304 lines
37 KiB
XML
304 lines
37 KiB
XML
<document id="D01D90D0DA366D3C247B90E5A207B6B6" ID-DOI="10.5281/zenodo.13203792" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203792" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658458551" checkinUser="felipe" docAuthor="Minkina, Ÿukasz, Anichtchenko, Alexander, Vasiljeva, Anastasija & Skelley, Paul E." docDate="2022" docId="03BD44335E56FFE4FC8C212806F3FE68" docLanguage="en" docName="BalticJColeopterol.22.1.81-101.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Rhyparus stebnickae Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="90" masterDocId="FF843C4B5E51FFEEFF8A22690417FFB0" masterDocTitle="Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females" masterLastPageNumber="101" masterPageNumber="81" pageNumber="88" updateTime="1722698086404" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="A7B681C52F8ECED8E5D0802FE31ECFA7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A7B9751C144E553BC5063DB47C215F81">
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<mods:title id="981AB9C2CA099461032279C19AB03DA4">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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</mods:titleInfo>
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<mods:name id="EF591177506FABAEEB4555181A16F421" type="personal">
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<mods:role id="487FD8239F97BEBBDA1C35A79366FC76">
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<mods:roleTerm id="426CFD1FAD6FD2C6E5B18C371F3A9F52">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E7AAD0514AE7FDCBA81781A1BDF0F9F1">Minkina, Ÿukasz</mods:namePart>
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</mods:name>
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<mods:name id="9676384517CCB97ACB9809527141A4ED" type="personal">
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<mods:role id="00BAA1BE066279875B610622B09A5995">
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<mods:roleTerm id="4412DC542A2D5A0487D905D8079FBC35">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4A332311D780D7D8821896D65CA022D9">Anichtchenko, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="9A3CC33F6BC20A694E2862B6930EC96E" type="personal">
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<mods:role id="83CA2EC3707AC315F01007F1FC206556">
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<mods:roleTerm id="14DECAB72D8F4C1D23CF23CD4E3673CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9BEC763129613E0BE4BC1D1FA9B9EFBB">Vasiljeva, Anastasija</mods:namePart>
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</mods:name>
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<mods:name id="8F9E9855879207BC64FE07AB887854BD" type="personal">
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<mods:role id="961B8027B1A3529FBA15673352C6F69C">
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<mods:roleTerm id="0DF603C236D761CE6F94B06EBBDCBD6F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EAFC97DB5AE4E80365902589F5404814">Skelley, Paul E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F18B8BC65D6995A9DC7012E03F2C4FCF">text</mods:typeOfResource>
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<mods:relatedItem id="5B34B1581E45CD79540A80175A47E936" type="host">
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<mods:titleInfo id="B7C87B5765298362F04A20C3E524E7B8">
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<mods:title id="6E0B8866BD1146B669C7AF3BBE493A5B">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="97C1A3908688DDD32D0BE35719D19191">
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<mods:date id="14C67EDF9DC6A6CB2F68E62F17D1FF3A">2022</mods:date>
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<mods:detail id="9957CFA9B42825B42CA97B0D4F168B2B" type="volume">
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<mods:number id="2BD6FF7C4687C6AD74EE1540EDF8E7DA">22</mods:number>
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</mods:detail>
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<mods:detail id="FEA057D7EC60DEC72E8BB973C9468C01" type="issue">
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<mods:number id="D2BF6A1295F0644BF3BBD8B1EF7F7759">1</mods:number>
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</mods:detail>
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<mods:extent id="18937828319EAC1F006E4A501CF6D1C6" unit="page">
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<mods:start id="D3C3CE461B54B73BB89B2951BB846585">81</mods:start>
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<mods:end id="DFFB504FAAB840712BCDA0FBEEE3AE47">101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="945CF9B72F53C7F85ED1A3C98CE3D844">journal article</mods:classification>
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<mods:identifier id="EBA7B168AC174F8090CF1EDDC70593A3" type="DOI">10.5281/zenodo.13203792</mods:identifier>
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<mods:identifier id="476D9E96ADE056B865237B9CB93811D9" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="9CE91EA9AACEED9AD7BC282C07C0DBD2" type="Zenodo-Dep">13203792</mods:identifier>
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</mods:mods>
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<treatment id="03BD44335E56FFE4FC8C212806F3FE68" LSID="urn:lsid:plazi:treatment:03BD44335E56FFE4FC8C212806F3FE68" httpUri="http://treatment.plazi.org/id/03BD44335E56FFE4FC8C212806F3FE68" lastPageId="10" lastPageNumber="90" pageId="7" pageNumber="88">
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<subSubSection id="C30EA6AE5E56FFE9FC8C21280050FCE8" box="[774,1095,833,857]" pageId="7" pageNumber="88" type="nomenclature">
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<paragraph id="8BABF5255E56FFE9FC8C21280050FCE8" blockId="7.[774,1095,833,888]" box="[774,1095,833,857]" pageId="7" pageNumber="88">
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<heading id="D0E342495E56FFE9FC8C21280050FCE8" box="[774,1095,833,857]" fontSize="10" level="2" pageId="7" pageNumber="88" reason="7">
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||
<taxonomicName id="4C148EA65E56FFE9FC8C212807F9FCE9" ID-CoL="9WHZG" authorityName="Minkina & Anichtchenko & Vasiljeva & Skelley" authorityYear="2022" box="[774,1006,833,857]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
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<emphasis id="B96029375E56FFE9FC8C212807F9FCE9" bold="true" box="[774,1006,833,857]" italics="true" pageId="7" pageNumber="88">Rhyparus stebnickae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A253944C5E56FFE9FC7E21280050FCE8" box="[1012,1095,833,856]" pageId="7" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
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||
</heading>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C30EA6AE5E56FFE9FC8C210807FDFCC8" box="[774,1002,865,888]" pageId="7" pageNumber="88" type="description">
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||
<paragraph id="8BABF5255E56FFE9FC8C210807FDFCC8" blockId="7.[774,1095,833,888]" box="[774,1002,865,888]" pageId="7" pageNumber="88">
|
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(
|
||
<figureCitation id="132FE9A05E56FFE9FC8421080786FCC8" box="[782,913,865,888]" captionStart="Figs" captionStartId="8.[302,352,1693,1716]" captionTargetBox="[450,1200,751,1654]" captionTargetId="figure-239@8.[448,1205,750,1660]" captionTargetPageId="8" captionText="Figs. 10–12. Rhyparus stebnickae sp. nov.,, holotype: 10 – dorsal view; 11 – ventral view; 12 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203802" httpUri="https://zenodo.org/record/13203802/files/figure.png" pageId="7" pageNumber="88">Figs. 10–12</figureCitation>
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,
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||
<figureCitation id="132FE9A05E56FFE9FC17210807ADFCC8" box="[925,954,865,888]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 20–25. Heads: 20 – R. bacanensis sp. nov.,, holotype; 21 – R. buruensis sp. nov.,, holotype; 22 – R. fijiensis sp. nov.,, holotype; 23 – R. stebnickae sp. nov.,, holotype; 24 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 25 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203808" httpUri="https://zenodo.org/record/13203808/files/figure.png" pageId="7" pageNumber="88">23</figureCitation>
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,
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<figureCitation id="132FE9A05E56FFE9FC4C210807F5FCC8" box="[966,994,865,888]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with blackwhite miniatures of pygidium on leftdown side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="7" pageNumber="88">29</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C30EA6AE5E56FFE9FC8C21C900EBF96A" pageId="7" pageNumber="88" type="materials_examined">
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||
<paragraph id="8BABF5255E56FFE9FC8C21C907EEFC48" blockId="7.[774,1285,928,1016]" pageId="7" pageNumber="88">
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||
<materialsCitation id="3B7CFF785E56FFE9FC8C21C907E2FC48" country="Indonesia" county="Arfak Mts" elevation="1190" location="Dueibei" municipality="Manokwari distr." pageId="7" pageNumber="88" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
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||
<emphasis id="B96029375E56FFE9FC8C21C9078EFC08" bold="true" box="[774,921,928,952]" pageId="7" pageNumber="88">
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||
<typeStatus id="54AF4B875E56FFE9FC8C21C9072AFC08" box="[774,829,928,952]" pageId="7" pageNumber="88">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<collectingCountry id="F303B5B55E56FFE9FC1721C80012FC08" box="[925,1029,929,952]" name="Indonesia" pageId="7" pageNumber="88">Indonesia</collectingCountry>
|
||
,
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||
<collectingRegion id="49D03BC75E56FFE9FB8721C800A9FC08" box="[1037,1214,929,952]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="88">West Papua prov.</collectingRegion>
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||
,
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||
<collectingCounty id="62CA8DA95E56FFE9FB4221C80725FC68" pageId="7" pageNumber="88">Arfak Mts</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BCF6F5F5E56FFE9FCB721A807E2FC68" box="[829,1013,961,984]" pageId="7" pageNumber="88">Manokwari distr.</collectingMunicipality>
|
||
,
|
||
<location id="8ECBA3FE5E56FFE9FB8921A8004FFC68" LSID="urn:lsid:plazi:treatment:03BD44335E56FFE4FC8C212806F3FE68:8ECBA3FE5E56FFE9FB8921A8004FFC68" box="[1027,1112,961,984]" country="Indonesia" county="Arfak Mts" municipality="Manokwari distr." name="Dueibei" pageId="7" pageNumber="88" stateProvince="Papua Barat">Dueibei</location>
|
||
env., cca
|
||
<locationDeviation id="9A19EC075E56FFE9FB4B21A80789FC48" location="Warmare" pageId="7" pageNumber="88">
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||
<quantity id="4CEC58C05E56FFE9FB4B21A80112FC68" box="[1217,1285,961,984]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="88" unit="km" value="20.0">20 km</quantity>
|
||
S of Warmare
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||
</locationDeviation>
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||
,
|
||
<quantity id="4CEC58C05E56FFE9FC22218807E2FC48" box="[936,1013,993,1016]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="7" pageNumber="88" unit="m" value="1190.0">
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||
<elevation id="003912165E56FFE9FC22218807E2FC48" box="[936,1013,993,1016]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="7" pageNumber="88" unit="m" value="1190.0">1190m</elevation>
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||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E56FFE9FC8C264800EBF96A" blockId="7.[774,1286,1057,1754]" pageId="7" pageNumber="88">
|
||
<emphasis id="B96029375E56FFE9FC8C264807A7FB89" bold="true" box="[774,944,1057,1081]" pageId="7" pageNumber="88">Type material.</emphasis>
|
||
<typeStatus id="54AF4B875E56FFE9FC322648000BFB88" box="[952,1052,1057,1080]" pageId="7" pageNumber="88" type="holotype">Holotype</typeStatus>
|
||
():
|
||
<collectingCountry id="F303B5B55E56FFE9FBD2264800D3FB88" box="[1112,1220,1057,1080]" name="Indonesia" pageId="7" pageNumber="88">Indonesia</collectingCountry>
|
||
,
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<collectingRegion id="49D03BC75E56FFE9FB5A2648079EFBEA" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="88">West Papua prov.</collectingRegion>
|
||
| Arfak Mts,
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||
<quantity id="4CEC58C05E56FFE9FBA8262A007CFBEA" box="[1058,1131,1091,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="7" pageNumber="88" unit="m" value="1190.0">1190m</quantity>
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||
alt. | Dueibei env.,
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||
<date id="FFAAD3E55E56FFE9FCB5260A07F1FBCA" box="[831,998,1123,1146]" pageId="7" pageNumber="88" value="2008-02-10" valueMax="2008-02-28" valueMin="2008-02-10">10.ii28.ii.2008</date>
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||
| cca
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||
<quantity id="4CEC58C05E56FFE9FBAA260A0074FBCA" box="[1056,1123,1123,1146]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="88" unit="km" value="20.0">20 km</quantity>
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||
S of Warmare | Manokwari distr, St. Jakl lgt. || (
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||
<emphasis id="B96029375E56FFE9FBED26E90060FB2A" bold="true" box="[1127,1143,1152,1178]" pageId="7" pageNumber="88">
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||
<collectionCode id="ED056DE05E56FFE9FBED26E90060FB2A" box="[1127,1143,1152,1178]" pageId="7" pageNumber="88">Ÿ</collectionCode>
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||
</emphasis>
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||
<collectionCode id="ED056DE05E56FFE9FBFD26EA00AEFB2A" box="[1143,1209,1155,1178]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="7" pageNumber="88" type="Herbarium">MCN</collectionCode>
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||
).
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||
<typeStatus id="54AF4B875E56FFE9FB4626EA072BFB0A" pageId="7" pageNumber="88" type="paratype">Paratypes</typeStatus>
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||
(10exx.):
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||
<specimenCount id="9D123EAC5E56FFE9FC2E26CA07C5FB0A" box="[932,978,1187,1210]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
: Coll. I.R.Sc.N.B. |
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||
<collectingCountry id="F303B5B55E56FFE9FB1826CA0759FB6A" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
|
||
| Wau,
|
||
<collectingRegion id="49D03BC75E56FFE9FC1926AA07F5FB6A" box="[915,994,1219,1242]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe</collectingRegion>
|
||
prob., | Wau Ecology Institute, |
|
||
<quantity id="4CEC58C05E56FFE9FC98268A0772FB4A" box="[786,869,1251,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="88" unit="m" value="1200.0">1200 m</quantity>
|
||
, at light | I.G.: 26373 |
|
||
<date id="FFAAD3E55E56FFE9FBD2268A00DCFB4A" box="[1112,1227,1251,1274]" pageId="7" pageNumber="88" value="1981-06-18">18.vi.1981</date>
|
||
| leg. J. Van Goethem || (
|
||
<collectionCode id="ED056DE05E56FFE9FC4D276A0000FAAA" box="[967,1047,1283,1306]" pageId="7" pageNumber="88">DUBC</collectionCode>
|
||
);
|
||
<specimenCount id="9D123EAC5E56FFE9FBAD276A0040FAAA" box="[1063,1111,1283,1306]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
: Coll.R.Isc.N.B. | Canopy mission P.N.G. |
|
||
<collectingRegion id="49D03BC75E56FFE9FB9A274A00DEFA8A" box="[1040,1225,1315,1338]" country="Papua New Guinea" name="Madang" pageId="7" pageNumber="88">Madang province</collectingRegion>
|
||
| Batteta
|
||
<date id="FFAAD3E55E56FFE9FCBE272A07BAFAEA" box="[820,941,1347,1370]" pageId="7" pageNumber="88" value="1993-06-19">19 vi1993</date>
|
||
| light M1 | leg. Olivier Missa || (
|
||
<collectionCode id="ED056DE05E56FFE9FC84270A074CFACA" box="[782,859,1379,1402]" pageId="7" pageNumber="88">DUBC</collectionCode>
|
||
);
|
||
<specimenCount id="9D123EAC5E56FFE9FCE0270A078EFACA" box="[874,921,1379,1402]" count="2" pageId="7" pageNumber="88" type="generic">2ex.</specimenCount>
|
||
:
|
||
<collectingCountry id="F303B5B55E56FFE9FC15270A0072FACA" box="[927,1125,1379,1402]" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
|
||
: |
|
||
<collectingRegion id="49D03BC75E56FFE9FBFE270A00E9FACA" box="[1140,1278,1379,1402]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe Prov.</collectingRegion>
|
||
: Wau; |
|
||
<date id="FFAAD3E55E56FFE9FCD927EA07D6FA2A" box="[851,961,1411,1434]" pageId="7" pageNumber="88" value="1982-02-21">21.ii.1982</date>
|
||
|| light trap | R.T.Bell, Coll. || (
|
||
<specimenCount id="9D123EAC5E56FFE9FC8527CA072CFA0A" box="[783,827,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
<collectionCode id="ED056DE05E56FFE9FCB527CA079AFA0A" box="[831,909,1443,1466]" pageId="7" pageNumber="88">CNCW</collectionCode>
|
||
;
|
||
<specimenCount id="9D123EAC5E56FFE9FC1227CA07D2FA0A" box="[920,965,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
NHNG);
|
||
<specimenCount id="9D123EAC5E56FFE9FBA327CA004FFA0A" box="[1065,1112,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
: Papua Nlle
|
||
<collectingCountry id="F303B5B55E56FFE9FB5F27CA073CFA6A" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinée</collectingCountry>
|
||
| Morobe i.(19)80 | env de Gurakor | W.G.Ullrich || (
|
||
<collectionCode id="ED056DE05E56FFE9FCC3278A079FFA4A" box="[841,904,1507,1530]" pageId="7" pageNumber="88">ISEA</collectionCode>
|
||
);
|
||
<specimenCount id="9D123EAC5E56FFE9FC12278A07D0FA4A" box="[920,967,1507,1530]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
: New
|
||
<collectingCountry id="F303B5B55E56FFE9FB8F278A0043FA4A" box="[1029,1108,1507,1530]" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinea</collectingCountry>
|
||
: (NE) | Wau,
|
||
<collectingRegion id="49D03BC75E56FFE9FB52278A0778F9AA" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe distr.</collectingRegion>
|
||
|
|
||
<quantity id="4CEC58C05E56FFE9FCF7246A07DEF9AA" box="[893,969,1539,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="88" unit="m" value="1200.0">1200m</quantity>
|
||
, 2 10.xi.(19)61 || J. & M. Sedlacek | M.V. light trap || H.F. Howden, 1992 || (
|
||
<emphasis id="B96029375E56FFE9FC8524290708F9EA" bold="true" box="[783,799,1600,1626]" pageId="7" pageNumber="88">
|
||
<collectionCode id="ED056DE05E56FFE9FC8524290708F9EA" box="[783,799,1600,1626]" pageId="7" pageNumber="88">Ÿ</collectionCode>
|
||
</emphasis>
|
||
<collectionCode id="ED056DE05E56FFE9FC94242A074AF9EA" box="[798,861,1603,1626]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="7" pageNumber="88" type="Herbarium">MCN</collectionCode>
|
||
); 3exx.:
|
||
<collectingCountry id="F303B5B55E56FFE9FC25242A0078F9EA" box="[943,1135,1603,1626]" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
|
||
| Wau,
|
||
<quantity id="4CEC58C05E56FFE9FB3C242A0111F9EA" box="[1206,1286,1603,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="7" pageNumber="88" unit="ft" value="4000.0">4000 ft.</quantity>
|
||
|
|
||
<date id="FFAAD3E55E56FFE9FC9A240A07B2F9CA" box="[784,933,1635,1658]" pageId="7" pageNumber="88" value="1974-06-24" valueMax="1974-06-30" valueMin="1974-06-24">24 30.VI.1974</date>
|
||
| H.F. Howden ||
|
||
<taxonomicName id="4C148EA65E56FFE9FBC1240A00BAF9CA" authorityName="Westwood" authorityYear="1845" box="[1099,1197,1635,1658]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">Rhyparus</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C148EA65E56FFE9FB36240A073EF92A" authority="Ar." authorityName="Arrow" authorityYear="1935" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="striatus">striatus Ar.</taxonomicName>
|
||
| dt. Stebnicka (2exx.
|
||
<collectionCode id="ED056DE05E56FFE9FB9424EA0066F92A" box="[1054,1137,1667,1690]" pageId="7" pageNumber="88">CMNC</collectionCode>
|
||
;
|
||
<specimenCount id="9D123EAC5E56FFE9FBF624EA00BEF92A" box="[1148,1193,1667,1690]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
<collectionCode id="ED056DE05E56FFE9FB3B24EA00EEF92A" box="[1201,1273,1667,1690]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="7" pageNumber="88" type="Museum">FSCA</collectionCode>
|
||
);
|
||
<specimenCount id="9D123EAC5E56FFE9FC8C24CA0720F90A" box="[774,823,1699,1722]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
|
||
: Papua – N.
|
||
<collectingCountry id="F303B5B55E56FFE9FC4024CA000FF90A" box="[970,1048,1699,1722]" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinea</collectingCountry>
|
||
| Mainyanda
|
||
<date id="FFAAD3E55E56FFE9FB3824CA00E1F90A" box="[1202,1270,1699,1722]" pageId="7" pageNumber="88" value="1980-01">i.1980</date>
|
||
|
|
||
<quantity id="4CEC58C05E56FFE9FC8C24AA0753F96A" box="[774,836,1731,1754]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="88" unit="km" value="25.0">25km</quantity>
|
||
W Bulolo | W.G. Ullrich
|
||
<quantity id="4CEC58C05E56FFE9FBD824AA0098F96A" box="[1106,1167,1731,1754]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="88" unit="m" value="600.0">600m</quantity>
|
||
|| (
|
||
<collectionCode id="ED056DE05E56FFE9FB2524AA00E6F96A" box="[1199,1265,1731,1754]" pageId="7" pageNumber="88">ISEA</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30EA6AE5E59FFE7FEA422A90048FB59" lastPageId="9" lastPageNumber="90" pageId="8" pageNumber="89" type="description">
|
||
<paragraph id="8BABF5255E59FFE6FEA422A9073AFE68" blockId="8.[302,813,192,472]" pageId="8" pageNumber="89">
|
||
<emphasis id="B96029375E59FFE6FEA422A9067FFF68" bold="true" box="[302,616,192,216]" pageId="8" pageNumber="89">
|
||
Description of the
|
||
<typeStatus id="54AF4B875E59FFE6FE7422A90674FF68" box="[510,611,192,216]" pageId="8" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Dorsum (
|
||
<figureCitation id="132FE9A05E59FFE6FD5B22A80737FF68" box="[721,800,193,216]" captionStart="Figs" captionStartId="8.[302,352,1693,1716]" captionTargetBox="[450,1200,751,1654]" captionTargetId="figure-239@8.[448,1205,750,1660]" captionTargetPageId="8" captionText="Figs. 10–12. Rhyparus stebnickae sp. nov.,, holotype: 10 – dorsal view; 11 – ventral view; 12 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203802" httpUri="https://zenodo.org/record/13203802/files/figure.png" pageId="8" pageNumber="89">Fig. 10</figureCitation>
|
||
). Length:
|
||
<quantity id="4CEC58C05E59FFE6FE0F228805CFFF48" box="[389,472,225,248]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="89" unit="mm" value="6.5">6.5 mm</quantity>
|
||
; maximum width:
|
||
<quantity id="4CEC58C05E59FFE6FD13228806FCFF48" box="[665,747,225,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="89" unit="mm" value="1.9">1.9 mm</quantity>
|
||
. Body moderate to largesized for members of this genus, elongate, not so distinctly convex, flattened in central part; shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E59FFE6FEA4206807C5FE08" blockId="8.[302,812,513,696]" lastBlockId="8.[850,1361,193,440]" pageId="8" pageNumber="89">
|
||
Head (
|
||
<figureCitation id="132FE9A05E59FFE6FEF8206805ACFDA8" box="[370,443,513,536]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 20–25. Heads: 20 – R. bacanensis sp. nov.,, holotype; 21 – R. buruensis sp. nov.,, holotype; 22 – R. fijiensis sp. nov.,, holotype; 23 – R. stebnickae sp. nov.,, holotype; 24 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 25 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203808" httpUri="https://zenodo.org/record/13203808/files/figure.png" pageId="8" pageNumber="89">Fig. 23</figureCitation>
|
||
) shiny; transversely subhexagonal; clypeus trapezoidal in outline, anteriorly weakly sinuate, on sides weakly upturned as obtuse, quite distinct tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of very indistinct, very short, convergent ridges, nearly on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E59FFE6FCD823880147FD08" blockId="8.[850,1361,481,696]" pageId="8" pageNumber="89">Pronotum shiny; with eight low but distinct costae and seven intercostae, with two lateral, rounded lobes on each side.Anterior lobes somewhat lower and distinctly narrower than posterior; posterior lobes on the top are the widest part of pronotum. Costae of third and fourth pair not interrupted in basal part of apical half, very gently convergent,</paragraph>
|
||
<caption id="DF6BA5AD5E59FFE6FEA424F40685F964" ID-DOI="http://doi.org/10.5281/zenodo.13203802" ID-Zenodo-Dep="13203802" httpUri="https://zenodo.org/record/13203802/files/figure.png" pageId="8" pageNumber="89" startId="8.[302,352,1693,1716]" targetBox="[450,1200,751,1654]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BABF5255E59FFE6FEA424F40685F964" blockId="8.[302,1360,1693,1748]" pageId="8" pageNumber="89">
|
||
Figs. 10–12.
|
||
<taxonomicName id="4C148EA65E59FFE6FE3624F7068AF905" authorityName="Minkina & Anichtchenko & Vasiljeva & Skelley" authorityYear="2022" box="[444,669,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
|
||
<emphasis id="B96029375E59FFE6FE3624F7068AF905" box="[444,669,1694,1717]" italics="true" pageId="8" pageNumber="89">Rhyparus stebnickae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A253944C5E59FFE6FD2E24F406E0F904" box="[676,759,1693,1716]" pageId="8" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,, holotype: 10 – dorsal view; 11 – ventral view; 12 – lateral view. Scale lines: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BABF5255E58FFE7FD8A22F807FFFF14" blockId="9.[512,1000,145,164]" box="[512,1000,145,164]" pageId="9" pageNumber="90">
|
||
<emphasis id="B96029375E58FFE7FD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="9" pageNumber="90">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FF6922A806F5FE4D" blockId="9.[227,738,193,509]" pageId="9" pageNumber="90">distinctly convergent in the middle of apical third; median pair of costae in apical half become very low but without any cutpoint of interruption; second pair of costae distinctly interrupted in basal part of apical half; costae on each side with small punctures bearing very small macrosetae.All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, very distinctly concentrated around median part, all intercostae in basal part with distinct, dense punctation.</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FF69204E0628FD8E" blockId="9.[227,575,551,574]" box="[227,575,551,574]" pageId="9" pageNumber="90">Scutellum almost imperceptible.</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FF6920010624FB38" blockId="9.[227,737,616,1160]" pageId="9" pageNumber="90">Elytra shiny. Each elytron with six elevated but flattened on the top costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of punctures; on third intercostae there is short additional costae with additional row of punctures untill half of length of elytra. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures here. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and mediointernal caudal bulbs not divided; mediointernal caudal bulb shortened, transverse, internally stretched.</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FF6926DB05D9FAFB" blockId="9.[227,737,1202,1355]" pageId="9" pageNumber="90">
|
||
Pygidium (
|
||
<figureCitation id="132FE9A05E58FFE7FEDF26DB05B7FB79" box="[341,416,1202,1225]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with blackwhite miniatures of pygidium on leftdown side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="9" pageNumber="90">Fig. 29</figureCitation>
|
||
) with dense, irregularly spaced punctation, with very distinct longitudinal rib in the middle and deep excision on its sides; with distinctly longitudinal apex in the shape of volcano with sinuate top.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FF69271C0763FEA9" blockId="9.[227,737,1397,1745]" lastBlockId="9.[774,1285,193,281]" pageId="9" pageNumber="90">
|
||
Venter (
|
||
<figureCitation id="132FE9A05E58FFE7FEB3271C0590FA3C" box="[313,391,1397,1420]" captionStart="Figs" captionStartId="8.[302,352,1693,1716]" captionTargetBox="[450,1200,751,1654]" captionTargetId="figure-239@8.[448,1205,750,1660]" captionTargetPageId="8" captionText="Figs. 10–12. Rhyparus stebnickae sp. nov.,, holotype: 10 – dorsal view; 11 – ventral view; 12 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203802" httpUri="https://zenodo.org/record/13203802/files/figure.png" pageId="9" pageNumber="90">Fig. 11</figureCitation>
|
||
) shiny. Mesometaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of mesometaventral plate dense, regularly spaced, irregular in size; all punctures bearing short macrosetae. Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are slightly larger than on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FC8C232A009AFE6C" blockId="9.[774,1285,323,476]" pageId="9" pageNumber="90">Mesofemora with two distinct tubercles on lower border; metafemora with very weak tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FC8C206F0055FC74" blockId="9.[774,1286,518,964]" pageId="9" pageNumber="90">
|
||
<emphasis id="B96029375E58FFE7FC8C206F076DFDAE" bold="true" box="[774,890,518,542]" pageId="9" pageNumber="90">Variation.</emphasis>
|
||
Length of body: 6.2 7.0 mm. Intercostae of pronotum and elytra usually are shiny, but sometimes may be somewhat less shiny. Lateral lobes of pronotum may be more or less weakly developed. Density of punctation of pronotal and elytral intercostae is quite distinctly variable. Degree of concentration of punctures in median intercostae of pronotum is very weakly variable. Usually elytral costae are flattened and wide, rarely may be somewhat more distinctly convex and quite narrow. Additional rows of fine punctures observed usually on third elytral intercostae – between main rows, but there are specimens with simple punctures on all rows.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E58FFE7FC8C21870048FB59" blockId="9.[774,1285,1006,1257]" pageId="9" pageNumber="90">
|
||
<emphasis id="B96029375E58FFE7FC8C218707F1FBB6" bold="true" box="[774,998,1006,1030]" pageId="9" pageNumber="90">Sexual dimorphism.</emphasis>
|
||
Shape of mesometaventral plate (in both sexes), apex of mesotibiae (in both sexes) and shape of last abdominal ventrite and pygidium of males are typical for sexual dimorphism in genus
|
||
<taxonomicName id="4C148EA65E58FFE7FC2E2618001FFB38" authorityName="Westwood" authorityYear="1845" box="[932,1032,1137,1160]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96029375E58FFE7FC2E2618001FFB38" box="[932,1032,1137,1160]" italics="true" pageId="9" pageNumber="90">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
. Last abdominal ventrite and pygidium of females is distinctly elongate at apex (
|
||
<figureCitation id="132FE9A05E58FFE7FCCF26DB0784FB79" box="[837,915,1202,1225]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with blackwhite miniatures of pygidium on leftdown side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="9" pageNumber="90">Fig. 29</figureCitation>
|
||
). Last abdominal ventrite and pygidium of males are not modified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30EA6AE5E58FFE4FC8C277A06F3FE68" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90" type="etymology">
|
||
<paragraph id="8BABF5255E58FFE7FC8C277A00FAFADB" blockId="9.[774,1285,1298,1387]" pageId="9" pageNumber="90">
|
||
<emphasis id="B96029375E58FFE7FC8C277A0791FA9B" bold="true" box="[774,902,1299,1323]" pageId="9" pageNumber="90">Etymology.</emphasis>
|
||
Patronymic. Dedicated to
|
||
<emphasis id="B96029375E58FFE7FB12277A0112FA9A" bold="true" box="[1176,1285,1298,1322]" pageId="9" pageNumber="90">Zdzis˘awa</emphasis>
|
||
Stebnicka who noticed unusual sexual dimorphism in the genus
|
||
<taxonomicName id="4C148EA65E58FFE7FC5C273D002AFADB" authorityName="Westwood" authorityYear="1845" box="[982,1085,1364,1387]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">Rhyparus</taxonomicName>
|
||
at the first time.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E58FFE4FC8C27FC06F3FE68" blockId="9.[774,1285,1429,1745]" lastBlockId="10.[302,813,193,472]" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
|
||
<emphasis id="B96029375E58FFE7FC8C27FC0773FA1D" bold="true" box="[774,868,1429,1453]" pageId="9" pageNumber="90">Affinity.</emphasis>
|
||
See discussion and tables 1 and 2. Additional problem is that some specimens of
|
||
<taxonomicName id="4C148EA65E58FFE7FB6727DF07F3FA5D" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96029375E58FFE7FB6727DF074EFA5E" italics="true" pageId="9" pageNumber="90">R. gracilis</emphasis>
|
||
Arrow, 1905
|
||
</taxonomicName>
|
||
(which them
|
||
<taxonomicName id="4C148EA65E58FFE7FBFC27BE0112FA5E" authorityName="Minkina & Anichtchenko & Vasiljeva & Skelley" authorityYear="2022" box="[1142,1285,1495,1518]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
|
||
<emphasis id="B96029375E58FFE7FBFC27BE0112FA5E" box="[1142,1285,1495,1518]" italics="true" pageId="9" pageNumber="90">R. stebnickae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A253944C5E58FFE7FC8C279E074DF9BE" box="[774,858,1527,1550]" pageId="9" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected on the same localities on
|
||
<collectingCountry id="F303B5B55E58FFE7FC8C247E07CFF99E" box="[774,984,1559,1582]" name="Papua New Guinea" pageId="9" pageNumber="90">Papua New Guinea</collectingCountry>
|
||
) may have similar structure of elytral costae, identique structure of caudal bulbs and somewhat similar shape of lateral lobes of pronotum (posterior lateral lobes can be widely, regularly rounded, weakly developed, with very small sinuation between them and anterior lateral lobes). Anyway females of
|
||
<taxonomicName id="4C148EA65E5BFFE4FDD822A806D5FF68" authorityName="Arrow" authorityYear="1905" box="[594,706,193,216]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96029375E5BFFE4FDD822A806D5FF68" box="[594,706,193,216]" italics="true" pageId="10" pageNumber="91">R. gracilis</emphasis>
|
||
</taxonomicName>
|
||
have normally developed pygidium and last abdominal ventrite. Males of
|
||
<taxonomicName id="4C148EA65E5BFFE4FE6323680642FEA8" authorityName="Arrow" authorityYear="1905" box="[489,597,257,280]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96029375E5BFFE4FE6323680642FEA8" box="[489,597,257,280]" italics="true" pageId="10" pageNumber="91">R. gracilis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by: more distinctly convex body, posterior lateral lobes of pronotum never more distinctly developed than anterior, punctures in median intercostae of pronotum never so distinctly concentrated nearby the middle (never create so distinct longitudinal line) and elytral intercostae usually matt.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |