageId="323" pageNumber="324" year="2006">
<bibRefCitation id="EF854B016A22FE86AE57FDF4FDD038FA" author="Ahrens, D." box="[329,526,1438,1462]" pageId="323" pageNumber="324" refId="ref329946" refString="Ahrens, D. (2006 b) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book" year="2006">Ahrens 2006b: 237</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AB510E16A22FE86AD06FDF4FD5F38FA" author="Ahrens, D." box="[536,641,1438,1462]" page="24" pageId="323" pageNumber="324" year="2007">
<bibRefCitation id="EF854B016A22FE86AD06FDF4FD5F38FA" author="Ahrens, D." box="[536,641,1438,1462]" pageId="323" pageNumber="324" pagination="1 - 51" refId="ref330155" refString="Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1" type="journal article" year="2007">2007b: 24</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36F06A22FE86AF89FDD6FCEB3899" blockId="323.[151,821,1407,1493]" box="[151,821,1468,1493]" pageId="323" pageNumber="324">
<treatmentCitationGroup id="AB0411DE6A22FE86AF89FDD6FCEB3899" box="[151,821,1468,1493]" pageId="323" pageNumber="324">
<taxonomicName id="4C144D736A22FE86AF89FDD6FDF23899" authority="Miyake, 1991: 9" authorityName="Miyake" authorityPageNumber="9" authorityYear="1991" box="[151,556,1468,1493]" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="kojimai">
<emphasis id="B960EAE26A22FE86AF89FDD6FEA33898" box="[151,381,1468,1492]" italics="true" pageId="323" pageNumber="324">Cephaloserica kojimai</emphasis>
Miyake, 1991: 9
</taxonomicName>
, syn. by
<treatmentCitation id="0AB510E16A22FE86AD9BFDD6FCEF3899" author="Ahrens, D." box="[645,817,1468,1493]" page="71" pageId="323" pageNumber="324" year="2002">
<bibRefCitation id="EF854B016A22FE86AD9BFDD6FCEF3899" author="Ahrens, D." box="[645,817,1468,1493]" pageId="323" pageNumber="324" pagination="53 - 91" refId="ref329519" refString="Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91." type="journal article" year="2002">Ahrens 2002: 71</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A22FE86AF89FE69FD4A3A71" pageId="323" pageNumber="324" type="materials_examined">
<paragraph id="8BAB36F06A22FE86AF89FE69FD403B51" blockId="323.[151,1437,1538,2033]" box="[151,670,1538,1565]" pageId="323" pageNumber="324">
<emphasis id="B960EAE26A22FE86AF89FE69FE613B50" bold="true" box="[151,447,1538,1565]" pageId="323" pageNumber="324">
<typeStatus id="54AF88526A22FE86AF89FE69FF0C3B51" box="[151,210,1539,1565]" pageId="323" pageNumber="324">Type</typeStatus>
material examined.
</emphasis>
See
<bibRefCitation id="EF854B016A22FE86AEEAFE69FD443B51" author="Ahrens, D." box="[500,666,1539,1565]" pageId="323" pageNumber="324" pagination="53 - 91" refId="ref329519" refString="Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91." type="journal article" year="2002">Ahrens (2002)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAB36F06A22FE86AFD9FE4CFD4A3A71" blockId="323.[151,1437,1538,2033]" pageId="323" pageNumber="324">
<emphasis id="B960EAE26A22FE86AFD9FE4CFDE93B0C" bold="true" box="[199,567,1574,1600]" pageId="323" pageNumber="324">Additional material examined.</emphasis>
<materialsCitation id="3B7C3CAD6A22FE86AD5EFE4CFE4A3B29" ID-GBIF-Occurrence="3034556539" collectingDate="1991-05-19" collectionCode="MNAFU" collectorName="Wang Yinglun &amp; Tian Binggang" country="China" location="Menglun" pageId="323" pageNumber="324" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan">
<emphasis id="B960EAE26A22FE86AD5EFE4CFD4C3B0C" bold="true" box="[576,658,1574,1600]" pageId="323" pageNumber="324">
<collectingCountry id="F30376606A22FE86AD5EFE4CFD533B0C" box="[576,653,1574,1600]" name="China" pageId="323" pageNumber="324">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A22FE86AD85FE4DFD003B0D" box="[667,734,1575,1601]" pageId="323" pageNumber="324" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D12FD796A22FE86ADF5FE4DFCC93B0D" box="[747,791,1575,1601]" pageId="323" pageNumber="324" type="female">1 ♀</specimenCount>
“
<location id="8ECB602B6A22FE86AC37FE4DFC4D3B0D" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AC37FE4DFC4D3B0D" box="[809,915,1575,1601]" country="China" name="Menglun" pageId="323" pageNumber="324" stateProvince="Yunnan">Menglun</location>
,
<collectingRegion id="49D0F8126A22FE86ACBEFE4DFC273B0D" box="[928,1017,1575,1601]" country="China" name="Yunnan" pageId="323" pageNumber="324">Yunnan</collectingRegion>
,
<date id="FFAA10306A22FE86AB1BFE4DFBA43B0D" box="[1029,1146,1575,1601]" pageId="323" pageNumber="324" value="1991-05-19">
<collectingDate id="EFEEE9D86A22FE86AB1BFE4DFBA43B0D" box="[1029,1146,1575,1601]" pageId="323" pageNumber="324" value="1991-05-19">19.V.1991</collectingDate>
</date>
, leg.
<collectorName id="26E153266A22FE86ABA6FE4DFA833B0D" box="[1208,1373,1575,1601]" pageId="323" pageNumber="324">Wang Yinglun</collectorName>
,
<collectorName id="26E153266A22FE86AA74FE4DFED93B29" pageId="323" pageNumber="324">Tian Binggang</collectorName>
” (
<collectionCode id="ED05AE356A22FE86AE3FFE21FE523B29" box="[289,396,1611,1637]" pageId="323" pageNumber="324">MNAFU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6A22FE86AEBFFE21FB1B3B29" ID-GBIF-Occurrence="3034556756" box="[417,1221,1610,1637]" collectingDate="1963-05-30" collectionCode="CAU" collectorName="Yang Jikun" country="China" location="Mt. Yanshan" pageId="323" pageNumber="324" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangxi">
<specimenCount id="9D12FD796A22FE86AEBFFE21FE113B29" box="[417,463,1611,1637]" pageId="323" pageNumber="324" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D12FD796A22FE86AEC3FE21FDD63B29" box="[477,520,1611,1637]" pageId="323" pageNumber="324" type="female">1 ♀</specimenCount>
“
<location id="8ECB602B6A22FE86AD02FE21FD713B29" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AD02FE21FD713B29" box="[540,687,1611,1637]" country="China" name="Mt. Yanshan" pageId="323" pageNumber="324" stateProvince="Guangxi">Mt. Yanshan</location>
,
<collectingRegion id="49D0F8126A22FE86ADA2FE21FCC13B29" box="[700,799,1611,1637]" country="China" name="Guangxi" pageId="323" pageNumber="324">Guangxi</collectingRegion>
,
<date id="FFAA10306A22FE86AC32FE20FC7F3B28" box="[812,929,1610,1636]" pageId="323" pageNumber="324" value="1963-05-30">
<collectingDate id="EFEEE9D86A22FE86AC32FE20FC7F3B28" box="[812,929,1610,1636]" pageId="323" pageNumber="324" value="1963-05-30">30.V.1963</collectingDate>
</date>
, leg.
<collectorName id="26E153266A22FE86ACFEFE21FBBC3B29" box="[992,1122,1611,1637]" pageId="323" pageNumber="324">Yang Jikun</collectorName>
” (
<collectionCode id="ED05AE356A22FE86AB62FE21FB623B29" box="[1148,1212,1611,1637]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="323" pageNumber="324">CAU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6A22FE86ABCCFE21FD183BC5" ID-GBIF-Occurrence="3034555448" collectingDate="1942-06-26" collectionCode="BMH" collectorName="T. Maa" country="China" location="China Shaowu City" pageId="323" pageNumber="324" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D12FD796A22FE86ABCCFE21FACD3B29" box="[1234,1299,1611,1637]" pageId="323" pageNumber="324" type="female">2 ♀♀</specimenCount>
“Fukien, S.
<location id="8ECB602B6A22FE86AF89FE05FEAD3BC5" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AF89FE05FEAD3BC5" box="[151,371,1647,1673]" country="China" name="China Shaowu City" pageId="323" pageNumber="324">China Shaowu City</location>
<date id="FFAA10306A22FE86AE64FE05FDD63BC4" box="[378,520,1646,1673]" pageId="323" pageNumber="324" value="1942-06-26">
<collectingDate id="EFEEE9D86A22FE86AE64FE05FDD63BC4" box="[378,520,1646,1673]" pageId="323" pageNumber="324" value="1942-06-26">1942, 26.VI.</collectingDate>
</date>
<collectorName id="26E153266A22FE86AD10FE05FD813BC5" box="[526,607,1647,1673]" pageId="323" pageNumber="324">T. Maa</collectorName>
” (
<collectionCode id="ED05AE356A22FE86AD67FE05FD603BC5" box="[633,702,1647,1673]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13650" name="Museum" pageId="323" pageNumber="324" type="Herbarium">BMH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6A22FE86ADCFFE05FF323BE1" ID-GBIF-Occurrence="3034556211" collectingDate="1942-05-16" collectionCode="BMH" collectorName="Fukien S. China Yungan &amp; T. C. Maa" country="China" location="Siying" pageId="323" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD796A22FE86ADCFFE05FD253BC5" box="[721,763,1647,1673]" pageId="323" pageNumber="324" type="male">1 ♂</specimenCount>
“
<collectorName id="26E153266A22FE86AC12FE05FBFD3BC5" box="[780,1059,1647,1673]" pageId="323" pageNumber="324">Fukien S. China Yungan</collectorName>
,
<location id="8ECB602B6A22FE86AB30FE05FBA93BC5" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AB30FE05FBA93BC5" box="[1070,1143,1647,1673]" country="China" name="Siying" pageId="323" pageNumber="324">Siying</location>
pan
<date id="FFAA10306A22FE86ABB0FE04FAFF3BC4" box="[1198,1313,1646,1672]" pageId="323" pageNumber="324" value="1942-05-16">
<collectingDate id="EFEEE9D86A22FE86ABB0FE04FAFF3BC4" box="[1198,1313,1646,1672]" pageId="323" pageNumber="324" value="1942-05-16">16.V.1942</collectingDate>
</date>
<collectorName id="26E153266A22FE86AA39FE05FA4C3BC5" box="[1319,1426,1647,1673]" pageId="323" pageNumber="324">T.C. Maa</collectorName>
” (
<collectionCode id="ED05AE356A22FE86AF81FEF9FF3A3BE1" box="[159,228,1683,1709]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13650" name="Museum" pageId="323" pageNumber="324" type="Herbarium">BMH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6A22FE86AFE8FEF9FBCB3BE1" ID-GBIF-Occurrence="3034556663" box="[246,1045,1683,1709]" collectingDate="1940-07-01" collectionCode="BMH" collectorName="Fukien S. China Yungan &amp; T. C. Maa" country="China" location="Siying" pageId="323" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD796A22FE86AFE8FEF9FEC13BE1" box="[246,287,1683,1709]" pageId="323" pageNumber="324" type="male">1 ♂</specimenCount>
“
<collectorName id="26E153266A22FE86AE30FEF9FD9E3BE1" box="[302,576,1683,1709]" pageId="323" pageNumber="324">Fukien S. China Yungan</collectorName>
,
<location id="8ECB602B6A22FE86AD54FEF9FD4D3BE1" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AD54FEF9FD4D3BE1" box="[586,659,1683,1709]" country="China" name="Siying" pageId="323" pageNumber="324">Siying</location>
pan
<date id="FFAA10306A22FE86ADD8FEF9FC9F3BE1" box="[710,833,1683,1709]" pageId="323" pageNumber="324" value="1940-07-01">
<collectingDate id="EFEEE9D86A22FE86ADD8FEF9FC9F3BE1" box="[710,833,1683,1709]" pageId="323" pageNumber="324" value="1940-07-01">1.VII.1940</collectingDate>
</date>
<collectorName id="26E153266A22FE86AC58FEF9FC713BE1" box="[838,943,1683,1709]" pageId="323" pageNumber="324">T.C. Maa</collectorName>
” (
<collectionCode id="ED05AE356A22FE86ACD9FEF9FBD33BE1" box="[967,1037,1683,1709]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13650" name="Museum" pageId="323" pageNumber="324" type="Herbarium">BMH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A22FE86AB00FEF8FE943BB9" ID-GBIF-Occurrence="3034555648" collectingDate="2012-05-12" collectionCode="CASH" collectorName="A. Skale" country="Vietnam" elevation="70" latitude="21.384167" location="Me Linh" longLatPrecision="21" longitude="105.71222" municipality="Ngoc Thanh" pageId="323" pageNumber="324" specimenCount="20" stateProvince="Thai Nguyen">
<emphasis id="B960EAE26A22FE86AB00FEF8FB533BE0" bold="true" box="[1054,1165,1682,1708]" pageId="323" pageNumber="324">
<collectingCountry id="F30376606A22FE86AB00FEF8FB563BE0" box="[1054,1160,1682,1708]" name="Vietnam" pageId="323" pageNumber="324">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A22FE86AB8DFEF9FB083BE1" box="[1171,1238,1683,1709]" pageId="323" pageNumber="324" type="generic">20 ex.</specimenCount>
“N-Vietnam
<collectingRegion id="49D0F8126A22FE86AA77FEF9FEE83B9D" country="Vietnam" name="Thai Nguyen" pageId="323" pageNumber="324">Thai Nguyen Prov.</collectingRegion>
, vic
<collectingMunicipality id="6BCFAC8A6A22FE86AE71FEDDFE203B9D" box="[367,510,1719,1745]" pageId="323" pageNumber="324">Ngoc Thanh</collectingMunicipality>
,
<location id="8ECB602B6A22FE86AD12FEDDFDB13B9D" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AD12FEDDFDB13B9D" box="[524,623,1719,1745]" country="Vietnam" latitude="21.384167" longLatPrecision="21" longitude="105.71222" municipality="Ngoc Thanh" name="Me Linh" pageId="323" pageNumber="324" stateProvince="Thai Nguyen">Me Linh</location>
(IEBR station),
<date id="FFAA10306A22FE86AC30FEDDFC7D3B9D" box="[814,931,1719,1745]" pageId="323" pageNumber="324" value="2012-05-12">
<collectingDate id="EFEEE9D86A22FE86AC30FEDDFC7D3B9D" box="[814,931,1719,1745]" pageId="323" pageNumber="324" value="2012-05-12">12.V.2012</collectingDate>
</date>
,
<geoCoordinate id="EE2050376A22FE86ACAFFEDDFBED3B9D" box="[945,1075,1719,1745]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="323" pageNumber="324" precision="15" seconds="3" value="21.384167">21°23’3’’N</geoCoordinate>
,
<geoCoordinate id="EE2050376A22FE86AB5FFEDCFB023B9D" box="[1089,1244,1718,1745]" degrees="105" direction="east" minutes="42" orientation="longitude" pageId="323" pageNumber="324" precision="15" seconds="44" value="105.71222">105°42’44’’E</geoCoordinate>
,
<quantity id="4CEC9B156A22FE86ABF7FEDCFA9D3B9C" box="[1257,1347,1718,1744]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="323" pageNumber="324" unit="m" value="70.0" valueMax="80.0" valueMin="60.0">
<elevation id="0039D1C36A22FE86ABF7FEDCFA9D3B9C" box="[1257,1347,1718,1744]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="323" pageNumber="324" unit="m" value="70.0" valueMax="80.0" valueMin="60.0">60-80m</elevation>
</quantity>
, leg.
<collectorName id="26E153266A22FE86AA9FFEDDFF063BB9" pageId="323" pageNumber="324">A. Skale</collectorName>
” (
<collectionCode id="ED05AE356A22FE86AFEFFEB1FE9F3BB9" box="[241,321,1755,1781]" pageId="323" pageNumber="324">CASH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6A22FE86AE4BFEB1FC6A3BB9" ID-GBIF-Occurrence="3034556794" box="[341,948,1754,1781]" collectionCode="MNHN" country="Vietnam" pageId="323" pageNumber="324" specimenCount="7">
<specimenCount id="9D12FD796A22FE86AE4BFEB1FE533BB9" box="[341,397,1755,1781]" pageId="323" pageNumber="324" type="generic">7 ex.</specimenCount>
“Museum Paris Tonkin Langue 1886” (
<collectionCode id="ED05AE356A22FE86AC51FEB1FC753BB9" box="[847,939,1755,1781]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="323" pageNumber="324">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A22FE86ACA1FEB0FD7D3A55" ID-GBIF-Occurrence="3034556837" collectingDate="1993-05-11" collectingDateMax="1993-05-15" collectingDateMin="1993-05-11" collectionCode="CP" collectorName="Pha-Khab Pacholatko &amp; Dembicky" country="Thailand" location="Thailand" pageId="323" pageNumber="324" specimenCount="1" specimenCount-male="1">
<emphasis id="B960EAE26A22FE86ACA1FEB0FBEB3BB8" bold="true" box="[959,1077,1754,1780]" pageId="323" pageNumber="324">
<collectingCountry id="F30376606A22FE86ACA1FEB0FBEE3BB8" box="[959,1072,1754,1780]" name="Thailand" pageId="323" pageNumber="324">Thailand</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A22FE86AB22FEB1FBB93BB9" box="[1084,1127,1755,1781]" pageId="323" pageNumber="324" type="male">1 ♂</specimenCount>
“THAI
<date id="FFAA10306A22FE86ABD8FEB0FA833BB9" box="[1222,1373,1754,1781]" pageId="323" pageNumber="324" value="1993-05-11" valueMax="1993-05-15" valueMin="1993-05-11">
<collectingDate id="EFEEE9D86A22FE86ABD8FEB0FA833BB9" box="[1222,1373,1754,1781]" pageId="323" pageNumber="324" value="1993-05-11" valueMax="1993-05-15" valueMin="1993-05-11">11-15.V.1993</collectingDate>
</date>
<collectorName id="26E153266A22FE86AA7AFEB1FE553A55" pageId="323" pageNumber="324">Nan-Pha-Khab Pacholátko</collectorName>
&amp;
<collectorName id="26E153266A22FE86AEAFFE95FDF83A55" box="[433,550,1791,1817]" pageId="323" pageNumber="324">Dembicky</collectorName>
leg.” (
<collectionCode id="ED05AE356A22FE86AD6DFE95FD443A55" box="[627,666,1791,1817]" pageId="323" pageNumber="324">CP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A22FE86ADB1FE95FD4F3A71" ID-GBIF-Occurrence="3034556780" collectingDate="1997-05-31" collectionCode="CP" collectorName="Vit Kuban" country="Laos" elevation="600" latitude="21.0" location="Laos" longLatPrecision="1266" longitude="101.416664" pageId="323" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Louangnamtha">
<emphasis id="B960EAE26A22FE86ADB1FE95FD2F3A55" bold="true" box="[687,753,1791,1817]" pageId="323" pageNumber="324">
<collectingCountry id="F30376606A22FE86ADB1FE95FD333A55" box="[687,749,1791,1817]" name="Laos" pageId="323" pageNumber="324">Laos</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A22FE86ADE7FE95FCFA3A55" box="[761,804,1791,1817]" pageId="323" pageNumber="324" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6A22FE86AC28FE95FCAF3A55" LSID="urn:lsid:plazi:treatment:03BD87E66A22FE80AF89FD7CFD103E75:8ECB602B6A22FE86AC28FE95FCAF3A55" box="[822,881,1791,1817]" country="Laos" latitude="21.0" longLatPrecision="1266" longitude="101.416664" name="Laos" pageId="323" pageNumber="324" stateProvince="Louangnamtha">Laos</location>
,
<date id="FFAA10306A22FE86AC60FE95FC323A55" box="[894,1004,1791,1817]" pageId="323" pageNumber="324" value="1997-05-31">
<collectingDate id="EFEEE9D86A22FE86AC60FE95FC323A55" box="[894,1004,1791,1817]" pageId="323" pageNumber="324" value="1997-05-31">31.v.1997</collectingDate>
</date>
<collectingRegion id="49D0F8126A22FE86ACEAFE95FB423A55" box="[1012,1180,1791,1817]" country="Laos" name="Louangnamtha" pageId="323" pageNumber="324">Louangnamtha</collectingRegion>
pr.
<geoCoordinate id="EE2050376A22FE86ABD6FE95FAF73A55" box="[1224,1321,1791,1817]" degrees="21" direction="north" minutes="00" orientation="latitude" pageId="323" pageNumber="324" precision="925" value="21.0">21°00’N</geoCoordinate>
<geoCoordinate id="EE2050376A22FE86AA2FFE94FA433A55" box="[1329,1437,1790,1817]" degrees="101" direction="east" minutes="25" orientation="longitude" pageId="323" pageNumber="324" precision="925" value="101.416664">101°25’E</geoCoordinate>
<collectingRegion id="49D0F8126A22FE86AF89FF49FE933A71" box="[151,333,1827,1853]" country="Laos" name="Louangnamtha" pageId="323" pageNumber="324">Louang Namtha</collectingRegion>
<collectorName id="26E153266A22FE86AE4DFF49FE193A71" box="[339,455,1827,1853]" pageId="323" pageNumber="324">Vit Kuban</collectorName>
leg.,
<quantity id="4CEC9B156A22FE86AD1BFF48FD993A70" box="[517,583,1826,1852]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="323" pageNumber="324" unit="m" value="600.0">
<elevation id="0039D1C36A22FE86AD1BFF48FD993A70" box="[517,583,1826,1852]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="323" pageNumber="324" unit="m" value="600.0">600m</elevation>
</quantity>
” (
<collectionCode id="ED05AE356A22FE86AD7FFF49FD563A71" box="[609,648,1827,1853]" pageId="323" pageNumber="324">CP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A22FE80AFD9FF2CFD793E59" lastPageId="325" lastPageNumber="326" pageId="323" pageNumber="324" type="description">
<paragraph id="8BAB36F06A22FE86AFD9FF2CFCFC3AC9" blockId="323.[151,1437,1538,2033]" pageId="323" pageNumber="324">
<emphasis id="B960EAE26A22FE86AFD9FF2CFEAB3A2C" bold="true" box="[199,373,1862,1888]" pageId="323" pageNumber="324">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156A22FE86AEC0FF2CFDE63A2D" box="[478,568,1862,1889]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="323" pageNumber="324" unit="mm" value="5.6">5.6 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156A22FE86AC19FF2DFCBF3A2D" box="[775,865,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="323" pageNumber="324" unit="mm" value="4.1">4.1 mm</quantity>
, width:
<quantity id="4CEC9B156A22FE86ACDEFF2DFBC53A2D" box="[960,1051,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="323" pageNumber="324" unit="mm" value="3.4">3.4 mm</quantity>
. Body short-oval, reddish to dark brown and shiny; antenna yellow, dorsal surface glabrous.
</paragraph>
<paragraph id="8BAB36F06A22FE81AFD9FFE5FCED3C0D" blockId="323.[151,1437,1538,2033]" lastBlockId="324.[151,1437,151,321]" lastPageId="324" lastPageNumber="325" pageId="323" pageNumber="324">
Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle; margins distinctly reflexed, anteriorly distinctly emarginate medially; surface flat, finely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and moderately curved; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus short and moderately wide, finely punctate, with one terminal seta. Frons with fine, dense punctures, with a few single setae beside eyes. Eyes moderately large, ratio of diameter/interocular width: 0.6. Antenna with 10 antennomeres; club with three antennomeres, little shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126A25FE81AE43F94DFE623C0D" box="[349,444,295,321]" country="Myanmar" name="Chin" pageId="324" pageNumber="325">Mentum</collectingRegion>
anteriorly elevated and flattened.
</paragraph>
<caption id="DF6B66786A25FE81AF89FE54FDF13BD2" ID-DOI="http://doi.org/10.5281/zenodo.4496522" ID-Zenodo-Dep="4496522" httpUri="https://zenodo.org/record/4496522/files/figure.png" pageId="324" pageNumber="325" startId="324.[151,250,1598,1622]" targetBox="[161,1422,358,1563]" targetPageId="324">
<paragraph id="8BAB36F06A25FE81AF89FE54FDF13BD2" blockId="324.[151,1436,1598,1694]" pageId="324" pageNumber="325">
<emphasis id="B960EAE26A25FE81AF89FE54FEBF3B1A" bold="true" box="[151,353,1598,1622]" pageId="324" pageNumber="325">FIGURE 77. A–D:</emphasis>
<taxonomicName id="4C144D736A25FE81AE77FE54FD773B1A" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[361,681,1598,1622]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="324" pageNumber="325" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B960EAE26A25FE81AE77FE54FDCE3B1A" box="[361,528,1598,1622]" italics="true" pageId="324" pageNumber="325">Maladera parva</emphasis>
(
<bibRefCitation id="EF854B016A25FE81AD01FE54FD7D3B1A" author="Moser, J." box="[543,675,1598,1622]" pageId="324" pageNumber="325" pagination="325 - 343" refId="ref334156" refString="Moser, J. (1908) Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe Entomologique de Belgique, 52, 325 - 343. https: // doi. org / 10.5962 / bhl. part. 5012" type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
(Paralectotype: Central Tonkin, Chiem Hoa);
<emphasis id="B960EAE26A25FE81AB99FE54FB1F3B1A" bold="true" box="[1159,1217,1598,1622]" pageId="324" pageNumber="325">E–H:</emphasis>
<taxonomicName id="4C144D736A25FE81ABD7FE55FF083B36" authority="(Frey, 1972)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1972" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="324" pageNumber="325" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="B960EAE26A25FE81ABD7FE55FA863B1A" box="[1225,1368,1598,1622]" italics="true" pageId="324" pageNumber="325">M. punctulata</emphasis>
(Frey, 1972)
</taxonomicName>
.
<emphasis id="B960EAE26A25FE81AFC1FE08FEC83B36" bold="true" box="[223,278,1634,1658]" pageId="324" pageNumber="325">A, E:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26A25FE81AD7FFE08FD423B36" bold="true" box="[609,668,1634,1658]" pageId="324" pageNumber="325">C, G:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26A25FE81ACEAFE08FBF73B36" bold="true" box="[1012,1065,1634,1658]" pageId="324" pageNumber="325">B, F:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26A25FE81AA39FE08FABC3B36" bold="true" box="[1319,1378,1634,1658]" pageId="324" pageNumber="325">D, H:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06A25FE81AFD9FEA7FC093A1F" blockId="324.[151,1437,1740,2019]" pageId="324" pageNumber="325">Pronotum subtrapezoidal, widest at base, lateral margins strongly convex and convergent anteriorly; anterior angles weakly produced and sharp; posterior angles blunt; anterior margin moderately convexly produced medially, anterior marginal line fine and complete; surface finely and densely punctate; anterior and lateral margins densely setose. Scutellum wide and dull, triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BAB36F06A25FE81AFD9FF36FE333AAE" blockId="324.[151,1437,1740,2019]" pageId="324" pageNumber="325">Elytra oblong, widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, dense punctures concentrated along striae, glabrous; epipleural margin robust, ending at blunt external apical angle of elytra, apex of elytra concavely trunctae; epipleura densely setose; apical border chitinous, without a rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06A24FE80AFD9F8FDFF303CC5" blockId="325.[151,1437,151,825]" pageId="325" pageNumber="326">Ventral surface shiny, metasternum and metacoxa with fine and dense punctures, sparsely setose; metacoxa glabrous except for a several long setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.47. Abdominal sternites, in addition to fine and dense punctures, each with a distinct transverse row of punctures each bearing a short and fine seta, otherwise punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Pygidium moderately convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.</paragraph>
<paragraph id="8BAB36F06A24FE80AFD9F9F9FC7B3FBD" blockId="325.[151,1437,151,825]" pageId="325" pageNumber="326">Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur shiny, anterior margin acute, lacking an adjacent serrated line; posterior ventral margin convex, weakly widened in apical half and smooth, dorsal posterior margin smooth, finely sparsely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/2.55, dorsal margin sharply carinate, with two group of spines, basal at one third, apical at two thirds of metatibial length; lateral face longitudinally convex, with sparse and fine punctures; ventral margin serrate, with three equidistant spines; medial face impunctate; apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres glabrous ventrally and with a strongly serrated ridge, beside which is not a strong longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia short, bidentate, moderately long. Protarsal claws symmetric.</paragraph>
<paragraph id="8BAB36F06A24FE80AFD9FA91FD793E59" blockId="325.[151,1437,151,825]" box="[199,679,763,789]" pageId="325" pageNumber="326">
Aedeagus:
<figureCitation id="132F2A756A24FE80AE5AFA91FE4A3E59" box="[324,404,763,789]" captionStart="FIGURE 77" captionStartId="324.[151,250,1598,1622]" captionTargetBox="[161,1422,358,1563]" captionTargetId="figure-93@324.[151,1436,345,1575]" captionTargetPageId="324" captionText="FIGURE 77. A–D: Maladera parva (Moser, 1908) (Paralectotype: Central Tonkin, Chiem Hoa); E–H: M. punctulata (Frey, 1972). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496522" httpUri="https://zenodo.org/record/4496522/files/figure.png" pageId="325" pageNumber="326">Fig. 77</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756A24FE80AD23FA91FD7A3E59" box="[573,676,763,789]" captionStart="FIGURE 77" captionStartId="324.[151,250,1598,1622]" captionTargetBox="[161,1422,358,1563]" captionTargetId="figure-93@324.[151,1436,345,1575]" captionTargetPageId="324" captionText="FIGURE 77. A–D: Maladera parva (Moser, 1908) (Paralectotype: Central Tonkin, Chiem Hoa); E–H: M. punctulata (Frey, 1972). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496522" httpUri="https://zenodo.org/record/4496522/files/figure.png" pageId="325" pageNumber="326">Fig. 77D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A24FE80AFD9FB74FD103E75" box="[199,718,798,825]" pageId="325" pageNumber="326" type="distribution">
<paragraph id="8BAB36F06A24FE80AFD9FB74FD103E75" blockId="325.[151,1437,151,825]" box="[199,718,798,825]" pageId="325" pageNumber="326">
<emphasis id="B960EAE26A24FE80AFD9FB74FEBF3E74" bold="true" box="[199,353,798,824]" pageId="325" pageNumber="326">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756A24FE80AEC8FB75FDE83E75" box="[470,566,799,825]" captionStart="FIGURE 130" captionStartId="366.[151,250,1942,1966]" captionTargetBox="[232,1356,1126,1912]" captionTargetId="figure-64@366.[226,1362,1122,1918]" captionTargetPageId="366" captionText="FIGURE 130. Distribution of Maladera species with uncertain systematic status: Maladera panyuensis sp. n., M. parva (Moser, 1908), Maladera punctulata (Frey, 1972), M. rotunda (Arrow, 1946), M. tibialis (Brenske, 1898), M. trifidiforceps sp. n., M. wandingana sp. n." figureDoi="http://doi.org/10.5281/zenodo.4496670" httpUri="https://zenodo.org/record/4496670/files/figure.png" pageId="325" pageNumber="326">Fig. 130</figureCitation>
) and
<tableCitation id="C696034B6A24FE80AD6AFB75FD163E75" box="[628,712,799,825]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="325" pageNumber="326">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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