treatments-xml/data/03/BD/87/03BD87A2C662A210F899FEF7F5EF4982.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.3739808" ID-GBIF-Dataset="b7e93c79-44a6-4d9c-8b78-dcb62033fb0d" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3739808" approvalRequired="13" approvalRequired_for_treatments="13" checkinTime="1585778501182" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03BD87A2C662A210F899FEF7F5EF4982" docLanguage="en" docName="hbmw_9_Hipposideridae_210.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Hipposideros beatus K. Andersen 1906" docType="treatment" docVersion="16" lastPageNumber="247" masterDocId="FF84FFDAC676A204FFF8FF9AFFEF4346" masterDocTitle="Hipposideridae" masterLastPageNumber="258" masterPageNumber="227" pageNumber="247" updateTime="1658320025573" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Hipposideridae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>227</mods:start>
<mods:end>258</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3739808</mods:identifier>
<mods:identifier type="GBIF-Dataset">b7e93c79-44a6-4d9c-8b78-dcb62033fb0d</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3739808</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6470474" ID-GBIF-Taxon="163433800" ID-Zenodo-Dep="6470474" LSID="urn:lsid:plazi:treatment:03BD87A2C662A210F899FEF7F5EF4982" httpUri="http://treatment.plazi.org/id/03BD87A2C662A210F899FEF7F5EF4982" lastPageNumber="247" pageId="20" pageNumber="247">
<heading pageId="20" pageNumber="247">
<subSubSection box="[1889,1969,365,426]" pageId="20" pageNumber="247" type="multiple">
<paragraph blockId="20.[1887,3216,365,522]" box="[1889,1969,365,426]" pageId="20" pageNumber="247">
<figureCitation box="[1889,1969,365,426]" captionStart="Plate 18: Hipposideridae" captionStartId="18.[172,215,4473,4502]" captionTargetBox="[0,3699,0,4866]" captionTargetPageId="17" captionText="47. Short-tailed Leaf-nosed Bat (Hipposideros curtus), 49. Aellens Leaf-nosed Bat (Hipposideros marisae), 50. Schneiders Leaf-nosed Bat (Hipposideros speoris), 51. Spurred Leaf-nosed Bat (Hipposideros calcaratus), 52. Fawn-colored Leaf-nosed Bat (Hipposideros ceruinus), 53. Coxs Leaf-nosed Bat (Hipposideros coxi), 54. Cantors Leaf-nosed Bat (Hipposideros galeritus), 55. Aba Leaf-nosed Bat (Hipposideros abae), 56. Benito Leaf-nosed Bat (Hipposideros beatus), 57. Sundevalls Leaf-nosed Bat (Hipposideros caffer), 58. Sooty Leaf-nosed (Bat Hipposiderosfuliginosus), 59. Lamottes Leaf-nosed Bat (Hipposideros lamottei), 60. Noacks Leaf-nosed Bat (Hipposideros ruber), 61. Maghreb Leaf-nosed Bat (Hipposideros tephrus), 62. Dusky Leaf-nosed Bat (Hipposideros ater), 63. Bicolored Leaf-nosed Bat (Hipposideros bicolor), 64. Short-headed Leaf-nosed Bat (Hipposideros breviceps), 65. Least Leaf-nosed Bat (Hipposideros cineraceus), 66. Large Mindanao Leaf-nosed Bat (Hipposideros coronatus)" figureDoi="http://doi.org/10.5281/zenodo.3739814" httpUri="https://zenodo.org/record/3739814/files/figure.png" pageId="20" pageNumber="247">56.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1994,2622,365,426]" pageId="20" pageNumber="247" type="vernacular_names">
<paragraph blockId="20.[1887,3216,365,522]" box="[1994,2622,365,426]" pageId="20" pageNumber="247">
<vernacularName box="[1994,2622,365,426]" pageId="20" pageNumber="247">Benito Leaf-nosed Bat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[2687,3188,365,426]" pageId="20" pageNumber="247" type="nomenclature">
<paragraph blockId="20.[1887,3216,365,522]" box="[2687,3188,365,426]" pageId="20" pageNumber="247">
<taxonomicName authorityName="K. Andersen" authorityYear="1906" box="[2687,3188,365,426]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="beatus">
<emphasis box="[2687,3188,365,426]" italics="true" pageId="20" pageNumber="247">Hipposideros beatus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="vernacular_names">
<paragraph blockId="20.[1887,3216,365,522]" box="[1889,3216,441,468]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1889,1991,443,468]" pageId="20" pageNumber="247">French:</emphasis>
<vernacularName box="[2004,2291,441,468]" pageId="20" pageNumber="247">Phyllorhine du Benito</vernacularName>
/
<emphasis bold="true" box="[2317,2441,443,468]" pageId="20" pageNumber="247">German:</emphasis>
<vernacularName box="[2453,2742,441,468]" pageId="20" pageNumber="247">Benito-Rundblattnase</vernacularName>
/
<emphasis bold="true" box="[2768,2893,443,468]" pageId="20" pageNumber="247">Spanish:</emphasis>
<vernacularName box="[2906,3216,441,468]" pageId="20" pageNumber="247">Hiposidérido de Benito</vernacularName>
</paragraph>
<paragraph blockId="20.[1887,3216,365,522]" box="[1888,2521,495,522]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1888,2222,497,522]" pageId="20" pageNumber="247">Other common names:</emphasis>
<vernacularName box="[2234,2521,495,522]" pageId="20" pageNumber="247">Benito Roundleaf Bat</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="20" pageNumber="247" type="reference_group">
<paragraph blockId="20.[2712,3516,594,1166]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[2713,2926,594,633]" pageId="20" pageNumber="247">Taxonomy.</emphasis>
<taxonomicName authority="K. Andersen, 1906" authorityName="K. Andersen" authorityYear="1906" class="Mammalia" family="Hipposideridae" genus="Hipposiderus" isUncertain="true" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="beatus">Hipposiderus [sic] beatus K. Andersen, 1906</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="materials_examined">
<paragraph blockId="20.[2712,3516,594,1166]" pageId="20" pageNumber="247">
<materialsCitation ID-GBIF-Occurrence="3753598325" pageId="20" pageNumber="247">
<quantity box="[2995,3147,647,686]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="20" pageNumber="247" unit="mi" value="15.0">15 miles</quantity>
[=
<quantity box="[3208,3327,647,686]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="247" unit="km" value="24.0">24 km</quantity>
] from Benito River,” Rfo Muni,
<collectingCountry box="[3141,3496,701,740]" name="Equatorial Guinea" pageId="20" pageNumber="247">Equatorial Guinea</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="discussion">
<paragraph blockId="20.[2712,3516,594,1166]" pageId="20" pageNumber="247">
<taxonomicName authorityName="K. Andersen" authorityYear="1906" box="[2714,3060,755,794]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="beatus">Hipposideros beatus</taxonomicName>
was formerly included in the
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[2861,2974,808,847]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="bicolor">bicolor</taxonomicName>
species group, but is now placed in the
<taxonomicName baseAuthorityName="Noack" baseAuthorityYear="1893" box="[2984,3076,861,900]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="ruber">ruber</taxonomicName>
species group. Validity of subspecies has not been tested with molecular data. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="distribution">
<caption inLine="true" pageId="20" pageNumber="247" startId="20.[2712,2922,1022,1061]" targetBox="[1884,2683,599,1160]" targetPageId="20">
<paragraph blockId="20.[2712,3516,594,1166]" box="[2712,3253,1022,1061]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[2712,3253,1022,1061]" pageId="20" pageNumber="247">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="20.[2712,3516,594,1166]" lastBlockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<taxonomicName authority="K Andersen, 1906" authorityName="K. Andersen" authorityYear="1906" box="[2713,3331,1075,1114]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="species" species="beatus">H. b. beatus K Andersen, 1906</taxonomicName>
— patchily in
<collectingCountry box="[2819,3070,1127,1166]" name="Sierra Leone" pageId="20" pageNumber="247">Sierra Leone</collectingCountry>
,
<collectingCountry box="[3091,3232,1127,1166]" name="Liberia" pageId="20" pageNumber="247">Liberia</collectingCountry>
,
<collectingCountry box="[3253,3397,1127,1166]" name="Guinea-Bissau" pageId="20" pageNumber="247">Guinea</collectingCountry>
,
<collectingCountry name="Ivory Coast" pageId="20" pageNumber="247">Ivory Coast</collectingCountry>
,
<collectingCountry box="[2015,2144,1179,1218]" name="Ghana" pageId="20" pageNumber="247">Ghana</collectingCountry>
,
<collectingCountry box="[2164,2308,1179,1218]" name="Nigeria" pageId="20" pageNumber="247">Nigeria</collectingCountry>
,
<collectingCountry box="[2328,2533,1179,1218]" name="Cameroon" pageId="20" pageNumber="247">Cameroon</collectingCountry>
,
<collectingCountry box="[2553,2907,1179,1218]" name="Equatorial Guinea" pageId="20" pageNumber="247">Equatorial Guinea</collectingCountry>
, and
<collectingCountry box="[3011,3142,1179,1218]" name="Gabon" pageId="20" pageNumber="247">Gabon</collectingCountry>
.
</paragraph>
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<taxonomicName authority="Verschuren, 1957" authorityName="Verschuren" authorityYear="1957" box="[1887,2490,1231,1270]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="247" phylum="Chordata" rank="subSpecies" species="beatus" subSpecies="maximus">H. b. maximus Verschuren, 1957</taxonomicName>
patchily in
<collectingCountry box="[2751,3191,1231,1270]" name="Republic of the Congo" pageId="20" pageNumber="247">Republic of the Congo</collectingCountry>
,
<collectingCountry box="[3212,3419,1231,1270]" name="Democratic Republic of the Congo" pageId="20" pageNumber="247">DR Congo</collectingCountry>
, and extreme NW
<collectingCountry box="[2140,2293,1286,1325]" name="Uganda" pageId="20" pageNumber="247">Uganda</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="description">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1886,2232,1339,1378]" pageId="20" pageNumber="247">Descriptive notes.</emphasis>
Head-body 43-50 mm, tail 20-31 mm, ear 12-17 mm, hindfoot 6-9 mm, forearm 39-48 mm; weight 6-9 g. The Benito Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with a large but simple noseleaf that has two lateral leaflets. A frontal sac is present in both sexes. Intemarial septum is not swollen and does not partially cover the nares. Ears are separate, and relatively small. Fur is fine and short, dark brown dorsally, paler ventrally.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="biology_ecology">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1885,2039,1658,1697]" pageId="20" pageNumber="247">Habitat.</emphasis>
Inhabits lowland rainforest, and is usually associated with rivers and streams. May also occupy riparian forest in
<collectingCountry box="[2546,2686,1711,1750]" name="Guinea-Bissau" pageId="20" pageNumber="247">Guinea</collectingCountry>
savanna.
</paragraph>
</subSubSection>
<subSubSection box="[1885,3070,1765,1804]" pageId="20" pageNumber="247" type="food_feeding">
<paragraph blockId="20.[1882,3513,1179,2756]" box="[1885,3070,1765,1804]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1885,2240,1765,1804]" pageId="20" pageNumber="247">Food and Feeding.</emphasis>
The Benito Leaf-nosed Bat is insectivorous.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="breeding">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1885,2069,1818,1857]" pageId="20" pageNumber="247">Breeding.</emphasis>
In
<collectingCountry box="[2142,2273,1818,1857]" name="Gabon" pageId="20" pageNumber="247">Gabon</collectingCountry>
, mating seems to occur in June-July with births of single young four months later in October—November. Females reach sexual maturity at six months, males at one year of age.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="activity">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1882,2201,1977,2016]" pageId="20" pageNumber="247">Activity patterns.</emphasis>
The Benito Leaf-nosed Bat roosts during the day under boulders and eroded roots of trees, in hollow trees, in holes in the ground, culverts under roads, and cavities under small rocks. It does not roost in large, deep caves. Echolocation call includes a F component at c.130-140 kHz.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="biology_ecology">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1884,2856,2190,2229]" pageId="20" pageNumber="247">Movements, Home range and Social organization.</emphasis>
Benito Leaf-nosed Bats roost singly, in pairs or in small family groups, typically numbering 3-4 individuals. In one study in
<collectingCountry box="[2051,2183,2296,2335]" name="Gabon" pageId="20" pageNumber="247">Gabon</collectingCountry>
, these family groups consisted of a mated pair with one young from the current year and one from the previous breeding season. These pairs broke up at the beginning of the next breeding season when new pairings were formed.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="conservation">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1884,2355,2455,2494]" pageId="20" pageNumber="247">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCNRed List. At present, the only significant threat appears to be the loss of rainforest The Benito Leaf-nosed Bat is present in several protected areas and forest reserves.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="247" type="bibRefCitation_list">
<paragraph blockId="20.[1882,3513,1179,2756]" pageId="20" pageNumber="247">
<emphasis bold="true" box="[1884,2092,2621,2650]" pageId="20" pageNumber="247">Bibliography.</emphasis>
Brosset (1966b, 1982), Grubb eta/. (1998), Happold, D.C.D. (1987), Happold, M. (2013v), Hill (1963a), Koopman (1984a), Monadjem, Richards &amp; Denys (2016), Rosevear (1965), Schütter et al. (1982), Thom &amp; Kerbis Peterhans (2009), Verschuren (1957).
</paragraph>
</subSubSection>
</treatment>
</document>